{"status":"ok","message-type":"work","message-version":"1.0.0","message":{"indexed":{"date-parts":[[2026,9,30]],"date-time":"2026-09-30T08:01:26Z","timestamp":1790755286644,"version":"4.1.0"},"reference-count":84,"publisher":"Wiley","issue":"10","license":[{"start":{"date-parts":[[2026,9,30]],"date-time":"2026-09-30T00:00:00Z","timestamp":1790726400000},"content-version":"vor","delay-in-days":0,"URL":"http:\/\/onlinelibrary.wiley.com\/termsAndConditions#vor"},{"start":{"date-parts":[[2026,9,30]],"date-time":"2026-09-30T00:00:00Z","timestamp":1790726400000},"content-version":"tdm","delay-in-days":0,"URL":"http:\/\/doi.wiley.com\/10.1002\/tdm_license_1.1"}],"funder":[{"DOI":"10.13039\/501100000038","name":"Natural Sciences and Engineering Research Council of Canada","doi-asserted-by":"publisher","id":[{"id":"10.13039\/501100000038","id-type":"DOI","asserted-by":"publisher"}]},{"DOI":"10.13039\/100012258","name":"Polar Knowledge Canada","doi-asserted-by":"publisher","id":[{"id":"10.13039\/100012258","id-type":"DOI","asserted-by":"publisher"}]},{"DOI":"10.13039\/100019889","name":"Weston Family Foundation","doi-asserted-by":"publisher","id":[{"id":"10.13039\/100019889","id-type":"DOI","asserted-by":"publisher"}]}],"content-domain":{"domain":["onlinelibrary.wiley.com"],"crossmark-restriction":true},"short-container-title":["Global Change Biology"],"published-print":{"date-parts":[[2026,10]]},"abstract":"<jats:title>ABSTRACT<\/jats:title>\n                  <jats:p>\n                    Climate change drives variable shifts in mammalian body sizes, yet the mechanisms underlying this variation remain unclear. Two explanations prevail: (1) higher temperatures reduce thermal constraints, favouring smaller bodies; or (2) warming enhances primary productivity, supporting larger bodies. We propose that species' overwintering strategies, which mitigate exposure to winter temperatures, determine which mechanism predominates. Using ~50\u2009years of body mass data, we tested these hypotheses in three small mammals with contrasting natural histories in the rapidly warming Yukon, Canada. Snowshoe hares (\n                    <jats:styled-content style=\"fixed-case\">\n                      <jats:italic>Lepus americanus<\/jats:italic>\n                    <\/jats:styled-content>\n                    ), exposed to ambient winter conditions, declined in mass over time and responded negatively to temperature and snow. In subnivean species, deer mice (\n                    <jats:styled-content style=\"fixed-case\">\n                      <jats:italic>Peromyscus maniculatus<\/jats:italic>\n                    <\/jats:styled-content>\n                    ) showed no long\u2010term trend but increased body mass with growing degree days, whereas red\u2010backed voles (\n                    <jats:styled-content style=\"fixed-case\">\n                      <jats:italic>Clethrionomys rutilus<\/jats:italic>\n                    <\/jats:styled-content>\n                    ) increased with temperature and snow. The results demonstrate that overwintering ecology mediates climate\u2010driven body mass change, emphasizing the role of natural history in shaping species' responses to warming.\n                  <\/jats:p>","DOI":"10.1111\/gcb.71132","type":"journal-article","created":{"date-parts":[[2026,9,30]],"date-time":"2026-09-30T07:44:03Z","timestamp":1790754243000},"update-policy":"https:\/\/doi.org\/10.1002\/crossmark_policy","source":"Crossref","is-referenced-by-count":0,"title":["Overwintering Strategies Drive Climate Change Impacts on Body Mass in Northern Mammals"],"prefix":"10.1111","volume":"32","author":[{"ORCID":"https:\/\/orcid.org\/0009-0004-1007-5927","authenticated-orcid":false,"given":"Olivia","family":"Leach","sequence":"first","affiliation":[{"name":"Department of Biology Memorial University of Newfoundland and Labrador  St. John's Newfoundland and Labrador Canada"}],"role":[{"vocabulary":"crossref","role":"author"}]},{"ORCID":"https:\/\/orcid.org\/0000-0002-3014-2056","authenticated-orcid":false,"given":"Michael J. L.","family":"Peers","sequence":"additional","affiliation":[{"name":"Department of Biology Memorial University of Newfoundland and Labrador  St. John's Newfoundland and Labrador Canada"}],"role":[{"vocabulary":"crossref","role":"author"}]},{"ORCID":"https:\/\/orcid.org\/0009-0001-7798-8884","authenticated-orcid":false,"given":"Abigail","family":"Earle","sequence":"additional","affiliation":[{"name":"Department of Biology Memorial University of Newfoundland and Labrador  St. John's Newfoundland and Labrador Canada"}],"role":[{"vocabulary":"crossref","role":"author"}]},{"ORCID":"https:\/\/orcid.org\/0000-0001-5908-3793","authenticated-orcid":false,"given":"Yasmine N.","family":"Majchrzak","sequence":"additional","affiliation":[{"name":"Department of Biology Memorial University of Newfoundland and Labrador  St. John's Newfoundland and Labrador Canada"}],"role":[{"vocabulary":"crossref","role":"author"}]},{"ORCID":"https:\/\/orcid.org\/0000-0003-1959-1077","authenticated-orcid":false,"given":"Rudy","family":"Boonstra","sequence":"additional","affiliation":[{"name":"Department of Biological Sciences University of Toronto Scarborough  Toronto Ontario Canada"}],"role":[{"vocabulary":"crossref","role":"author"}]},{"ORCID":"https:\/\/orcid.org\/0000-0001-6317-038X","authenticated-orcid":false,"given":"Stan","family":"Boutin","sequence":"additional","affiliation":[{"name":"Department of Biological Sciences University of Alberta  Edmonton Alberta Canada"}],"role":[{"vocabulary":"crossref","role":"author"}]},{"ORCID":"https:\/\/orcid.org\/0000-0002-5710-042X","authenticated-orcid":false,"given":"Dennis L.","family":"Murray","sequence":"additional","affiliation":[{"name":"Department of Biology Trent University  Peterborough Ontario Canada"}],"role":[{"vocabulary":"crossref","role":"author"}]},{"ORCID":"https:\/\/orcid.org\/0000-0003-2681-6852","authenticated-orcid":false,"given":"Thomas S.","family":"Jung","sequence":"additional","affiliation":[{"name":"Department of Environment Government of Yukon  Whitehorse Yukon Canada"},{"name":"Department of Renewable Resources University of Alberta  Edmonton Alberta Canada"}],"role":[{"vocabulary":"crossref","role":"author"}]},{"given":"B. Scott","family":"Gilbert","sequence":"additional","affiliation":[{"name":"Applied Science and Management Yukon University  Whitehorse Yukon Canada"}],"role":[{"vocabulary":"crossref","role":"author"}]},{"given":"Alice J.","family":"Kenney","sequence":"additional","affiliation":[{"name":"Department of Zoology University of British Columbia  Vancouver British Columbia Canada"}],"role":[{"vocabulary":"crossref","role":"author"}]},{"ORCID":"https:\/\/orcid.org\/0000-0002-8534-4317","authenticated-orcid":false,"given":"Eric Vander","family":"Wal","sequence":"additional","affiliation":[{"name":"Department of Biology Memorial University of Newfoundland and Labrador  St. John's Newfoundland and Labrador Canada"}],"role":[{"vocabulary":"crossref","role":"author"}]}],"member":"311","published-online":{"date-parts":[[2026,9,30]]},"reference":[{"key":"e_1_2_10_2_1","first-page":"229","volume-title":"Snow Ecology: An Interdisciplinary Examination of Snow Covered Ecosystems","author":"Aitchison C. W.","year":"2001"},{"key":"e_1_2_10_3_1","volume-title":"\u00dcber die Verh\u00e4ltnisse der W\u00e4rme\u00f6konomie der Thiere zu ihrer Gr\u00f6sse","author":"Bergmann C.","year":"1848"},{"key":"e_1_2_10_4_1","unstructured":"Black H. C.1965.\u201cAn Analysis of a Population of Snowshoe Hares Lepus americanus washingtoniiBaird in Western Oregon.\u201d Ph.D. thesis Oregon State University Corvallis."},{"issue":"4","key":"e_1_2_10_5_1","doi-asserted-by":"crossref","first-page":"165","DOI":"10.1046\/j.1472-4642.1999.00046.x","article-title":"Geographic Gradients in Body Size: A Clarification of Bergmann's Rule","volume":"5","author":"Blackburn T. M.","year":"1999","journal-title":"Diversity and Distributions"},{"issue":"1","key":"e_1_2_10_6_1","doi-asserted-by":"crossref","first-page":"335","DOI":"10.2307\/5176","article-title":"Species Number, Population Density and Body Size Relationships in Natural Communities","volume":"59","author":"Blackburn T. M.","year":"1990","journal-title":"Journal of Animal Ecology"},{"issue":"6","key":"e_1_2_10_7_1","doi-asserted-by":"crossref","first-page":"1269","DOI":"10.1111\/j.1365-2656.2006.01149.x","article-title":"Population Limitation of the Northern Red\u2010Backed Vole in the Boreal Forests of Northern Canada","volume":"75","author":"Boonstra R.","year":"2006","journal-title":"Journal of Animal Ecology"},{"key":"e_1_2_10_8_1","first-page":"216","volume-title":"Ecosystem Dynamics of the Boreal Forest: The Kluane Project","author":"Boonstra R.","year":"2001"},{"issue":"1","key":"e_1_2_10_9_1","doi-asserted-by":"crossref","first-page":"69","DOI":"10.2307\/3545676","article-title":"Population Changes of the Vertebrate Community During a Snowshoe Hare Cycle in Canada's Boreal Forest","volume":"74","author":"Boutin S.","year":"1995","journal-title":"Oikos"},{"issue":"1","key":"e_1_2_10_10_1","doi-asserted-by":"crossref","first-page":"29","DOI":"10.1111\/eva.12121","article-title":"Climate Change and Mammals: Evolutionary Versus Plastic Responses","volume":"7","author":"Boutin S.","year":"2014","journal-title":"Evolutionary Applications"},{"issue":"2","key":"e_1_2_10_11_1","first-page":"378","article-title":"glmmTMB Balances Speed and Flexibility Among Packages for Zero\u2010Inflated Generalized Linear Mixed Modeling","volume":"9","author":"Brooks M. 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M.","year":"2022","journal-title":"Ecological Applications"},{"issue":"7316","key":"e_1_2_10_17_1","doi-asserted-by":"crossref","first-page":"704","DOI":"10.1038\/nature09407","article-title":"Global Metabolic Impacts of Recent Climate Warming","volume":"467","author":"Dillon M. E.","year":"2010","journal-title":"Nature"},{"issue":"5","key":"e_1_2_10_18_1","doi-asserted-by":"crossref","first-page":"1499","DOI":"10.1111\/j.1365-2486.2012.02644.x","article-title":"Size Increase in High Elevation Ground Squirrels Over the Last Century","volume":"18","author":"Eastman L. M.","year":"2012","journal-title":"Global Change Biology"},{"issue":"2","key":"e_1_2_10_19_1","doi-asserted-by":"crossref","first-page":"115","DOI":"10.1139\/cjz-2015-0074","article-title":"Differential Sex\u2010Related Winter Energetics in Free\u2010Ranging Snowshoe Hares (\n                     Lepus americanus\n                  )","volume":"94","author":"Ellsworth E.","year":"2016","journal-title":"Canadian Journal of Zoology"},{"issue":"10","key":"e_1_2_10_20_1","doi-asserted-by":"crossref","first-page":"2098","DOI":"10.1111\/1365-2435.13630","article-title":"They Like It Cold, but Only in Winter: Climate\u2010Mediated Effects on a Hibernator","volume":"34","author":"Fietz J.","year":"2020","journal-title":"Functional Ecology"},{"issue":"6","key":"e_1_2_10_21_1","doi-asserted-by":"crossref","first-page":"285","DOI":"10.1016\/j.tree.2011.03.005","article-title":"Declining Body Size: A Third Universal Response to Warming?","volume":"26","author":"Gardner J. 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S.","year":"1981","journal-title":"Oecologia"},{"issue":"4","key":"e_1_2_10_25_1","doi-asserted-by":"crossref","first-page":"250","DOI":"10.1111\/j.1600-0587.1991.tb00659.x","article-title":"Population Dynamics of Clethrionomys and Peromyscus in Southwestern Yukon 1973\u20131989","volume":"14","author":"Gilbert B. S.","year":"1991","journal-title":"Ecography"},{"key":"e_1_2_10_26_1","first-page":"1","article-title":"Experimental Reduction of Wolves in the Yukon: Ungulate Responses and Management Implications","volume":"152","author":"Hayes R. D.","year":"2003","journal-title":"Wildlife Monographs"},{"key":"e_1_2_10_27_1","first-page":"143","volume-title":"Ecosystem Dynamics of the Boreal Forest: The Kluane Project","author":"Hodges K.","year":"2001"},{"issue":"5","key":"e_1_2_10_28_1","doi-asserted-by":"crossref","first-page":"961","DOI":"10.1093\/icb\/icx119","article-title":"To Everything There Is a Season: Summer\u2010to\u2010Winter Food Webs and the Functional Traits of Keystone Species","volume":"57","author":"Humphries M. M.","year":"2017","journal-title":"Integrative and Comparative Biology"},{"issue":"12","key":"e_1_2_10_29_1","doi-asserted-by":"crossref","first-page":"907","DOI":"10.1139\/cjz-2024-0020","article-title":"Seasonal Dynamics of Small Mammal Populations: Resource Availability and Cold Exposure Interact to Govern Abundance","volume":"102","author":"Jolly S. R.","year":"2024","journal-title":"Canadian Journal of Zoology"},{"key":"e_1_2_10_30_1","doi-asserted-by":"crossref","first-page":"2648","DOI":"10.1890\/08-1494.1","article-title":"PanTHERIA: A Species\u2010Level Database of Life History, Ecology, and Geography of Extant and Recently Extinct Mammals","volume":"90","author":"Jones K. E.","year":"2009","journal-title":"Ecology"},{"issue":"4","key":"e_1_2_10_31_1","doi-asserted-by":"crossref","first-page":"663","DOI":"10.1007\/s13364-021-00596-1","article-title":"Snow Tracking Provides Insights on Hunting Behaviour of Marten (\n                     Martes americana\n                  )","volume":"66","author":"Jung T. S.","year":"2021","journal-title":"Mammal Research"},{"key":"e_1_2_10_32_1","doi-asserted-by":"crossref","first-page":"11","DOI":"10.1093\/oso\/9780195133936.001.0001","volume-title":"Ecosystem Dynamics of the Boreal Forest: The Kluane Project","author":"Krebs C. J.","year":"2001"},{"issue":"1","key":"e_1_2_10_33_1","doi-asserted-by":"crossref","first-page":"87","DOI":"10.1111\/1365-2656.12720","article-title":"Using Experimentation to Understand the 10\u2010Year Snowshoe Hare Cycle in the Boreal Forest of North America","volume":"87","author":"Krebs C. J.","year":"2018","journal-title":"Journal of Animal Ecology"},{"key":"e_1_2_10_34_1","doi-asserted-by":"crossref","DOI":"10.3389\/fevo.2023.1187222","article-title":"Long\u2010Term Monitoring in the Boreal Forest Reveals High Spatio\u2010Temporal Variability Among Primary Ecosystem Constituents","volume":"11","author":"Krebs C. J.","year":"2023","journal-title":"Frontiers in Ecology and Evolution"},{"issue":"2","key":"e_1_2_10_35_1","doi-asserted-by":"crossref","first-page":"194","DOI":"10.1007\/BF00379239","article-title":"A Natural Feeding Experiment on a Declining Snowshoe Hare Population","volume":"70","author":"Krebs C. J.","year":"1986","journal-title":"Oecologia"},{"issue":"1","key":"e_1_2_10_36_1","doi-asserted-by":"crossref","first-page":"27","DOI":"10.1111\/1749-4877.12718","article-title":"Long\u2010Term Monitoring of Cycles in \n                     Clethrionomys rutilus\n                   in the Yukon Boreal Forest","volume":"19","author":"Krebs C. J.","year":"2024","journal-title":"Integrative Zoology"},{"key":"e_1_2_10_37_1","unstructured":"Lawrence I. E.1955.\u201cAn Ecological Study of the Snowshoe Hare Lepus americanus bairdiiHayden in the Medicine Bow National Forest of Wyoming.\u201d M.Sc. thesis University of Wyoming Laramie."},{"key":"e_1_2_10_38_1","doi-asserted-by":"crossref","first-page":"16","DOI":"10.1093\/acprof:oso\/9780199208784.003.0003","volume-title":"Sex, Size and Gender Roles: Evolutionary Studies of Sexual Size Dimorphism","author":"Lindenfors P.","year":"2007"},{"issue":"60","key":"e_1_2_10_39_1","doi-asserted-by":"crossref","DOI":"10.21105\/joss.03139","article-title":"Performance: An R Package for Assessment, Comparison and Testing of Statistical Models","volume":"6","author":"L\u00fcdecke D.","year":"2021","journal-title":"Journal of Open Source Software"},{"issue":"1","key":"e_1_2_10_40_1","doi-asserted-by":"crossref","first-page":"166","DOI":"10.1007\/s11692-024-09625-8","article-title":"Contemporary Body Size Variation of Neotropical Rodents: Environmental and Genetic Effects","volume":"51","author":"Mart\u00ednez J. J.","year":"2024","journal-title":"Evolutionary Biology"},{"issue":"6662","key":"e_1_2_10_41_1","doi-asserted-by":"crossref","first-page":"1067","DOI":"10.1126\/science.adg6006","article-title":"Widespread Shifts in Body Size Within Populations and Assemblages","volume":"381","author":"Martins I. S.","year":"2023","journal-title":"Science"},{"key":"e_1_2_10_42_1","first-page":"1","article-title":"Environmental Change and Long\u2010Term Body Mass Declines in an Alpine Mammal","volume":"11","author":"Mason T. H.","year":"2014","journal-title":"Frontiers in Zoology"},{"issue":"4","key":"e_1_2_10_43_1","doi-asserted-by":"crossref","first-page":"291","DOI":"10.1016\/S0168-1923(97)00027-0","article-title":"Growing Degree\u2010Days: One Equation, Two Interpretations","volume":"87","author":"McMaster G. S.","year":"1997","journal-title":"Agricultural and Forest Meteorology"},{"issue":"1","key":"e_1_2_10_44_1","doi-asserted-by":"crossref","first-page":"13","DOI":"10.1007\/s00442-010-1621-5","article-title":"Geographic and Temporal Correlations of Mammalian Size Reconsidered: A Resource Rule","volume":"164","author":"McNab B. K.","year":"2010","journal-title":"Oecologia"},{"issue":"2","key":"e_1_2_10_45_1","doi-asserted-by":"crossref","first-page":"240","DOI":"10.1111\/j.1466-8238.2008.00437.x","article-title":"Global Change and Carnivore Body Size: Data Are Stasis","volume":"18","author":"Meiri S.","year":"2009","journal-title":"Global Ecology and Biogeography"},{"issue":"1","key":"e_1_2_10_46_1","doi-asserted-by":"crossref","first-page":"221","DOI":"10.1007\/BF03192489","article-title":"Maximizing Survivorship in Cold: Thermogenic Profiles of Non\u2010Hibernating Mammals","volume":"47","author":"Merritt J. F.","year":"2002","journal-title":"Acta Theriologica"},{"issue":"7","key":"e_1_2_10_47_1","doi-asserted-by":"crossref","first-page":"2455","DOI":"10.1111\/j.1365-2486.2010.02382.x","article-title":"Advancing Breeding Phenology in Response to Environmental Change in a Wild Red Deer Population","volume":"17","author":"Moyes K.","year":"2011","journal-title":"Global Change Biology"},{"issue":"7305","key":"e_1_2_10_48_1","doi-asserted-by":"crossref","first-page":"482","DOI":"10.1038\/nature09210","article-title":"Coupled Dynamics of Body Mass and Population Growth in Response to Environmental Change","volume":"466","author":"Ozgul A.","year":"2010","journal-title":"Nature"},{"issue":"5939","key":"e_1_2_10_49_1","doi-asserted-by":"crossref","first-page":"464","DOI":"10.1126\/science.1173668","article-title":"The Dynamics of Phenotypic Change and the Shrinking Sheep of St. Kilda","volume":"325","author":"Ozgul A.","year":"2009","journal-title":"Science"},{"key":"e_1_2_10_50_1","doi-asserted-by":"crossref","first-page":"37","DOI":"10.1038\/nature01286","article-title":"A Globally Coherent Fingerprint of Climate Change Impacts Across Natural Systems","volume":"421","author":"Parmesan C.","year":"2003","journal-title":"Nature"},{"issue":"5","key":"e_1_2_10_51_1","doi-asserted-by":"crossref","first-page":"260","DOI":"10.1890\/120222","article-title":"The Subnivium: A Deteriorating Seasonal Refugium","volume":"11","author":"Pauli J. N.","year":"2013","journal-title":"Frontiers in Ecology and the Environment"},{"issue":"6332","key":"e_1_2_10_52_1","doi-asserted-by":"crossref","DOI":"10.1126\/science.aai9214","article-title":"Biodiversity Redistribution Under Climate Change: Impacts on Ecosystems and Human Well\u2010Being","volume":"355","author":"Pecl G. T.","year":"2017","journal-title":"Science"},{"key":"e_1_2_10_53_1","first-page":"126","volume-title":"Yukon Climate Change Indicators and Key Findings 2022","author":"Perrin A.","year":"2022"},{"issue":"1","key":"e_1_2_10_54_1","doi-asserted-by":"crossref","first-page":"60","DOI":"10.1038\/scientificamerican0160-60","article-title":"Animals in the Snow","volume":"202","author":"Pruitt W. O.","year":"1960","journal-title":"Scientific American"},{"key":"e_1_2_10_55_1","volume-title":"R: A Language and Environment for Statistical Computing","author":"R Core Team","year":"2024"},{"key":"e_1_2_10_56_1","doi-asserted-by":"crossref","first-page":"526","DOI":"10.1038\/sj.hdy.6885430","article-title":"Heritability of Body Mass Varies With Age and Season in Wild Bighorn Sheep","volume":"83","author":"R\u00e9ale D.","year":"1999","journal-title":"Heredity"},{"issue":"1515","key":"e_1_2_10_57_1","doi-asserted-by":"crossref","first-page":"591","DOI":"10.1098\/rspb.2002.2224","article-title":"Genetic and Plastic Responses of a Northern Mammal to Climate Change","volume":"270","author":"R\u00e9ale D.","year":"2003","journal-title":"Proceedings of the Royal Society of London, Series B: Biological Sciences"},{"issue":"3","key":"e_1_2_10_58_1","doi-asserted-by":"crossref","first-page":"680","DOI":"10.1093\/jmammal\/gyac004","article-title":"Small Mammal Dynamics in Snow\u2010Covered Forests","volume":"103","author":"Scott A. M.","year":"2022","journal-title":"Journal of Mammalogy"},{"issue":"50","key":"e_1_2_10_59_1","doi-asserted-by":"crossref","DOI":"10.1073\/pnas.2310855120","article-title":"Melting Climates Shrink North American Small Mammals","volume":"120","author":"Searing K. B.","year":"2023","journal-title":"Proceedings of the National Academy of Sciences of the United States of America"},{"issue":"2029","key":"e_1_2_10_60_1","article-title":"Ecological Consequences of Body Size Reduction Under Warming","volume":"291","author":"Sentis A.","year":"2024","journal-title":"Proceedings of the Biological Sciences"},{"issue":"8","key":"e_1_2_10_61_1","doi-asserted-by":"crossref","first-page":"401","DOI":"10.1038\/nclimate1259","article-title":"Shrinking Body Size as an Ecological Response to Climate Change","volume":"1","author":"Sheridan J. 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