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However, the regulatory mechanisms governing these traits, particularly in woody plants, remain unclear. In this study, overexpression of a citrus\n                    <jats:italic>microRNA156<\/jats:italic>\n                    (\n                    <jats:italic>miR156<\/jats:italic>\n                    ) family member,\n                    <jats:italic>Ci\u2010miR156c<\/jats:italic>\n                    , resulted in significant phenotypic changes in citrus, including reduced plant height and delayed flowering.\n                    <jats:italic>miR156<\/jats:italic>\n                    \u2010mediated repression of\n                    <jats:italic>SQUAMOSA PROMOTER\u2010BINDING PROTEIN\u2010LIKE<\/jats:italic>\n                    (\n                    <jats:italic>SPL<\/jats:italic>\n                    ) genes is a highly conserved regulatory mechanism in plants. Yeast one\u2010hybrid and dual\u2010luciferase assays, along with other related experiments, indicated that the\n                    <jats:italic>Ci\u2010miR156c\u2010CiSPL7<\/jats:italic>\n                    module targets the bZIP transcription factor (\n                    <jats:italic>CiFD<\/jats:italic>\n                    ) to regulate citrus flowering. Additionally, the\n                    <jats:italic>Ci\u2010miR156c\u2010CiSPL6<\/jats:italic>\n                    module regulates plant height by targeting\n                    <jats:italic>GA 20\u2010oxidase 2<\/jats:italic>\n                    (\n                    <jats:italic>CiGA20ox2<\/jats:italic>\n                    ), a key gibberellin biosynthesis gene. The\n                    <jats:italic>Ci\u2010miR156c\u2010CiSPL3<\/jats:italic>\n                    module also influences plant height by regulating the KNOX family gene (\n                    <jats:italic>CiKN6<\/jats:italic>\n                    ), which further regulates\n                    <jats:italic>CiGA20ox2<\/jats:italic>\n                    expression. Overexpression of\n                    <jats:italic>CiKN6<\/jats:italic>\n                    in citrus induced dwarfism, whereas its suppression increased height in transgenic plants, reinforcing its role in plant height regulation. Exogenous gibberellin and its inhibitor treatment further confirmed that the\n                    <jats:italic>miR156<\/jats:italic>\n                    \u2010\n                    <jats:italic>SPLs<\/jats:italic>\n                    module regulates citrus plant height by inhibiting gibberellin biosynthesis. These findings highlight the role of the\n                    <jats:italic>miR156\u2010SPLs<\/jats:italic>\n                    module in controlling citrus flowering and plant height.\n                  <\/jats:p>","DOI":"10.1111\/pbi.70238","type":"journal-article","created":{"date-parts":[[2025,6,30]],"date-time":"2025-06-30T08:34:27Z","timestamp":1751272467000},"page":"4271-4289","update-policy":"https:\/\/doi.org\/10.1002\/crossmark_policy","source":"Crossref","is-referenced-by-count":14,"title":["<scp>miR156<\/scp>\n                    \u2010\n                    <scp>SPLs<\/scp>\n                    module regulates flowering and controls plant height by modulating gibberellin biosynthesis in citrus"],"prefix":"10.1111","volume":"23","author":[{"given":"Min","family":"Chen","sequence":"first","affiliation":[{"name":"National Key Laboratory for Germplasm Innovation &amp; Utilization of Horticultural Crops, College of Horticulture and Forestry Science Huazhong Agricultural 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