{"status":"ok","message-type":"work","message-version":"1.0.0","message":{"indexed":{"date-parts":[[2026,1,29]],"date-time":"2026-01-29T14:12:06Z","timestamp":1769695926293,"version":"3.49.0"},"reference-count":107,"publisher":"Wiley","license":[{"start":{"date-parts":[[2026,1,28]],"date-time":"2026-01-28T00:00:00Z","timestamp":1769558400000},"content-version":"vor","delay-in-days":0,"URL":"http:\/\/creativecommons.org\/licenses\/by-nc-nd\/4.0\/"},{"start":{"date-parts":[[2026,1,28]],"date-time":"2026-01-28T00:00:00Z","timestamp":1769558400000},"content-version":"tdm","delay-in-days":0,"URL":"http:\/\/doi.wiley.com\/10.1002\/tdm_license_1.1"}],"funder":[{"DOI":"10.13039\/501100004488","name":"Hrvatska Zaklada za Znanost","doi-asserted-by":"publisher","award":["HRZZ\u2010IP\u20102022\u201010\u20105975"],"award-info":[{"award-number":["HRZZ\u2010IP\u20102022\u201010\u20105975"]}],"id":[{"id":"10.13039\/501100004488","id-type":"DOI","asserted-by":"publisher"}]},{"DOI":"10.13039\/501100004488","name":"Hrvatska Zaklada za Znanost","doi-asserted-by":"publisher","award":["HRZZ\u2010IP\u20102024\u201005\u20107157"],"award-info":[{"award-number":["HRZZ\u2010IP\u20102024\u201005\u20107157"]}],"id":[{"id":"10.13039\/501100004488","id-type":"DOI","asserted-by":"publisher"}]}],"content-domain":{"domain":["onlinelibrary.wiley.com"],"crossmark-restriction":true},"short-container-title":["Journal of Anatomy"],"abstract":"<jats:title>Abstract<\/jats:title>\n                  <jats:p>Early development of the human pallium is shaped by transient cellular compartments that regulate the emergence of cytoarchitectonically distinct cortical types. At these stages, regional identity depends less on mature cortical layers\u2014which form later in gestation\u2014and more on the organization of transient compartments spanning from the pia to the ventricular surface. These early compartments arise within separate pallial sectors, including the medial pallium that produces the archicortex and mesocortex and the dorsal pallium that expands into the neocortex. Although recent studies have described molecular and cellular features within individual compartments, the spatiotemporal relationships among them that govern cortex\u2010type specification remain insufficiently understood. We analyzed the distribution of proliferative, neuronal, fibrillar, and synaptic markers across the ventricular zone (VZ), subventricular zone (SVZ), cortical plate (CP), subplate (SP), and marginal zone (MZ) in prospective neocortical, mesocortical, and archicortical regions of the human fetal pallium. Development was examined across key phases: late preplate (7.5 PCW), initial CP formation (8 PCW), first CP condensation (9.5\u201312 PCW), SP formation (13\u201314 PCW), and the onset of typical fetal lamination (15 PCW). During the preplate phase, an expanded plexiform MZ containing the earliest TBR2+ and TBR1+ cells, along with tangential medial fibers, delineates the emerging archicortical (hippocampal) and mesocortical (entorhinal) sectors, which occupy much of the interhemispheric pallium. In contrast, the dorsal isocortical pallium shows a broad influx of TBR1+ neurons through abventricular compartments, while in the hippocampal primordium, these cells remain confined to the superficial zone. By 8 PCW, the first prospective pyramidal neurons form a disk\u2010shaped CP in the midlateral neopallium, with a basal\u2010to\u2010dorsal gradient reflecting SVZ proliferative activity. At the same time, the ventral archicortex shows a distinct pattern characterized by MZ expansion, accumulation of \u201cdormant\u201d TBR2+ cells, absence of a defined CP, and progressive merging of the SVZ and MZ. Between 9.5 and 12 PCW, with the onset of the second proliferative wave, a CP emerges in the ventral hippocampal anlage. The future entorhinal mesocortex differentiates unusually early, displaying large multipolar \u201cpromoter\u201d neurons in the superficial CP and the first appearance of a fibrillar lamina dissecans. By 13 PCW, hippocampal fields are discernible, and in the neocortex, the major developmental event is SP formation through merging of the deep CP and presubplate. Although hippocampal CP formation initially lags behind that of the neopallium, its subsequent differentiation\u2014and that of adjacent transitional cortices\u2014is accelerated. By 15 PCW, prospective archicortical, mesocortical, and neocortical sectors are clearly recognizable. Our findings suggest that cortex\u2010type divisions arise from coordinated interactions among proliferative, migratory\u2010fibrillar, synaptic, and postmigratory compartments. Allocortical regions exhibit distinctive developmental trajectories, including MZ enlargement, tapering of SVZ and SP and a characteristic tempo of CP formation. This compartment\u2010based perspective highlights early transitional belt\u2010like zones and spatiotemporal gradients across the pallium and identifies the ventral mesocortical (prospective entorhinal) area as an early organizing hub at the interface of allocortical and transmodal connectivity.<\/jats:p>","DOI":"10.1111\/joa.70111","type":"journal-article","created":{"date-parts":[[2026,1,29]],"date-time":"2026-01-29T02:55:53Z","timestamp":1769655353000},"update-policy":"https:\/\/doi.org\/10.1002\/crossmark_policy","source":"Crossref","is-referenced-by-count":0,"title":["Orchestrated molecular changes of proliferative, migratory\u2010fibrillar, synaptic, and postmigratory compartments align with precocious cortex\u2010type specification in the early human pallium"],"prefix":"10.1111","author":[{"given":"Janja","family":"Kopi\u0107","sequence":"first","affiliation":[{"name":"Croatian Institute for Brain Research, School of Medicine University of Zagreb  Zagreb Croatia"}],"role":[{"role":"author","vocabulary":"crossref"}]},{"given":"Fran","family":"Bo\u017ei\u0107","sequence":"additional","affiliation":[{"name":"Croatian Institute for Brain Research, School of Medicine University of Zagreb  Zagreb Croatia"}],"role":[{"role":"author","vocabulary":"crossref"}]},{"given":"Alisa","family":"Junakovi\u0107 Munjas","sequence":"additional","affiliation":[{"name":"University Department of Neurology, University Hospital Centre \u201cSestre Milosrdnice\u201d  Zagreb Croatia"}],"role":[{"role":"author","vocabulary":"crossref"}]},{"given":"\u017deljka","family":"Krsnik","sequence":"additional","affiliation":[{"name":"Croatian Institute for Brain Research, School of Medicine University of Zagreb  Zagreb Croatia"}],"role":[{"role":"author","vocabulary":"crossref"}]},{"given":"Ivica","family":"Kostovi\u0107","sequence":"additional","affiliation":[{"name":"Croatian Institute for Brain Research, School of Medicine University of Zagreb  Zagreb Croatia"}],"role":[{"role":"author","vocabulary":"crossref"}]}],"member":"311","published-online":{"date-parts":[[2026,1,28]]},"reference":[{"key":"e_1_2_9_2_1","doi-asserted-by":"publisher","DOI":"10.1515\/REVNEURO.2005.16.4.359"},{"key":"e_1_2_9_3_1","doi-asserted-by":"publisher","DOI":"10.1007\/s00429\u2010016\u20101343\u20105"},{"key":"e_1_2_9_4_1","doi-asserted-by":"publisher","DOI":"10.1093\/cercor\/bhx185"},{"key":"e_1_2_9_5_1","doi-asserted-by":"publisher","DOI":"10.1016\/0306\u20104522(89)90424\u20107"},{"key":"e_1_2_9_6_1","doi-asserted-by":"publisher","DOI":"10.1016\/j.neuron.2004.12.036"},{"key":"e_1_2_9_7_1","doi-asserted-by":"publisher","DOI":"10.1002\/(SICI)1096-9861(19960401)367:2<274::AID-CNE9>3.0.CO;2-2"},{"key":"e_1_2_9_8_1","doi-asserted-by":"publisher","DOI":"10.1111\/j.1460\u20109568.2008.06475.x"},{"key":"e_1_2_9_9_1","volume-title":"Neocortical development","author":"Bayer S.A.","year":"1991"},{"key":"e_1_2_9_10_1","doi-asserted-by":"publisher","DOI":"10.1073\/pnas.100228510711"},{"key":"e_1_2_9_11_1","doi-asserted-by":"publisher","DOI":"10.1002\/hipo.450030305"},{"key":"e_1_2_9_12_1","doi-asserted-by":"publisher","DOI":"10.3389\/fcell.2022.976549"},{"issue":"3","key":"e_1_2_9_13_1","doi-asserted-by":"crossref","first-page":"407","DOI":"10.1007\/BF00306883","article-title":"Zur Pigmentarchitektonik der Gro\u00dfhirnrinde des Menschen","volume":"127","author":"Braak H.","year":"1972","journal-title":"Zeitschrift f\u00fcr Zellforschung Und Mikroskopische Anatomie"},{"key":"e_1_2_9_14_1","doi-asserted-by":"publisher","DOI":"10.1007\/BF00690836"},{"key":"e_1_2_9_15_1","doi-asserted-by":"publisher","DOI":"10.1016\/0168\u20100102(92)90014\u20104"},{"key":"e_1_2_9_16_1","doi-asserted-by":"publisher","DOI":"10.1038\/nrn2252"},{"key":"e_1_2_9_17_1","doi-asserted-by":"publisher","DOI":"10.1016\/j.neuron.2019.07.009"},{"key":"e_1_2_9_18_1","first-page":"11145","article-title":"Layer\u2010specific programs of development in neocortical","volume":"92","author":"Clasca F.","year":"1995","journal-title":"Science"},{"key":"e_1_2_9_19_1","doi-asserted-by":"publisher","DOI":"10.1016\/j.semcdb.2017.08.033"},{"key":"e_1_2_9_20_1","doi-asserted-by":"publisher","DOI":"10.1016\/j.neuron.2014.12.060"},{"key":"e_1_2_9_21_1","doi-asserted-by":"publisher","DOI":"10.1126\/science.aai8178"},{"key":"e_1_2_9_22_1","doi-asserted-by":"publisher","DOI":"10.1093\/cercor\/9.6.586"},{"key":"e_1_2_9_23_1","doi-asserted-by":"publisher","DOI":"10.1073\/pnas.1610078113"},{"key":"e_1_2_9_24_1","doi-asserted-by":"publisher","DOI":"10.1002\/cne.902230102"},{"key":"e_1_2_9_25_1","doi-asserted-by":"publisher","DOI":"10.1523\/JNEUROSCI.08\u201008\u201002729.1988"},{"key":"e_1_2_9_26_1","doi-asserted-by":"publisher","DOI":"10.1523\/JNEUROSCI.2899\u201004.2005"},{"issue":"3","key":"e_1_2_9_27_1","doi-asserted-by":"crossref","first-page":"296","DOI":"10.1001\/archneurpsyc.1947.02300320047002","article-title":"A rational subdivision of the cerebral cortex","volume":"58","author":"Filimonoff I.N.","year":"1947","journal-title":"Archives of Neurology and Psychiatry"},{"key":"e_1_2_9_28_1","doi-asserted-by":"publisher","DOI":"10.1126\/science.1064252"},{"key":"e_1_2_9_29_1","doi-asserted-by":"publisher","DOI":"10.1146\/annurev.ne.11.030188.001033"},{"key":"e_1_2_9_30_1","doi-asserted-by":"publisher","DOI":"10.3389\/fnana.2014.00041"},{"key":"e_1_2_9_31_1","doi-asserted-by":"publisher","DOI":"10.1093\/cercor\/9.6.551"},{"key":"e_1_2_9_32_1","doi-asserted-by":"publisher","DOI":"10.1073\/pnas.0308600100"},{"key":"e_1_2_9_33_1","doi-asserted-by":"publisher","DOI":"10.1523\/jneurosci.20\u201015\u201005764.2000"},{"key":"e_1_2_9_34_1","doi-asserted-by":"publisher","DOI":"10.1002\/cne.23851"},{"key":"e_1_2_9_35_1","doi-asserted-by":"publisher","DOI":"10.1002\/(SICI)1096-9861(19960826)372:3<384::AID-CNE4>3.0.CO;2-Z"},{"key":"e_1_2_9_36_1","doi-asserted-by":"publisher","DOI":"10.1016\/S0896\u20106273(01)00211\u20102"},{"key":"e_1_2_9_37_1","doi-asserted-by":"publisher","DOI":"10.1002\/cne.900340104"},{"key":"e_1_2_9_38_1","volume-title":"Die Entwicklung des menschlichen Gehirns w\u00e4hrend der ersten Monate","author":"His W.","year":"1904"},{"key":"e_1_2_9_39_1","volume-title":"Beitr\u00e4ge zur Entwicklungsgeschichte des menschlichen Gehirns","author":"Hochstetter F.","year":"1919"},{"key":"e_1_2_9_40_1","doi-asserted-by":"publisher","DOI":"10.1523\/JNEUROSCI.4185\u201012.2013"},{"issue":"4","key":"e_1_2_9_41_1","first-page":"655","article-title":"The development of the human hippocampal fissure","volume":"101","author":"Humphrey T.","year":"1967","journal-title":"Journal of Anatomy"},{"key":"e_1_2_9_42_1","first-page":"711","volume-title":"The human nervous system","author":"Insausti R.","year":"2012"},{"key":"e_1_2_9_43_1","doi-asserted-by":"publisher","DOI":"10.1007\/978-3-642-03661-3_1"},{"key":"e_1_2_9_44_1","doi-asserted-by":"publisher","DOI":"10.1093\/cercor\/bhq219"},{"key":"e_1_2_9_45_1","doi-asserted-by":"publisher","DOI":"10.1007\/s00429-022-02606-7"},{"key":"e_1_2_9_46_1","doi-asserted-by":"publisher","DOI":"10.1007\/978-3-642-86247-2"},{"key":"e_1_2_9_47_1","doi-asserted-by":"publisher","DOI":"10.1038\/nature10523"},{"key":"e_1_2_9_48_1","doi-asserted-by":"publisher","DOI":"10.1016\/j.mcn.2005.08.022"},{"issue":"1","key":"e_1_2_9_49_1","doi-asserted-by":"crossref","first-page":"13","DOI":"10.1007\/s00429-024-02865-6","article-title":"Initial regional cytoarchitectonic differences in dorsal and orbitobasal human developing frontal cortex revealed by spatial transcriptomics","volume":"230","author":"Kopi\u0107 J.","year":"2024","journal-title":"Brain Structure & Function"},{"key":"e_1_2_9_50_1","doi-asserted-by":"publisher","DOI":"10.3390\/cells12020231"},{"key":"e_1_2_9_51_1","doi-asserted-by":"crossref","first-page":"381","DOI":"10.1007\/978-3-662-07815-0_16","volume-title":"Humanembryologie: lehrbuch und atlas der vorgeburtlichen entwicklung des menschen","author":"Kostovi\u0107 I.","year":"1990"},{"key":"e_1_2_9_52_1","doi-asserted-by":"publisher","DOI":"10.1016\/j.pneurobio.2020.101883"},{"key":"e_1_2_9_53_1","doi-asserted-by":"publisher","DOI":"10.1007\/s00429\u2010024\u201002838\u20109"},{"key":"e_1_2_9_54_1","doi-asserted-by":"publisher","DOI":"10.1111\/joa.12920"},{"key":"e_1_2_9_55_1","doi-asserted-by":"publisher","DOI":"10.1016\/j.ijdevneu.2010.09.005"},{"key":"e_1_2_9_56_1","doi-asserted-by":"publisher","DOI":"10.1002\/hipo.450030406"},{"key":"e_1_2_9_57_1","doi-asserted-by":"publisher","DOI":"10.1002\/cne.902970309"},{"key":"e_1_2_9_58_1","doi-asserted-by":"publisher","DOI":"10.1016\/j.neuroimage.2018.12.043"},{"key":"e_1_2_9_59_1","doi-asserted-by":"publisher","DOI":"10.1016\/0306-4522(89)90357-6"},{"key":"e_1_2_9_60_1","doi-asserted-by":"publisher","DOI":"10.1371\/journal.pone.0017753"},{"key":"e_1_2_9_61_1","doi-asserted-by":"publisher","DOI":"10.1126\/science.6199842"},{"key":"e_1_2_9_62_1","doi-asserted-by":"publisher","DOI":"10.1002\/cne.900950203"},{"key":"e_1_2_9_63_1","doi-asserted-by":"publisher","DOI":"10.1016\/j.ydbio.2007.08.038"},{"key":"e_1_2_9_64_1","doi-asserted-by":"publisher","DOI":"10.1007\/BF00305396"},{"key":"e_1_2_9_65_1","doi-asserted-by":"publisher","DOI":"10.1016\/0896\u20106273(88)90134\u20101"},{"key":"e_1_2_9_66_1","doi-asserted-by":"publisher","DOI":"10.1111\/j.1469\u20107580.2010.01266.x"},{"key":"e_1_2_9_67_1","doi-asserted-by":"publisher","DOI":"10.1016\/j.semcdb.2017.08.059"},{"key":"e_1_2_9_68_1","doi-asserted-by":"publisher","DOI":"10.1523\/jneurosci.20\u201005\u201001858.2000"},{"key":"e_1_2_9_69_1","doi-asserted-by":"publisher","DOI":"10.1016\/0006-8993(73)90741-5"},{"key":"e_1_2_9_70_1","doi-asserted-by":"publisher","DOI":"10.1111\/joa.13055"},{"key":"e_1_2_9_71_1","doi-asserted-by":"publisher","DOI":"10.1101\/cshperspect.a041503"},{"key":"e_1_2_9_72_1","doi-asserted-by":"publisher","DOI":"10.1098\/rsob.210042"},{"key":"e_1_2_9_73_1","doi-asserted-by":"publisher","DOI":"10.1002\/cne.902710306"},{"key":"e_1_2_9_74_1","doi-asserted-by":"publisher","DOI":"10.1002\/cne.901960110"},{"key":"e_1_2_9_75_1","doi-asserted-by":"publisher","DOI":"10.1016\/j.neuron.2007.10.010"},{"key":"e_1_2_9_76_1","doi-asserted-by":"crossref","first-page":"187","DOI":"10.1002\/0471973084.ch21","volume-title":"The Embryonic Human Brain","author":"O'Rahilly R.","year":"2006"},{"key":"e_1_2_9_77_1","doi-asserted-by":"publisher","DOI":"10.1093\/cercor\/bhp034"},{"key":"e_1_2_9_78_1","doi-asserted-by":"publisher","DOI":"10.1093\/cercor\/bhm124"},{"key":"e_1_2_9_79_1","doi-asserted-by":"publisher","DOI":"10.1523\/JNEUROSCI.20\u201016\u201006095.2000"},{"key":"e_1_2_9_80_1","doi-asserted-by":"publisher","DOI":"10.3389\/fnana.2017.00102"},{"key":"e_1_2_9_81_1","doi-asserted-by":"publisher","DOI":"10.1002\/1096\u20109861(20000828)424:3<409::AID\u2010CNE3>3.0.CO;2\u20107"},{"key":"e_1_2_9_82_1","doi-asserted-by":"publisher","DOI":"10.1038\/s41586-025-09010-1"},{"key":"e_1_2_9_83_1","doi-asserted-by":"publisher","DOI":"10.1126\/science.3291116"},{"key":"e_1_2_9_84_1","doi-asserted-by":"publisher","DOI":"10.1016\/j.tins.2009.01.007"},{"key":"e_1_2_9_85_1","first-page":"1","article-title":"Der Allocortex bei Tier und Mensch","volume":"34","author":"Rose M.","year":"1927","journal-title":"Journal f\u00fcr Psychologie Und Neurologie"},{"key":"e_1_2_9_86_1","doi-asserted-by":"publisher","DOI":"10.1093\/cercor\/9.6.521"},{"key":"e_1_2_9_87_1","doi-asserted-by":"publisher","DOI":"10.1002\/cne.24822"},{"key":"e_1_2_9_88_1","doi-asserted-by":"publisher","DOI":"10.1038\/nmeth.2019"},{"key":"e_1_2_9_89_1","doi-asserted-by":"publisher","DOI":"10.1016\/S0079\u20106123(07)63002\u20107"},{"key":"e_1_2_9_90_1","doi-asserted-by":"publisher","DOI":"10.1016\/0006\u20108993(73)90617\u20103"},{"key":"e_1_2_9_91_1","first-page":"3","volume-title":"Histology and histopathology of the nervous system","author":"Sidman R.L.","year":"1982"},{"key":"e_1_2_9_92_1","unstructured":"\u0160imi\u0107 G. Bjelo\u0161 M. Ra\u0161in M.R.et\u00a0al. (2001)Postnatal development of human CA1 pyramidal neurons. In:Abstracts of Annual Meeting of Society for Neuroscience. San Diego: Society for Neuroscience Program no. 250.\u2010x."},{"key":"e_1_2_9_93_1","doi-asserted-by":"publisher","DOI":"10.1002\/cne.25344"},{"key":"e_1_2_9_94_1","doi-asserted-by":"publisher","DOI":"10.1002\/cne.901290105"},{"key":"e_1_2_9_95_1","doi-asserted-by":"publisher","DOI":"10.1002\/cne.901310402"},{"key":"e_1_2_9_96_1","doi-asserted-by":"crossref","first-page":"213","DOI":"10.1007\/978-3-642-80890-6_8","volume-title":"Allocortex","author":"Stephan H.","year":"1975"},{"key":"e_1_2_9_97_1","doi-asserted-by":"publisher","DOI":"10.1038\/ncomms14167"},{"key":"e_1_2_9_98_1","doi-asserted-by":"publisher","DOI":"10.1016\/j.neuron.2021.06.016"},{"key":"e_1_2_9_99_1","doi-asserted-by":"publisher","DOI":"10.1038\/nn.4348"},{"key":"e_1_2_9_100_1","doi-asserted-by":"crossref","unstructured":"Tsyporin J. Zhang M. Qi C.et\u00a0al. (2025)\u201cCompeting Programs Shape Cortical Sensorimotor\u2010Association Axis Development\u201d.bioRxiv 2025\u201006.https:\/\/doi.org\/10.1101\/2025.06.26.660775.","DOI":"10.1101\/2025.06.26.660775"},{"key":"e_1_2_9_101_1","doi-asserted-by":"publisher","DOI":"10.1016\/0006\u20108993(75)90204\u20108"},{"key":"e_1_2_9_102_1","doi-asserted-by":"publisher","DOI":"10.1111\/j.1469\u20107580.2010.01260.x"},{"key":"e_1_2_9_103_1","doi-asserted-by":"publisher","DOI":"10.1002\/cne.70006"},{"key":"e_1_2_9_104_1","doi-asserted-by":"publisher","DOI":"10.1126\/science.3137660"},{"key":"e_1_2_9_105_1","doi-asserted-by":"publisher","DOI":"10.1111\/j.1749\u20106632.2000.tb06716.x"},{"key":"e_1_2_9_106_1","doi-asserted-by":"publisher","DOI":"10.1002\/(SICI)1096-9861(19990920)412:2<241::AID-CNE5>3.0.CO;2-X"},{"issue":"6420","key":"e_1_2_9_107_1","doi-asserted-by":"crossref","DOI":"10.1126\/science.aat8077","article-title":"Spatiotemporal transcriptomic divergence across human and macaque brain development","volume":"362","author":"Zhu Y.","year":"2018","journal-title":"Science"},{"key":"e_1_2_9_108_1","doi-asserted-by":"crossref","first-page":"836","DOI":"10.1016\/B978-0-12-374236-0.10023-9","volume-title":"The human nervous system","author":"Zilles K.","year":"2012"}],"container-title":["Journal of Anatomy"],"original-title":[],"language":"en","link":[{"URL":"https:\/\/onlinelibrary.wiley.com\/doi\/pdf\/10.1111\/joa.70111","content-type":"application\/pdf","content-version":"vor","intended-application":"text-mining"},{"URL":"https:\/\/onlinelibrary.wiley.com\/doi\/full-xml\/10.1111\/joa.70111","content-type":"application\/xml","content-version":"vor","intended-application":"text-mining"},{"URL":"https:\/\/onlinelibrary.wiley.com\/doi\/pdf\/10.1111\/joa.70111","content-type":"unspecified","content-version":"vor","intended-application":"similarity-checking"}],"deposited":{"date-parts":[[2026,1,29]],"date-time":"2026-01-29T02:55:59Z","timestamp":1769655359000},"score":1,"resource":{"primary":{"URL":"https:\/\/onlinelibrary.wiley.com\/doi\/10.1111\/joa.70111"}},"subtitle":[],"short-title":[],"issued":{"date-parts":[[2026,1,28]]},"references-count":107,"alternative-id":["10.1111\/joa.70111"],"URL":"https:\/\/doi.org\/10.1111\/joa.70111","archive":["Portico"],"relation":{},"ISSN":["0021-8782","1469-7580"],"issn-type":[{"value":"0021-8782","type":"print"},{"value":"1469-7580","type":"electronic"}],"subject":[],"published":{"date-parts":[[2026,1,28]]},"assertion":[{"value":"2025-11-06","order":0,"name":"received","label":"Received","group":{"name":"publication_history","label":"Publication History"}},{"value":"2026-01-09","order":2,"name":"accepted","label":"Accepted","group":{"name":"publication_history","label":"Publication History"}},{"value":"2026-01-28","order":3,"name":"published","label":"Published","group":{"name":"publication_history","label":"Publication History"}}],"article-number":"joa.70111"}}