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To assess how domestication will shape responses to long\u2010term [CO\n                    <jats:sub>2<\/jats:sub>\n                    ] enrichment, we compared wild tetraploid wheat\n                    <jats:styled-content style=\"fixed-case\">\n                      <jats:italic>Triticum turgidum<\/jats:italic>\n                    <\/jats:styled-content>\n                    ssp.\n                    <jats:italic>dicoccoides<\/jats:italic>\n                    and its cultivated derivative\n                    <jats:styled-content style=\"fixed-case\">\n                      <jats:italic>Triticum turgidum ssp. durum<\/jats:italic>\n                    <\/jats:styled-content>\n                    . Plants were grown under ambient (a[CO\n                    <jats:sub>2<\/jats:sub>\n                    ]: ~420\u2009\u03bcmol\u2009mol\n                    <jats:sup>\u22121<\/jats:sup>\n                    ) and naturally elevated (e[CO\n                    <jats:sub>2<\/jats:sub>\n                    ]: ~1000\u2009\u03bcmol\u2009mol\n                    <jats:sup>\u22121<\/jats:sup>\n                    ) [CO\n                    <jats:sub>2<\/jats:sub>\n                    ] at the Bossoleto CO\n                    <jats:sub>2<\/jats:sub>\n                    \u2010degassing vent in central Italy. Stomatal density (SD), stomatal size (SS), stomatal kinetics, and photosynthetic parameters, the maximum Rubisco carboxylation rate (\n                    <jats:italic>V<\/jats:italic>\n                    c\n                    <jats:sub>max<\/jats:sub>\n                    ), the maximum electron transport rate (\n                    <jats:italic>J<\/jats:italic>\n                    <jats:sub>max<\/jats:sub>\n                    ), and the quantum yield of photosystem II (\u03a6\n                    <jats:sub>PSII<\/jats:sub>\n                    ), were analyzed. Growth in e[CO\n                    <jats:sub>2<\/jats:sub>\n                    ] reduced SD in both subspecies, indicating an adaptive adjustment to limit transpirational water loss. However, the two taxa differed markedly in SS and stomatal behaviour.\n                    <jats:styled-content style=\"fixed-case\">\n                      <jats:italic>Triticum dicoccoides<\/jats:italic>\n                    <\/jats:styled-content>\n                    showed a slight increase in SS and a pronounced increase in stomatal physiological responsiveness, whereas\n                    <jats:styled-content style=\"fixed-case\">\n                      <jats:italic>T. durum<\/jats:italic>\n                    <\/jats:styled-content>\n                    exhibited reduced SS and slower stomatal responses. These findings indicate distinct coordination between stomatal morphology and physiological control of gas exchange. Neither apparent\n                    <jats:italic>V<\/jats:italic>\n                    c\n                    <jats:sub>max<\/jats:sub>\n                    nor apparent\n                    <jats:italic>J<\/jats:italic>\n                    <jats:sub>max<\/jats:sub>\n                    differed significantly between treatments, suggesting limited biochemical acclimation, whereas both subspecies displayed a significant increase in \u03a6\n                    <jats:sub>PSII<\/jats:sub>\n                    under e[CO\n                    <jats:sub>2<\/jats:sub>\n                    ], implying improved photochemical efficiency. Wild\n                    <jats:styled-content style=\"fixed-case\">\n                      <jats:italic>T. dicoccoides<\/jats:italic>\n                    <\/jats:styled-content>\n                    exhibited higher stomatal responsiveness and greater WUE, while cultivated\n                    <jats:styled-content style=\"fixed-case\">\n                      <jats:italic>T. durum<\/jats:italic>\n                    <\/jats:styled-content>\n                    maintained more stable gas exchange and carbon assimilation. This suggests that the morphological and physiological traits conducive to high photosynthesis and stomatal optimization of WUE may be mutually exclusive.\n                  <\/jats:p>","DOI":"10.1111\/ppl.70946","type":"journal-article","created":{"date-parts":[[2026,5,26]],"date-time":"2026-05-26T08:01:17Z","timestamp":1779782477000},"update-policy":"https:\/\/doi.org\/10.1002\/crossmark_policy","source":"Crossref","is-referenced-by-count":0,"title":["Responses of Wild and Cultivated Tetraploid Wheat to\n                    <scp>\n                      CO\n                      <sub>2<\/sub>\n                    <\/scp>\n                    Enrichment: Stomatal Optimization of Photosynthesis and Water Use Efficiency"],"prefix":"10.1111","volume":"178","author":[{"ORCID":"https:\/\/orcid.org\/0009-0000-3577-1591","authenticated-orcid":false,"given":"Eleonore","family":"Cinti","sequence":"first","affiliation":[{"name":"Department of Agricultural, Food and Environmental Sciences 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