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Heynh. Quantitative trait locus (QTL) analysis revealed that\n GA5<\/jats:italic>\n and\n BRC1<\/jats:italic>\n , as major genes regulating plant height and branching, respectively, underlie the variation in plant architecture of the Tibet ecotype.\n <\/jats:p>\n <\/jats:list-item>\n \n \n Loss\u2010of\u2010function of GA5 disrupts gibberellin biosynthesis, leading to dwarfism, and natural variations in the\n BRC1<\/jats:italic>\n promoter reduce its expression, promoting branching. A positive feedback loop formed by\n GA5<\/jats:italic>\n and\n BRC1<\/jats:italic>\n via the DELLA\u2010SPL9 module balances plant height and branching. Additionally, alleles of\n GA5<\/jats:italic>\n and\n BRC1<\/jats:italic>\n exhibit contrasting sensitivity to UV\u2010B radiation and low temperatures, suggesting that they have experienced strong ecological selections at high altitudes.\n <\/jats:p>\n <\/jats:list-item>\n \n \n Our findings reveal that\n GA5<\/jats:italic>\n and\n BRC1<\/jats:italic>\n form a feedback regulatory module that coordinately regulates the balance between plant height and branching, thereby governing developmental adaptation to high\u2010altitude environments.\n <\/jats:p>\n <\/jats:list-item>\n <\/jats:list>\n <\/jats:p>","DOI":"10.1111\/nph.70628","type":"journal-article","created":{"date-parts":[[2025,10,2]],"date-time":"2025-10-02T05:15:31Z","timestamp":1759382131000},"page":"2964-2980","update-policy":"https:\/\/doi.org\/10.1002\/crossmark_policy","source":"Crossref","is-referenced-by-count":4,"title":["Allelic variations and interactive feedback in major regulators of plant architecture confer 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