{"status":"ok","message-type":"work","message-version":"1.0.0","message":{"indexed":{"date-parts":[[2026,7,27]],"date-time":"2026-07-27T20:06:56Z","timestamp":1785182816238,"version":"3.55.0"},"reference-count":88,"publisher":"MIT Press","license":[{"start":{"date-parts":[[2026,7,2]],"date-time":"2026-07-02T00:00:00Z","timestamp":1782950400000},"content-version":"vor","delay-in-days":182,"URL":"https:\/\/creativecommons.org\/licenses\/by\/4.0\/"}],"funder":[{"DOI":"10.13039\/501100001691","name":"Japan Society for the Promotion of Science","doi-asserted-by":"publisher","award":["23KJ0477"],"award-info":[{"award-number":["23KJ0477"]}],"id":[{"id":"10.13039\/501100001691","id-type":"DOI","asserted-by":"publisher"}]}],"content-domain":{"domain":["direct.mit.edu"],"crossmark-restriction":true},"short-container-title":[],"published-print":{"date-parts":[[2026,7,27]]},"abstract":"<jats:title>Abstract<\/jats:title>\n                  <jats:p>Recent advances in neural networks trained on natural language have revealed that category-selective regions encode complex semantics and contextual information of natural scenes in addition to object categories. However, the limited interpretability of embeddings derived from these models complicates the characterization of the aspects of natural scenes that contribute to such semantic representations. Here we addressed this question by developing an analysis of the relationship between object co-occurrence in large-scale natural scene captions and corresponding fMRI responses predicted by caption-based encoding models, inspired by the fact that the joint presence of multiple objects generally shapes the overall content of a scene. We performed this analysis on the extrastriate body area (EBA), which responds strongly to human body parts. We found that human bodies co-occurring with sports-related objects drive the strongest predicted responses in the EBA among all image categories, whereas those co-occurring with vehicles or accessories elicit strong but weaker predicted responses. The findings from the co-occurrence analysis helped identify three key body-related features that contribute to the semantic representation in the EBA and fusiform body area: human body motion speed implied in static images of natural scenes is a primary contributor, and the number of people and body size are secondary contributors. Our framework, integrating object co-occurrence with caption-based encoding models, offers an interpretable approach for understanding high-level visual representations underlying natural scene perception.<\/jats:p>","DOI":"10.1162\/imag.a.1309","type":"journal-article","created":{"date-parts":[[2026,7,2]],"date-time":"2026-07-02T20:01:16Z","timestamp":1783022476000},"update-policy":"https:\/\/doi.org\/10.1162\/mitpressjournals.corrections.policy","source":"Crossref","is-referenced-by-count":0,"title":["Probing the content of semantic representations in body-selective regions"],"prefix":"10.1162","volume":"4","author":[{"given":"Ryuto","family":"Yashiro","sequence":"first","affiliation":[{"name":"Graduate School of Information Science and Technology, The University of Tokyo, Tokyo, Japan"},{"name":"Department of Education and Psychology, Freie Universit\u00e4t Berlin, Berlin, Germany"},{"name":"Institute of Cognitive Science, Universit\u00e4t Osnabr\u00fcck, Osnabr\u00fcck, Germany"}],"role":[{"vocabulary":"crossref","role":"author"}]},{"given":"Masataka","family":"Sawayama","sequence":"additional","affiliation":[{"name":"Graduate School of Information Science and Technology, The University of Tokyo, Tokyo, Japan"},{"name":"Graduate School of Information Science and Technology, Hokkaido University, Sapporo, Hokkaido, Japan"},{"name":"Prometech CG Research, Tokyo, Japan"}],"role":[{"vocabulary":"crossref","role":"author"}]},{"given":"Ayumu","family":"Yamashita","sequence":"additional","affiliation":[{"name":"Graduate School of Information Science and Technology, The University of Tokyo, Tokyo, Japan"}],"role":[{"vocabulary":"crossref","role":"author"}]},{"ORCID":"https:\/\/orcid.org\/0000-0002-4346-4791","authenticated-orcid":false,"given":"Kaoru","family":"Amano","sequence":"additional","affiliation":[{"name":"Graduate School of Information Science and Technology, The University of Tokyo, Tokyo, Japan"}],"role":[{"vocabulary":"crossref","role":"author"}]}],"member":"281","published-online":{"date-parts":[[2026,7,27]]},"reference":[{"issue":"4","key":"2026072715551303000_IMAG.a.1309-b1","doi-asserted-by":"crossref","first-page":"852","DOI":"10.1523\/JNEUROSCI.1378-19.2019","article-title":"The representation of two-body shapes in the human visual cortex","volume":"40","author":"Abassi","year":"2020","journal-title":"The Journal of Neuroscience"},{"issue":"1","key":"2026072715551303000_IMAG.a.1309-b2","doi-asserted-by":"crossref","first-page":"5531","DOI":"10.1038\/s41467-024-49073-8","article-title":"Occipital-temporal cortical tuning to semantic and affective features of natural images predicts associated behavioral responses","volume":"15","author":"Abdel-Ghaffar","year":"2024","journal-title":"Nature Communications"},{"issue":"1","key":"2026072715551303000_IMAG.a.1309-b3","doi-asserted-by":"crossref","first-page":"116","DOI":"10.1038\/s41593-021-00962-x","article-title":"A massive 7T fMRI dataset to bridge cognitive neuroscience and artificial intelligence","volume":"25","author":"Allen","year":"2022","journal-title":"Nature Neuroscience"},{"issue":"1","key":"2026072715551303000_IMAG.a.1309-b4","doi-asserted-by":"crossref","first-page":"1774","DOI":"10.1038\/s41467-018-04126-7","article-title":"Representation of multiple objects in macaque category-selective areas","volume":"9","author":"Bao","year":"2018","journal-title":"Nature Communications"},{"issue":"6439","key":"2026072715551303000_IMAG.a.1309-b5","doi-asserted-by":"crossref","first-page":"eaav9436","DOI":"10.1126\/science.aav9436","article-title":"Neural population control via deep image synthesis","volume":"364","author":"Bashivan","year":"2019","journal-title":"Science"},{"issue":"1","key":"2026072715551303000_IMAG.a.1309-b6","doi-asserted-by":"crossref","first-page":"4081","DOI":"10.1038\/s41467-021-24368-2","article-title":"Object representations in the human brain reflect the co-occurrence statistics of vision and language","volume":"12","author":"Bonner","year":"2021","journal-title":"Nature Communications"},{"issue":"5","key":"2026072715551303000_IMAG.a.1309-b7","doi-asserted-by":"crossref","first-page":"1615","DOI":"10.1002\/hbm.20626","article-title":"The Fusiform Face Area responds automatically to statistical regularities optimal for face categorization","volume":"30","author":"Caldara","year":"2009","journal-title":"Human Brain Mapping"},{"key":"2026072715551303000_IMAG.a.1309-b8","unstructured":"Chen, X., Fang, H., Lin, T.-Y., Vedantam, R., Gupta, S., Dollar, P., & Zitnick, C. 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Microsoft COCO captions: Data collection and evaluation server. arXiv. http:\/\/arxiv.org\/abs\/1504.00325"},{"issue":"1","key":"2026072715551303000_IMAG.a.1309-b9","doi-asserted-by":"crossref","first-page":"319","DOI":"10.1146\/annurev.neuro.22.1.319","article-title":"Space and attention in parietal cortex","volume":"22","author":"Colby","year":"1999","journal-title":"Annual Review of Neuroscience"},{"issue":"1","key":"2026072715551303000_IMAG.a.1309-b10","doi-asserted-by":"crossref","first-page":"9383","DOI":"10.1038\/s41467-024-53147-y","article-title":"A large-scale examination of inductive biases shaping high-level visual representation in brains and machines","volume":"15","author":"Conwell","year":"2024","journal-title":"Nature Communications"},{"issue":"5237","key":"2026072715551303000_IMAG.a.1309-b11","doi-asserted-by":"crossref","first-page":"802","DOI":"10.1126\/science.270.5237.802","article-title":"Superior parietal cortex activation during spatial attention shifts and visual feature conjunction","volume":"270","author":"Corbetta","year":"1995","journal-title":"Science (New York, N.Y.)"},{"issue":"5667","key":"2026072715551303000_IMAG.a.1309-b12","doi-asserted-by":"crossref","first-page":"115","DOI":"10.1126\/science.1093110","article-title":"Contextually evoked object-specific responses in human visual cortex","volume":"304","author":"Cox","year":"2004","journal-title":"Science"},{"issue":"9","key":"2026072715551303000_IMAG.a.1309-b13","doi-asserted-by":"crossref","first-page":"744","DOI":"10.1016\/j.tics.2021.05.010","article-title":"A computational neuroethology perspective on body and expression perception","volume":"25","author":"de Gelder","year":"2021","journal-title":"Trends in Cognitive Sciences"},{"issue":"27","key":"2026072715551303000_IMAG.a.1309-b14","doi-asserted-by":"crossref","first-page":"6539","DOI":"10.1523\/JNEUROSCI.3267-16.2017","article-title":"The hierarchical cortical organization of human speech processing","volume":"37","author":"de Heer","year":"2017","journal-title":"The Journal of Neuroscience: The Official Journal of the Society for Neuroscience"},{"issue":"8","key":"2026072715551303000_IMAG.a.1309-b15","doi-asserted-by":"crossref","first-page":"1220","DOI":"10.1038\/s42256-025-01072-0","article-title":"High-level visual representations in the human brain are aligned with large language models","volume":"7","author":"Doerig","year":"2025","journal-title":"Nature Machine Intelligence"},{"key":"2026072715551303000_IMAG.a.1309-b16","doi-asserted-by":"crossref","first-page":"e75726","DOI":"10.7554\/eLife.75726","article-title":"The normalization model predicts responses in the human visual cortex during object-based attention","volume":"12","author":"Doostani","year":"2023","journal-title":"eLife"},{"key":"2026072715551303000_IMAG.a.1309-b17","unstructured":"Dosovitskiy, A., Beyer, L., Kolesnikov, A., Weissenborn, D., Zhai, X., Unterthiner, T., Dehghani, M., Minderer, M., Heigold, G., Gelly, S., Uszkoreit, J., & Houlsby, N. (2020). An image is worth 16x16 words: Transformers for image recognition at scale. arXiv. http:\/\/arxiv.org\/abs\/2010.11929"},{"issue":"5539","key":"2026072715551303000_IMAG.a.1309-b18","doi-asserted-by":"crossref","first-page":"2470","DOI":"10.1126\/science.1063414","article-title":"A cortical area selective for visual processing of the human body","volume":"293","author":"Downing","year":"2001","journal-title":"Science (New York, N.Y.)"},{"issue":"3\u20134","key":"2026072715551303000_IMAG.a.1309-b19","doi-asserted-by":"crossref","first-page":"186","DOI":"10.1080\/17588928.2011.582945","article-title":"The role of occipitotemporal body-selective regions in person perception","volume":"2","author":"Downing","year":"2011","journal-title":"Cognitive Neuroscience"},{"key":"2026072715551303000_IMAG.a.1309-b20","doi-asserted-by":"crossref","first-page":"412","DOI":"10.1016\/j.neuroimage.2012.10.060","article-title":"The overlap of the EBA and the MT\/V5 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area?","volume":"23","author":"Hanson","year":"2004","journal-title":"NeuroImage"},{"issue":"4","key":"2026072715551303000_IMAG.a.1309-b30","doi-asserted-by":"crossref","first-page":"947","DOI":"10.1093\/cercor\/bhs091","article-title":"Deconstructing visual scenes in cortex: Gradients of object and spatial layout information","volume":"23","author":"Harel","year":"2013","journal-title":"Cerebral Cortex"},{"issue":"5539","key":"2026072715551303000_IMAG.a.1309-b31","doi-asserted-by":"crossref","first-page":"2425","DOI":"10.1126\/science.1063736","article-title":"Distributed and overlapping representations of faces and objects in ventral temporal cortex","volume":"293","author":"Haxby","year":"2001","journal-title":"Science"},{"key":"2026072715551303000_IMAG.a.1309-b32","doi-asserted-by":"crossref","first-page":"e82580","DOI":"10.7554\/eLife.82580","article-title":"THINGS-data, a multimodal collection of large-scale datasets for investigating object representations in human brain and 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C., Lo, W.-Y., Doll\u00e1r, P., & Girshick, R. (2023). Segment anything. arXiv. http:\/\/arxiv.org\/abs\/2304.02643","DOI":"10.1109\/ICCV51070.2023.00371"},{"issue":"39","key":"2026072715551303000_IMAG.a.1309-b40","doi-asserted-by":"crossref","first-page":"7545","DOI":"10.1523\/JNEUROSCI.0446-20.2020","article-title":"The functional organization of High-Level visual cortex determines the representation of complex visual stimuli","volume":"40","author":"Kliger","year":"2020","journal-title":"The Journal of Neuroscience: The Official Journal of the Society for Neuroscience"},{"issue":"2","key":"2026072715551303000_IMAG.a.1309-b41","doi-asserted-by":"crossref","first-page":"47","DOI":"10.1016\/j.tics.2003.12.001","article-title":"But still, it moves","volume":"8","author":"Kourtzi","year":"2004","journal-title":"Trends in Cognitive Sciences"},{"issue":"1","key":"2026072715551303000_IMAG.a.1309-b42","doi-asserted-by":"crossref","first-page":"48","DOI":"10.1162\/08989290051137594","article-title":"Activation in human MT\/MST by static images with implied motion","volume":"12","author":"Kourtzi","year":"2000","journal-title":"Journal of Cognitive Neuroscience"},{"issue":"1","key":"2026072715551303000_IMAG.a.1309-b43","doi-asserted-by":"crossref","first-page":"26","DOI":"10.1016\/j.tics.2012.10.011","article-title":"The ventral visual pathway: An expanded neural framework for the processing of object quality","volume":"17","author":"Kravitz","year":"2013","journal-title":"Trends in Cognitive Sciences"},{"key":"2026072715551303000_IMAG.a.1309-b44","doi-asserted-by":"crossref","first-page":"167","DOI":"10.1016\/j.conb.2019.04.002","article-title":"Interpreting encoding and decoding models","volume":"55","author":"Kriegeskorte","year":"2019","journal-title":"Current Opinion in Neurobiology"},{"issue":"6","key":"2026072715551303000_IMAG.a.1309-b45","doi-asserted-by":"crossref","first-page":"84","DOI":"10.1145\/3065386","article-title":"ImageNet classification with deep convolutional neural networks","volume":"60","author":"Krizhevsky","year":"2017","journal-title":"Communications of the ACM"},{"issue":"1","key":"2026072715551303000_IMAG.a.1309-b46","doi-asserted-by":"crossref","first-page":"6241","DOI":"10.1038\/s41467-024-50310-3","article-title":"Modeling short visual events through the BOLD moments video fMRI dataset and metadata","volume":"15","author":"Lahner","year":"2024","journal-title":"Nature Communications"},{"key":"2026072715551303000_IMAG.a.1309-b47","doi-asserted-by":"crossref","first-page":"135","DOI":"10.3389\/fncom.2015.00135","article-title":"Fourier power, subjective distance, and object categories all provide plausible models of BOLD responses in scene-selective visual areas","volume":"9","author":"Lescroart","year":"2015","journal-title":"Frontiers in Computational Neuroscience"},{"key":"2026072715551303000_IMAG.a.1309-b48","doi-asserted-by":"crossref","unstructured":"Lin, T.-Y., Maire, M., Belongie, S., Bourdev, L., Girshick, R., Hays, J., Perona, P., Ramanan, D., Lawrence Zitnick, C., & Doll\u00e1r, P. 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The Twelfth International Conference on Learning Representations. https:\/\/openreview.net\/pdf?id=mQYHXUUTkU"},{"key":"2026072715551303000_IMAG.a.1309-b53","doi-asserted-by":"crossref","first-page":"120240","DOI":"10.1016\/j.neuroimage.2023.120240","article-title":"Voxelwise encoding models of body stimuli reveal a representational gradient from low-level visual features to postural features in occipitotemporal cortex","volume":"277","author":"Marrazzo","year":"2023","journal-title":"NeuroImage"},{"key":"2026072715551303000_IMAG.a.1309-b54","doi-asserted-by":"crossref","first-page":"118545","DOI":"10.1016\/j.neuroimage.2021.118545","article-title":"Decoding the difference between explicit and implicit body expression representation in high level visual, prefrontal and inferior parietal 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GLMsingle","volume":"11","author":"Prince","year":"2022","journal-title":"eLife"},{"issue":"4","key":"2026072715551303000_IMAG.a.1309-b67","doi-asserted-by":"crossref","first-page":"466","DOI":"10.3390\/brainsci12040466","article-title":"Features and extra-striate body area representations of diagnostic body parts in anger and fear perception","volume":"12","author":"Ren","year":"2022","journal-title":"Brain Sciences"},{"key":"2026072715551303000_IMAG.a.1309-b68","unstructured":"Ren, T., Liu, S., Zeng, A., Lin, J., Li, K., Cao, H., Chen, J., Huang, X., Chen, Y., Yan, F., Zeng, Z., Zhang, H., Li, F., Yang, J., Li, H., Jiang, Q., & Zhang, L. (2024). Grounded SAM: Assembling open-world models for diverse visual tasks. arXiv. http:\/\/arxiv.org\/abs\/2401.14159"},{"issue":"12","key":"2026072715551303000_IMAG.a.1309-b200","doi-asserted-by":"crossref","first-page":"giaa133","DOI":"10.1093\/gigascience\/giaa133","article-title":"Fractional ridge regression: A fast, interpretable reparameterization of ridge regression","volume":"9","author":"Rokem","year":"2020","journal-title":"GigaScience"},{"issue":"1","key":"2026072715551303000_IMAG.a.1309-b69","doi-asserted-by":"crossref","first-page":"16","DOI":"10.1016\/S0960-9822(99)00259-6","article-title":"The functional neuroanatomy of implicit-motion perception or representational momentum","volume":"10","author":"Senior","year":"2000","journal-title":"Current Biology: CB"},{"issue":"1\u20132","key":"2026072715551303000_IMAG.a.1309-b70","doi-asserted-by":"crossref","first-page":"81","DOI":"10.1080\/13506280143000331","article-title":"Representational momentum and the brain: An investigation into the functional necessity of V5\/MT","volume":"9","author":"Senior","year":"2002","journal-title":"Visual Cognition"},{"issue":"6","key":"2026072715551303000_IMAG.a.1309-b71","doi-asserted-by":"crossref","first-page":"e38118","DOI":"10.1371\/journal.pone.0038118","article-title":"The constructive nature of affective vision: Seeing fearful scenes activates extrastriate body area","volume":"7","author":"Sinke","year":"2012","journal-title":"PLoS One"},{"key":"2026072715551303000_IMAG.a.1309-b72","doi-asserted-by":"crossref","first-page":"408","DOI":"10.1016\/j.neuroimage.2013.06.032","article-title":"The fusiform face area responds equivalently to faces and abstract shapes in the left and central visual fields","volume":"83","author":"Slotnick","year":"2013","journal-title":"NeuroImage"},{"key":"2026072715551303000_IMAG.a.1309-b73","unstructured":"Song, K., Tan, X., Qin, T., Lu, J., & Liu, T.-Y. (2020). MPNet: Masked and permuted pre-training for language understanding. arXiv. http:\/\/arxiv.org\/abs\/2004.09297"},{"issue":"5","key":"2026072715551303000_IMAG.a.1309-b74","doi-asserted-by":"crossref","first-page":"1025","DOI":"10.1016\/j.neuron.2013.06.034","article-title":"Natural scene statistics account for the representation of scene categories in human visual cortex","volume":"79","author":"Stansbury","year":"2013","journal-title":"Neuron"},{"issue":"Pt A","key":"2026072715551303000_IMAG.a.1309-b75","doi-asserted-by":"crossref","first-page":"188","DOI":"10.1016\/j.neuroimage.2017.06.035","article-title":"The feature-weighted receptive field: An interpretable encoding model for complex feature spaces","volume":"180","author":"St-Yves","year":"2018","journal-title":"NeuroImage"},{"issue":"2","key":"2026072715551303000_IMAG.a.1309-b76","doi-asserted-by":"crossref","first-page":"214","DOI":"10.1111\/j.1440-1819.2008.01757.x","article-title":"Enhanced activation in the extrastriate body area by goal-directed actions","volume":"62","author":"Takahashi","year":"2008","journal-title":"Psychiatry and Clinical Neurosciences"},{"issue":"23","key":"2026072715551303000_IMAG.a.1309-b77","doi-asserted-by":"crossref","first-page":"2130","DOI":"10.1016\/j.cub.2004.11.031","article-title":"Magnetic stimulation of extrastriate body area impairs visual processing of nonfacial body parts","volume":"14","author":"Urgesi","year":"2004","journal-title":"Current Biology: CB"},{"issue":"2","key":"2026072715551303000_IMAG.a.1309-b78","doi-asserted-by":"crossref","first-page":"101","DOI":"10.1080\/17470910701865367","article-title":"Specific and common brain regions involved in the perception of faces and bodies and the representation of their emotional expressions","volume":"4","author":"van de Riet","year":"2009","journal-title":"Social Neuroscience"},{"issue":"3","key":"2026072715551303000_IMAG.a.1309-b79","doi-asserted-by":"crossref","first-page":"661","DOI":"10.1006\/nimg.2001.0860","article-title":"Functional specificity of superior parietal mediation of spatial shifting","volume":"14","author":"Vandenberghe","year":"2001","journal-title":"NeuroImage"},{"issue":"2","key":"2026072715551303000_IMAG.a.1309-b80","doi-asserted-by":"crossref","first-page":"574","DOI":"10.1523\/JNEUROSCI.4032-13.2014","article-title":"Distinct neural mechanisms for body form and body motion discriminations","volume":"34","author":"Vangeneugden","year":"2014","journal-title":"The Journal of Neuroscience: The Official Journal of the Society for Neuroscience"},{"issue":"12","key":"2026072715551303000_IMAG.a.1309-b81","doi-asserted-by":"crossref","first-page":"1415","DOI":"10.1038\/s42256-023-00753-y","article-title":"Better models of human high-level visual cortex emerge from natural language supervision with a large and diverse dataset","volume":"5","author":"Wang","year":"2023","journal-title":"Nature Machine Intelligence"},{"issue":"4","key":"2026072715551303000_IMAG.a.1309-b82","doi-asserted-by":"crossref","first-page":"2183","DOI":"10.1016\/j.neuroimage.2011.03.041","article-title":"Not one extrastriate body area: Using anatomical landmarks, hMT+, and visual field maps to parcellate limb-selective activations in human lateral occipitotemporal cortex","volume":"56","author":"Weiner","year":"2011","journal-title":"NeuroImage"},{"issue":"7","key":"2026072715551303000_IMAG.a.1309-b83","doi-asserted-by":"crossref","first-page":"1765","DOI":"10.1162\/jocn.2010.21552","article-title":"Representation of action in occipito-temporal cortex","volume":"23","author":"Wiggett","year":"2011","journal-title":"Journal of Cognitive Neuroscience"},{"issue":"2","key":"2026072715551303000_IMAG.a.1309-b84","doi-asserted-by":"crossref","first-page":"103","DOI":"10.1016\/j.tics.2021.10.003","article-title":"Two \u201cwhat\u201d 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