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However, their functional roles in maize endosperm remain largely unexplored. Here, we characterize the recessive maize small\u2010kernel mutant\n                          <jats:italic>sks1<\/jats:italic>\n                          , which exhibits persistent endosperm cell hyperproliferation and compromised cell expansion during grain filling.\n                        <\/jats:p>\n                      <\/jats:list-item>\n                      <jats:list-item>\n                        <jats:p>\n                          Map\u2010based cloning reveals that\n                          <jats:italic>SKS1<\/jats:italic>\n                          encodes ZmEMF1a, a PcG protein that physically interacts with subunits of both PRC1 and PRC2. Integrated ChIP\u2010seq and RNA\u2010seq analyses were performed to investigate its epigenetic regulatory functions.\n                        <\/jats:p>\n                      <\/jats:list-item>\n                      <jats:list-item>\n                        <jats:p>\n                          ZmEMF1a orchestrates a stage\u2010specific epigenetic regulatory program: it predominantly mediates H3K27me3 deposition at 6\u2009d after pollination (DAP), while coordinately regulating the deposition of both H3K27me3 and H2Aub1 at 10 DAP. Loss of\n                          <jats:italic>ZmEMF1a<\/jats:italic>\n                          leads to ectopic hyperproliferation of differentiated endosperm tissues, specifically the basal endosperm transfer layer (BETL) and aleurone (AL), as well as elevated vitamin B content in the endosperm.\n                        <\/jats:p>\n                      <\/jats:list-item>\n                      <jats:list-item>\n                        <jats:p>Collectively, these findings establish ZmEMF1a as an epigenetic regulator that balances endosperm proliferation, cell fate specification, and nutrient accumulation through stage\u2010specific histone modifications, thereby offering promising targets for enhancing maize yield and nutritional quality.<\/jats:p>\n                      <\/jats:list-item>\n                    <\/jats:list>\n                  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