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The nutrient delivery to shallow basins supported productivity and, because of O\n            <jats:sub>2<\/jats:sub>\n            consumption by organic-matter respiration, anoxia development. However, several studies have shown that calcareous nannoplankton experienced a decrease during the T-OAE. Nannofossil fluxes measured in the Llanbedr (Mochras Farm) borehole, Wales, UK, were the highest prior to the negative C-isotope excursion, along with high amounts of taxa indicative of nutrient-rich environments (Biscutaceae). Such conditions attest to high productivity. Fluxes show the lowest values in the core of the event, along with a size decrease of\n            <jats:italic>Schizosphaerella<\/jats:italic>\n            and a peak in Calyculaceae. The recovery of nannofossil fluxes and\n            <jats:italic>Schizosphaerella<\/jats:italic>\n            size occurred concomitant with the return of C-isotopes to more positive values. Concomitantly, deep dwellers (\n            <jats:italic>Crepidolithus crassus<\/jats:italic>\n            ) dominated, indicating a recovery of the photic-zone productivity. These observations demonstrate that the cascade of environmental responses to the initial perturbation was more complex than previously considered. In spite of elevated nutrient delivery to epicontinental basins in the early Toarcian, carbonate and primary productions of nannoplankton were depressed in the core the T-OAE, probably because of prolonged thermohaline seawater stratification.\n          <\/jats:p>","DOI":"10.1144\/sp514-2021-19","type":"journal-article","created":{"date-parts":[[2021,8,6]],"date-time":"2021-08-06T09:50:14Z","timestamp":1628243414000},"page":"59-81","update-policy":"https:\/\/doi.org\/10.1144\/crossmark-policy","source":"Crossref","is-referenced-by-count":22,"title":["Primary v. carbonate production in the Toarcian, a case study from the Llanbedr (Mochras Farm) borehole, Wales"],"prefix":"10.1144","volume":"514","author":[{"given":"Alessandro","family":"Menini","sequence":"first","affiliation":[{"name":"Universit\u00e9 de Lyon, UCBL, ENSL, UJM, CNRS, LGL-TPE, F-69622, Villeurbanne, France"}]},{"ORCID":"https:\/\/orcid.org\/0000-0003-0990-1641","authenticated-orcid":false,"given":"Emanuela","family":"Mattioli","sequence":"additional","affiliation":[{"name":"Universit\u00e9 de Lyon, UCBL, ENSL, UJM, CNRS, LGL-TPE, F-69622, Villeurbanne, France"},{"name":"Institut Universitaire de France","place":["Paris, France"]}]},{"ORCID":"https:\/\/orcid.org\/0000-0001-6178-5401","authenticated-orcid":false,"given":"Stephen P.","family":"Hesselbo","sequence":"additional","affiliation":[{"name":"Camborne School of Mines, University of Exeter, Penryn Campus, Penryn, Cornwall TR10 9FE, UK"}]},{"ORCID":"https:\/\/orcid.org\/0000-0001-8170-0399","authenticated-orcid":false,"given":"Micha","family":"Ruhl","sequence":"additional","affiliation":[{"name":"Trinity College Dublin, The University of Dublin","place":["Dublin, Ireland"]}]},{"given":"Guillaume","family":"Suan","sequence":"additional","affiliation":[{"name":"Universit\u00e9 de Lyon, UCBL, ENSL, UJM, CNRS, LGL-TPE, F-69622, Villeurbanne, France"}]}],"member":"1881","published-online":{"date-parts":[[2021,9,22]]},"reference":[{"key":"e_1_3_5_2_1","doi-asserted-by":"publisher","DOI":"10.1144\/0016-76492010-025"},{"key":"e_1_3_5_3_1","doi-asserted-by":"publisher","DOI":"10.1086\/684831"},{"key":"e_1_3_5_4_1","doi-asserted-by":"publisher","DOI":"10.1016\/j.gr.2019.06.016"},{"key":"e_1_3_5_5_1","doi-asserted-by":"publisher","DOI":"10.1016\/S0012-821X(03)00278-4"},{"key":"e_1_3_5_6_1","first-page":"63","volume-title":"Atlas Tethys Paleoenvironmental Maps","author":"Bassoullet J.P.","year":"1993","unstructured":"BassoulletJ.P., ElmiS., PoissonA., CeccaF., BellionY., GuiraudR. and BaudinF. 1993. 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