{"status":"ok","message-type":"work","message-version":"1.0.0","message":{"indexed":{"date-parts":[[2026,10,31]],"date-time":"2026-10-31T22:47:09Z","timestamp":1793486829964,"version":"4.5.0"},"reference-count":1,"publisher":"Magnolia Press","issue":"1","content-domain":{"domain":[],"crossmark-restriction":false},"short-container-title":["Zootaxa"],"abstract":"The spider genus Porrhomma has been revised; it now includes 24 species. Two new species are described: Porrhomma altaica new species from the Altai Mountains, and Porrhomma nekolai new species from eastern Asia and North America. Bathyphantes charpentieri Lebert, 1877 was removed from synonymy with Porrhomma microphthalmum (O. Pickard-Cambridge, 1871). The following new synonymies are proposed: Bathyphantes charpentieri Lebert, 1877 is a junior synonym of Porrhomma convexum (Westring, 1851) new synonymy; Porrhomma gertschi Hackman, 1954 is a junior synonym of Porrhomma terrestre (Emerton, 1882) new synonymy; Porrhomma myops Simon, 1884 is a junior synonym of Porrhomma rosenhaueri (L. Koch, 1872) new synonymy; Porrhomma ocella Chamberlin & Ivie, 1943 is a junior synonym of Porrhomma convexum (Westring, 1851) new synonymy; Porrhomma omissum Miller, 1971 is a junior synonym of Porrhomma egeria Simon, 1884 new synonymy; Porrhomma pallidum affinis Miller & Kratochv\u00edl, 1940 is a junior synonym of Porrhomma pallidum Jackson, 1913 new synonymy; Porrhomma sodonta (Chamberlin, 1949) is a junior synonym of Porrhomma convexum (Westring, 1851) new synonymy. The synonymy of Porrhomma macrochelis (Emerton, 1917) with Macrargus multesimus (O. Pickard-Cambridge, 1875) is supported. Porrhomma subterraneum Simon, 1884 is considered a nomen dubium, because it was described according to a juvenile specimen. To date, Porrhomma indecorum Simon, 1910 and Porrhomma marphaense Wunderlich, 1983 are only known from findings of females; they are considered to be species inquirenda, and their names are declared nomina dubia. Keys are provided for males and females. Vulvae are drawn in dorsal, ventral and caudal views to understand their spatial structure.\u00a0<\/jats:p>","DOI":"10.11646\/zootaxa.4481.1.1","type":"journal-article","created":{"date-parts":[[2018,9,14]],"date-time":"2018-09-14T01:09:01Z","timestamp":1536887341000},"source":"Crossref","is-referenced-by-count":9,"title":["A review of the spider genus Porrhomma (Araneae, Linyphiidae)"],"prefix":"10.11646","volume":"4481","author":[{"given":"VLASTIMIL","family":"R\u016e\u017dI\u010cKA","sequence":"first","affiliation":[],"role":[{"vocabulary":"crossref","role":"author"}]}],"member":"4913","published-online":{"date-parts":[[2018,9,14]]},"reference":[{"key":"43594","unstructured":"

Arachnologische Gesellschaft (2017) Atlas of the European Arachnids. Available from: https:\/\/atlas.arages.de (accessed 30 November 2017)<\/p>

Banks, N. (1899) Arachnida. In<\/em>: Ashmead, W.H. (Ed.), Reports upon the insects, mites and myriapods collected by Dr. L. Stejneger and Mr. G.E.H. Barrett-Hamilton on the Commander Islands. Report of Fur-Seal Investigations 1896\u201397<\/em>, 1899, pp. 347\u2013350.<\/p>

Berland, L. (1931) Biospeologica LVI. Campagne sp\u00e9ologique de C. Bolivar et R. Jeannel dans l\u2019Am\u00e9rique du Nord (1928). 7. Arachnides Aran\u00e9ides. Archives de Zoologie Exp\u00e9rimentale et G\u00e9n\u00e9rale<\/em>, 71, 383\u2013388.<\/p>

Bianchi, C., Caporiacco, L. di, Massera, M.G. & Valle, A. (1949) Raccolte faunistiche della grotta della Spipola (Bologna). Commentationes Pontificia Academia Scientiarvm<\/em>, 13, 493\u2013527.<\/p>

Bishop, S.C. & Crosby, C.R. (1938) Studies in American spiders: Miscellaneous genera of Erigoneae, Part II. Journal of the New York Entomological Society<\/em>, 46, 55\u2013107.<\/p>

Blackwall, J. (1834) Characters of some undescribed species of Araneidae. London and Edinburgh Philosophical Magazine and Journal of Science<\/em>, 5 (3), 50\u201353.
https:\/\/doi.org\/10.1080\/14786443408648400<\/p>

Blackwall, J. (1841) The difference in the number of eyes with which spiders are provided proposed as the basis of their distribution into tribes; with descriptions of newly discovered species and the characters of a new family and three new genera of spiders. Transactions of the Linnean Society of London<\/em>, 18, 601\u2013670.
https:\/\/doi.org\/10.1111\/j.1095-8339.1838.tb00210.x<\/p>

Borges, P.A.V. & Wunderlich, J. (2008) Spider biodiversity patterns and their conservation in the Azorean archipelago, with descriptions of new species. Systematics and Biodiversity<\/em>, 6, 249\u2013282.
https:\/\/doi.org\/10.1017\/S1477200008002648<\/p>

Bosmans, R. (2005) Contribution to the knowledge of the Linyphiidae of the Maghreb. Part XI. Miscellaneous Linyphiid genera and additions (Araneae: Linyphiidae: Linyphiinae). Bulletin et Annales de la Soci\u00e9t\u00e9 Royale d\u2019 Entomologie de Belgique\/Koninklijke Belgische Vereniging voor Entomologie<\/em>, 141, 125\u2013161.<\/p>

Bourne, J.D. (1977a) A contribution to the study of the genus Porrhomma<\/em> (Araneae: Linyphidae). Notes on a population of P. egeria<\/em> (Simon) and other cavernicolous species. International Journal of Speleology<\/em>, 9, 89\u201396.<\/p>

Bourne, J.D. (1977b) Contribution a \u013e\u00e9tude du genre Porrhomma<\/em> (Araneae, Linyphiidae). Caract\u00e8res morphologiques, biom\u00e9trie et \u00e9cologie au niveau des populations de P. convexum<\/em> (Westring) et P. myops<\/em> (Simon). Mitteilungen der Schweizerischen entomologischen Gesellschaft<\/em>, 50, 153\u2013165.<\/p>

Breuss, W. (2009) A collection of spiders and harvestmen from two caves in Ontario and Newfounland, Canada (Araneae, Opiliones). Contributions to Natural History<\/em>, 12, 297\u2013313.<\/p>

Brignoli, P.M. (1979) Ragni d\u2019Italia XXXI. Specie cavernicole nuove o interessanti (Araneae). Quaderni del Museo di Speleologia \u00abV. Rivera\u00bb<\/em>, 5 (10), 3\u201348.<\/p>

Buchar, J. & R\u016f\u017ei\u010dka, V. (2002) Catalogue of spiders of the Czech Republic<\/em>. Peres, Praha, 351 pp.<\/p>

Buckle, D.J., Carroll, D., Crawford, R.L. & Roth, V.D. (2001) Linyphiidae and Pimoidae of America north of Mexico: Checklist, synonymy, and literature. In<\/em>: Paquin, P. & Buckle, D.J. (Eds.), Contributions \u00e0 la connaissance des Araign\u00e9es (Araneae) d\u2019Am\u00e9rique du Nord.<\/em> Fabreries<\/em>, 10 (Suppl\u00e9ment), 89\u2013191.<\/p>

Carter, J., Fowles, A. & Angele, C. (2010) Monitoring the population of the linyphiid spider Porrhomma rosenhaueri<\/em> (L. Koch, 1872) (Araneae: Linyphiidae) in Lesser Garth Cave, Cardiff, UK. Cave and Karst Science<\/em>, 37 (1), 3\u20138.<\/p>

Chamberlin, R.V. (1949) On some American spiders of the family Erigonidae. Annals of the Entomological Society of America<\/em>, 61, 483\u2013562.<\/p>

Chamberlin, R.V. & Ivie, W. (1933) Spiders of the Raft River Mountanins of Utah. Bulletin of the University of Utah<\/em>, 23 (4), 1\u201379.<\/p>

Chamberlin, R.V. & Ivie, W. (1943) New genera and species of North American Linyphiid spiders. Bulletin of the University of Utah<\/em>, 33 (10), 1\u201339.<\/p>

Chamberlin, R.V. & Ivie, W. (1947) The spiders of Alaska. Bulletin of the University of Utah<\/em>, 37 (10), 1\u2013103.<\/p>

Charitonov, D.E. (1928) Versuch eines Katalogs der russischen Spinnen. (Autoreferat). Izvestia Biologitscheskogo Nautschno-Issledovatelskogo Instituta pri Permskom Gosudarstvennom Universitete<\/em>, 6, 55\u201356. [in Russian, German summary]<\/p>

Coddington, J.A. (1990) Ontogeny and homology in the male palpus of orb-weaving spiders and their relatives, with comments on phylogeny (Araneoclada: Araneoidea, Deinopoidea). Smithsonian Contributions to Zoology<\/em>, 496, 1\u201352.
https:\/\/doi.org\/10.5479\/si.00810282.496<\/p>

Crosby, C.R. (1905) The spiders of the Rochport Cave, Mo. The Canadian Entomologist<\/em>, 37, 367\u2013369.
https:\/\/doi.org\/10.4039\/Ent37367-11<\/p>

Crawford, R.L. (1988) An annotated checklist of the spiders of Washington. Burke Museum Contributions in Anthropology and Natural History<\/em>, 5, 1\u201348.<\/p>

Dahl, M. (1938) Zur Verbreitung der Gattung Porrhomma<\/em> in deutschen H\u00f6hlen, Stollen, Bergwerken und Kellern un deren freilebenden Arten. Mitteilungen \u00fcber H\u00f6hlen- und Karstforschung<\/em>, 1938, 122\u2013132.<\/p>

Dahl, M. (1939) Berichtigung. Mitteilungen \u00fcber H\u00f6hlen- und Karstforschung<\/em>, 1939, 48.<\/p>

Dani\u015fman, T. & Ko\u015far, \u0130. (2013) A contribution to the knowledge of the linyphiid spider fauna of Turkey. Acta Zoologica Bulgarica<\/em>, 65 (4), 567\u2013570.<\/p>

Deltshev, C.C., \u0106ur\u010di\u0107, B.P.M. & Blagoev, G.A. (2003) The Spiders of Serbia<\/em>. Institute of Zoology, Belgrade, 834 pp.<\/p>

Demircan, N. & Top\u00e7u, A. (2016) First records for spider fauna of the European part of Turkey (Araneae). Serket<\/em>, 15, 85\u201391.<\/p>

Emerton, J.H. (1875) Notes on spiders from Caves in Kentucky, Virginia and Indiana. American Naturalist<\/em>, 9, 278\u2013281.
https:\/\/doi.org\/10.1086\/271488<\/p>

Emerton, J.H. (1882) New England spiders of the family Theridiidae. Transactions of the Connecticut Academy of Arts and Sciences<\/em>, 6, 1\u201386.
https:\/\/doi.org\/10.5962\/bhl.part.7410<\/p>

Emerton, J.H. (1917) New spiders from Canada and the adjoining states. The Canadian Entomologist<\/em>, 49, 261\u2013272.
https:\/\/doi.org\/10.4039\/Ent49261-8<\/p>

Eskov, K.Y. (1985) Spiders of the tundra zone of the USSR. In<\/em>: The fauna and ecology of spiders of the USSR. Proceedings of the Zoological Institute of the USSR Academy of Sciences<\/em>, 139, pp. 121\u2013128 [in Russian]<\/p>

Eskov, K.Y. (1988) Spiders (Aranei) of central Siberia. In<\/em>: Rogacheva, E.V. (Ed.), Materialy po faune Srednei Sibiri i prilezhaschikh raionov Mongolii<\/em>. Academia nauk, Moscow, pp. 101\u2013155. [in Russian]<\/p>

Eskov, K.Y. & Marusik, Y.M. (1993) New data on the taxonomy and faunistics of North Asian linyphiid spiders (Aranei Linyphiidae). Arthropoda Selecta<\/em>, 2 (4), 41\u201379.<\/p>

Esyunin, S.L. (2007) Remarks on the Ural spider (Arachnida, Aranei) fauna, 13. New records of linyphiid spider (Linyphiidae). Arthropoda Selecta<\/em>, 16 (3), 169\u2013172.<\/p>

Fage, L. (1931) Biospeologica LV. Araneae. Cinqui\u00e9me s\u00e9rie, pr\u00e9c\u00e9d\u00e9e d\u2019un essai sur l\u2019\u00e9volution souterraine et son d\u00e9terminisme. Archives de Zoologie Exp\u00e9rimentale et G\u00e9n\u00e9rale<\/em>, 71, 91\u2013291.<\/p>

Fage, L. (1933) Les arachnides cavernicoles de Belgique. Bulletin de la Soci\u00e9t\u00e9 entomologique de France<\/em>, 38, 53\u201356.<\/p>

Fickert, C. (1875) Myriopoden und Araneiden vom Kamme des Riesengebirges<\/em>. Thesis, Koeniglichen Universitaet, Wroc\u0142aw, 49 pp.<\/p>

Fuhn, I.E. & Oltean, C. (1970) Lista areneelor din r. s. Rom\u00e2nia. Studii \u015fi Comunic\u0103ri<\/em>, 1970, 157\u2013196.<\/p>

Gnelitsa, V.A. & Koponen, S. (2010) A new species of the genus Macrargus<\/em> (Araneae, Linyphiidae, Micronetinae) from the north-east of Ukraine and redescription of two related species. Vestnik Zoologii<\/em>, 44, 291\u2013299.
https:\/\/doi.org\/10.2478\/v10058-010-0019-5<\/p>

Hackman, W. (1954) The spiders of Newfounland. Acta Zoologica Fennica<\/em>, 79, 1\u201399.<\/p>

Harvey, P.R., Nellist, D.R. & Telfer, M.G. (2002) Provisional atlas of British spiders (Arachnida, Araneae). Vols. 1 & 2<\/em>. Biological Records Centre, Huntingdon, 406 pp.<\/p>

Helsdingen, P.J. van (1965) Sexual behaviour of Lepthyphantes leprosus<\/em> (Ohlert) (Araneida, Linyphiidae) with notes on the function of the genital organs. Zoologische Mededelingen<\/em>, 41 (2), 15\u201344.<\/p>

Helsdingen, P.J. van (1986) The Porrhomma microphthalmum<\/em> species-group. Bulletin of the British Arachnological Society<\/em>, 7, 11\u201316.<\/p>

Helsdingen, P.J. van & IJland, S. (2011) Porrhomma microcavense<\/em> Wunderlich, 1990 (Araneae, Linyphiidae) new for the Netherlands. Nieuwsbrief Spined<\/em>, 31, 23\u201326.<\/p>

Holm, \u00c5. (1944) Revision einiger norwegischer Spinnenarten und Bemerkungen \u00fcber deren Vorkommen in Schweden. Entomologisk tidskrift<\/em>, 65, 122\u2013134.<\/p>

Holm, \u00c5. (1960) On a collection of spiders from Alaska. Zoologiska Bidrag fr\u00e5n Uppsala<\/em>, 33, 109\u2013134.<\/p>

Holm, \u00c5. (1970) Notes on spiders collected by the \u201cVega\u201d expedition 1878\u20131880. Entomologica scandinavica<\/em>, 1, 188\u2013208.
https:\/\/doi.org\/10.1163\/187631270X00023<\/p>

Holm, \u00c5. (1973) On the spiders collected during the Swedish expeditions to Novaya Zemlya and Yenisey in 1875 and 1876. Zoologica Scripta<\/em>, 2, 71\u2013110.
https:\/\/doi.org\/10.1111\/j.1463-6409.1974.tb00741.x<\/p>

Holm, \u00c5. (1979) A taxonomic study of European and East African species of the genera Pelecopsis<\/em> and Trichopterna<\/em> (Araneae, Linyphiidae), with descriptions of a new genus and two new species of Pelecopsis<\/em> from Kenya. Zoologica Scripta<\/em>, 8, 255\u2013278.
https:\/\/doi.org\/10.1111\/j.1463-6409.1979.tb00638.x<\/p>

Hormiga, G. (2000) Higher level phylogenetics of Erigonine spiders (Araneae, Linyphiidae, Erigoninae). Smithsonian Contributions to Zoology<\/em>, 609, 1\u2013160.
https:\/\/doi.org\/10.5479\/si.00810282.609<\/p>

Ivie, W. (1967) Some synonyms in American spiders. Journal of the New York Entomological Society<\/em>, 75 (3), 126\u2013131.<\/p>

Jackson, A.R. (1913) On some new and obscure British spiders. Transactions and Annual Report of the Nottingham Naturalists\u2019 Society<\/em>, 60, 20\u201349.<\/p>

Jackson, A.R. (1916) On some arthropods observed in 1915. I. Arachnida. The Lancashire and Cheshire Naturalist<\/em>, 8, 355\u2013364.<\/p>

J\u00e4ger, P. (2005) Lengthening of embolus and copulatory duct: a review of an evolutionary trend in the spider family Sparassidae (Araneae). In<\/em>: Deltshev, C. & Stoev, P. (Eds.), European Arachnology 2005. Acta zoologica bulgarica<\/em>, 1 (Supplementum), pp. 49\u201362.<\/p>

Kaston, B.J. (1948) Spiders of Connecticut<\/em>. Bulletin of the Connecticut State Geological and Natural History Survey<\/em>, 70, 1\u2013874.<\/p>

Keyserling, E. (1886) Die Spinnen Amerikas. 2 (2). Theridiidae.<\/em> Verlag von Bauer & Raspe (E. K\u00fcster), N\u00fcrnberg, 295 pp.<\/p>

Koch, L. (1872) Apterologisches aus dem Frankischen Jura. Abhandlungen der Naturhistorischen Gesellschaft zu N\u00fcrnberg<\/em>, 5, 127\u2013152.<\/p>

Kolosv\u00e1ry, G. & Loksa, I. (1944) VIII. Beitrag zur Spinnenfauna Siebenb\u00fcrgens. Annales Historico-Naturales Musei Nationalis Hungarici<\/em>, 1, 11\u201328.<\/p>

Kratochv\u00edl, J. (1934) Liste g\u00e9n\u00e9rale des Araign\u00e9es cavernicoles en Yougoslavie. Pregled pe\u0107inskih paukova u Jugoslaviji. Prirodoslovne razprave<\/em>, 2 (4), 165\u2013226.<\/p>

Kritscher, E. (1968) Ein Beitrag zur Kenntnis der Araneean-Fauna Italiens. Memorie del Museo Civico di Storia Naturale di Verona<\/em>, 16, 271\u2013319.<\/p>

Kubcov\u00e1, L. & Dolej\u0161, P. (2016) In memoriam Prof. RNDr. Jan Buchar, DrSc. (1932\u20132015). Arachnologische Mitteilungen<\/em>, 51, iv\u2013xi.<\/p>

Kubcov\u00e1, L. & Schlaghamersk\u00fd, J. (2002) Zur Spinnenfauna der Stammregion stehenden Totholzes in s\u00fcdm\u00e4hrischen Auenw\u00e4ldern. Arachnologische Mitteilungen<\/em>, 24, 35\u201361.
https:\/\/doi.org\/10.5431\/aramit2403<\/p>

Lebert, H. (1877) Die Spinnen der Schweiz, ihr Bau, ihr Leben, ihre systematische \u00dcbersicht. Neue Denkschriften der Schweizerischen Naturforschenden Gesellschaft<\/em>, 27, 1\u2013321.<\/p>

Lessert, R. de (1910) Araign\u00e9es. Catalogue des invert\u00e9br\u00e9s de la Suisse, Fsc. 3<\/em>. Museum d\u2019Histoire Naturelle de Gen\u00e8ve, Gen\u00e8ve, 639 pp.<\/p>

Levi, H.W. (1965) Techniques for the study of spider genitalia. Psyche<\/em>, 72 (2), 152\u2013158.
https:\/\/doi.org\/10.1155\/1965\/94978<\/p>

Li, S. & Song, D. (1993) On the diagnostic characters of linyphiid spiders, with descriptions of some species (Araneae: Linyphiidae). Scientific Treatise on Systematic and Evolutionary Zoology<\/em>, 2, 247\u2013256. [in Chinese, English summary]<\/p>

Locket, G.H. & Millidge, A.F. (1953) British spiders. Vol. II.<\/em> Ray Society, London, 449 pp.<\/p>

Loksa, I. (1961) \u00d6kologisch-faunistische Untersuchungen in der Freiheith\u00f6hle bei \u00c9gersz\u00f6g. (Biospeologica Hungarica, XIII.) Acta Zoologica Academiae Scientiarum Hungaricae<\/em>, 7, 219\u2013230.<\/p>

Loksa, I. (1970) Die Spinnen der \u201eK\u00f6lyuk\u201c-H\u00f6hlen im B\u00fckkgebirge (Biospeologica Hungarica XXXIII). Annales Universitatis Scientiarum Budapestinensis de Rolando E\u00f6tv\u00f6s nominatae<\/em>, Sectio Biologica, 12, 269\u2013276.<\/p>

Marusik, Y.M. (2004) Fauna and populations of the petrophilous spiders (Arachnida: Araneae) of north-east Siberia and north-west Canada. In<\/em>: Logunov, D.V. & Penney, D. (Eds.), European Arachnology 2003<\/em>. Arthropoda Selecta<\/em>, Special Issue 1, pp. 185\u2013200.<\/p>

Marusik, Y.M, B\u00f6cher, J. & Koponen, S. (2006) The collection of Greenland spiders (Aranei) kept in the Zoological Museum, University of Copenhagen. Arthropoda Selecta<\/em>, 15 (1), 59\u201380.<\/p>

Marusik, Y.M. & Koponen, S. (2000) New data on spiders (Aranei) from the Maritime Province, Russian Far East. Arthropoda Selecta<\/em>, 9, 55\u201368.<\/p>

Marusik, Y.M. & Koponen, S. & Danilov, S.N. (2001) Taxonomic and faunistic notes on linyphiids of Transbaikalia and south Siberia (Araneae, Linyphiidae). Bulletin of the British Arachnological Society<\/em>, 12, 83\u201392.<\/p>

Merrett, P. (1994) Porrhomma cambridgei<\/em>, replacement name for Porrhomma oblongum<\/em> (O. P.-Cambridge, 1871), revalidated and redescribed from southern England (Araneae: Linyphiidae). Bulletin of the British Arachnological Society<\/em>, 9, 318\u2013320.<\/p>

Miller, F. (1971) \u0158\u00e1d Pavouci\u2014Araneida [Order Spiders\u2014Araneida]. In<\/em>: Daniel, M. & \u010cern\u00fd, V. (Eds.), Kl\u00ed\u010d zv\u00ed\u0159eny \u010cSSR IV.<\/em> [Key to the fauna of Czechoslovakia IV ]<\/em>. \u010cSAV, Praha, pp. 51\u2013306 [in Czech]<\/p>

Miller, F. & Kratochv\u00edl, J. (1940) Ein Beitrag zur Revision der mitteleurop\u00e4ischen Spinnenarten aus der Gattung Porrhomma<\/em> E. Sim. Zoologischer Anzeiger<\/em>, 130, 161\u2013190.<\/p>

Miller, F. & Obrtel, R. (1975) Soil surface spiders in a lowland forest. Acta scientiarum naturalium Academiae scientiarum bohemoslovacae, Brno<\/em>, 9 (4), 1\u201340.<\/p>

Miller, J.A. (2005) A redescription of Porrhomma cavernicola<\/em> Keyserling (Araneae, Linyphiidae) with notes on Appalachian troglobites. The Journal of Arachnology<\/em>, 33, 426\u2013438.
https:\/\/doi.org\/10.1636\/04-51.1<\/p>

Millidge, A.F. & Locket, G.H. (1952) New and rare British spiders. Proceedings of the Linnean Society of London<\/em>, 163, 59\u201378.
https:\/\/doi.org\/10.1111\/j.1095-8312.1952.tb00641.x<\/p>

Namkung, J., Yoo, J.S., Lee, S.Y., Lee, J.H., Paek, W.K. & Kim, S.T. (2009) Bibliographic check list of Korean spiders (Arachnida: Araneae) ver. 2010. Journal of Korean Nature<\/em>, 2 (3), 191\u2013285.<\/p>

Naumova, M.V., Lazarov, S.P., Petrov, B.P. & Deltshev, C.D. (2016) New faunistic data on the cave-dwelling spiders in the Balcan Peninsula (Araneae). Ecologica Montenegriana<\/em>, 7, 425\u2013438.<\/p>

Nekola, J.C. (1999) Paleorefugia and neorefugia: the influence of colonization history on community pattern and process. Ecology<\/em>, 80, 2459\u20132473.<\/p>

Ono, H., Matsuda, M. & Saito, H. (2009) Linyphiidae, Pimoidae. In<\/em>: Ono, H. (Ed.), The Spiders of Japan with keys to the families and genera and illustrations of the species<\/em>. Tokai University Press, Kanagawa, pp. 253\u2013344.<\/p>

Otto, S. (2017) Caucasian spiders. A faunistic database on the spiders of the Caucasus. Version 08.2017<\/em>. Available from: http:\/\/caucasus-spiders.info\/ (accessed 29 December 2017)<\/p>

Paquin, P. & Dup\u00e9rr\u00e9, N. (2003) Guide d\u2019identification des Araign\u00e9es (Araneae) du Qu\u00e9bec. Fabreries<\/em>, 11 (Suppl\u00e9ment), 1\u2013251.<\/p>

Pickard-Cambridge, F.O. (1894) New genera and species of British spiders. Annals and Magazine of Natural History<\/em>, Series 6, 13, 87\u2013111.
https:\/\/doi.org\/10.1080\/00222939408677672<\/p>

Pickard-Cambridge, F.O. (1895) Notes on British spiders, with descriptions of new species. Annals and Magazine of Natural History<\/em>, Series 6, 15, 25\u201341.
https:\/\/doi.org\/10.1080\/00222939508677845<\/p>

Pickard-Cambridge, O. (1871) Descriptions of some British spiders new to science; with a notice of others, of which some are now for the first time recorded as British species. Transactions of the Linnean Society of London<\/em>, 27, 393\u2013464.
https:\/\/doi.org\/10.1111\/j.1096-3642.1871.tb00218.x<\/p>

Pickard-Cambridge, O. (1875) On some new species of Erigone<\/em> from North America. Proceedings of the Zoological Society of London<\/em>, 1875, 393\u2013405.<\/p>

Pickard-Cambridge, O. (1878) Notes on British spiders with descriptions of new species. Annals and Magazine of Natural History<\/em>, Series 5, 1, 105\u2013128.
https:\/\/doi.org\/10.1080\/00222937808682302<\/p>

Roberts, M.J. (1987) The Spiders of Great Britain and Ireland. Vol. 2. Linyphiidae and Check List<\/em>. Harley Books, Colchester, 204 pp.<\/p>

Roewer, C.F. (1931) Arachnoideen aus s\u00fcdostalpinen H\u00f6hlen, gesammelt von Herrn Karl Strasser in den Jahren 1929 und 1930. Mitteilungen \u00fcber H\u00f6hlen- und Karstforschung<\/em>, 1931 (3), 69\u201380.<\/p>

Roewer, C.F. (1942) Katalog der Araneae von 1758 bis 1940. <\/em>1. Band. Natura, Bremen. Institut Royal des Sciences Naturelles de Belgique, Bruxelles, 1040 pp.<\/p>

Rozwa\u0142ka, R. (2010) Taxonomic position of the Bathyphantes pusiolus <\/em>(Fickert, 1875) (Araneae: Linyphiidae). Genus<\/em>, 21, 315\u2013317.<\/p>

R\u016f\u017ei\u010dka, V. (1982) Modifications to improve the efficiency of pitfall traps. Newsletter of the British Arachnological Society<\/em>, 34, 2\u20134.<\/p>

R\u016f\u017ei\u010dka, V. (1990) The spiders of stony debris. Acta Zoologica Fennica<\/em>, 190, 333\u2013337.<\/p>

R\u016f\u017ei\u010dka, V. (1996) Species composition and site distribution of spiders (Araneae) in a gneiss massif in the Dyje river valley. Revue suisse de Zoologie<\/em>, Supplement, 561\u2013569.<\/p>

R\u016f\u017ei\u010dka, V. (1999a) The first steps in subterranean evolution of spiders (Araneae) in Central Europe. Journal of Natural History<\/em>, 33, 255\u2013265.<\/p>

R\u016f\u017ei\u010dka, V. (1999b) The freezing scree slopes and their arachnofauna. In<\/em>: M\u00f6seler, B.M. & Molenda, R. (Eds.), Lebensraum Blockhlade<\/em>. Decheniana-Beihefte<\/em>, 37, 141\u2013147.<\/p>

R\u016f\u017ei\u010dka, V. (2002) Spatial distribution of spiders (Araneae) on scree slopes in K\u0159ivokl\u00e1tsko and Moravsk\u00fd Kras Protected Landscape Areas. Acta Societatis Zoologicae Bohemicae<\/em>, 66, 321\u2013328.<\/p>

R\u016f\u017ei\u010dka, V. (2009) The European species of the microphthalmum<\/em>-group in the genus Porrhomma<\/em> (Araneae: Linyphiidae). Contributions to Natural History<\/em>, 12, 1081\u20131094.<\/p>

R\u016f\u017ei\u010dka, V. (2011) Central European habitats inhabited by spiders with disjunctive distributions. Polish Journal of Ecology<\/em>, 59, 367\u2013380.<\/p>

R\u016f\u017ei\u010dka, V. & Dolansk\u00fd, J. (2016) Catching of spiders in shallow subterranean habitats in the Czech Republic. Arachnologische Mitteilungen<\/em>, 51, 43\u201348.
https:\/\/doi.org\/10.5431\/aramit5106<\/p>

R\u016f\u017ei\u010dka, V. & Hejkal, J. (1997) Succession of epigeic spider communities (Araneae) on spoil banks in North Bohemia. Acta Societatis Zoologicae Bohemicae<\/em>, 61, 381\u2013388.<\/p>

R\u016f\u017ei\u010dka, V. & Klime\u0161, L. (2005) Spider (Araneae) communities of scree slopes in the Czech Republic. The Journal of Arachnology<\/em>, 33, 280\u2013289.
https:\/\/doi.org\/10.1636\/04-44.1<\/p>

R\u016f\u017ei\u010dka, V., Mlejnek, R. & \u0160milauer, P. (2010) Local diversity versus<\/em> geographical distribution of arthropods occuring in a sandstone rock labyrinth. Polish Journal of Ecology<\/em>, 58, 533\u2013544.<\/p>

R\u016f\u017ei\u010dka, V., Mlejnek, R., Ju\u0159i\u010dkov\u00e1, L., Tajovsk\u00fd, P., \u0160milauer, P. & Zaj\u00ed\u010dek, P. (2016) Invertebrates of the Macocha Abyss (Moravian Karst). Acta Carsologica<\/em>, 45 (1), 71\u201384.
https:\/\/doi.org\/10.3986\/ac.v45i1.896<\/p>

R\u016f\u017ei\u010dka, V., \u0160milauer, P. & Mlejnek, R. (2013) Colonization of subterranean habitats by spiders in Central Europe. International Journal of Speleology<\/em>, 42, 133\u2013140.
https:\/\/doi.org\/10.5038\/1827-806X.42.2.5<\/p>

R\u016f\u017ei\u010dka, V., Zacharda, M., N\u011bmcov\u00e1, L., \u0160milauer, P. & Nekola, J.C. (2012) Periglacial microclimate in low-altitude scree slopes supports relict biodiversity. Journal of Natural History<\/em>, 46 (35\u201336), 2145\u20132157.
https:\/\/doi.org\/10.1080\/00222933.2012.707248<\/p>

Saito, H. (1977a) Cave spiders of Tochigi Prefecture, Japan. Atypus<\/em>, 69, 21\u201332. [in Japanese]<\/p>

Saito, H. (1977b) A new spider of the genus Porrhomma<\/em> (Araneae: Linyphiidae) from caves of Tochigi Prefecture, Japan. Acta Arachnologica<\/em>, 27, 48\u201352.<\/p>

Sanocka, E. (1982) Eye regression in Porrhomma moravicum<\/em> Mill. et Krat. (Aranei). Zoologica poloniae<\/em>, 29 (1\u20132), 13\u201321.<\/p>

Segers, H. (1988) Enkele nieuwe of zeldzame spinnensoorten voor de Belgische fauna (Araneae, Linyphiidae). Nieuwsbrief van de Belgische Arachnologische Vereniging<\/em>, 8, 43\u201345 [in French and English summary]<\/p>

\u0160ilhav\u00fd, V. (1958) Nov\u00fd druh pavouk\u016f z okol\u00ed T\u0159eb\u00ed\u010de Porrhoma<\/em> [sic] moderatum<\/em> n. sp. Nova species aranearum Moraviae occidentalis, \u010cSR: Porrhoma moderatum<\/em> n. sp. Vlastiv\u011bdn\u00fd sborn\u00edk Vyso\u010diny, Odd\u00edl v\u011bd p\u0159\u00edrodn\u00edch<\/em>, 2, 103\u2013107. [in Czech and Latin, German summary]<\/p>

Simon, E. (1872) Notice compl\u00e9mentaire sur les arachnides cavernicoles et hypog\u00e9s. Annales de la Soci\u00e9t\u00e9 Entomologique de France<\/em>, Serie 5e<\/sup>, Tome 2, 473\u2013488.<\/p>

Simon, E. (1884) Les arachnides de France. <\/em>V-2. Encyclop\u00e9die Roret, Paris, pp. 180\u2013420.<\/p>

Simon, E. (1910) Araneae et Opiliones (Seconde S\u00e9rie). Biospeologica. XV. Archives de Zoologie Exp\u00e9rimentale et G\u00e9n\u00e9rale<\/em>, S\u00e9rie 5, 5 (2), 49\u201366.<\/p>

Simon, E. (1913) Araneae et Opiliones (Quatr\u00e8me s\u00e9rie). Biospeologica, XXX. Archives de Zoologie Exp\u00e9rimentale et G\u00e9n\u00e9rale<\/em>, 52 (2), 359\u2013386.<\/p>

Song, D., Zhu, M. & Chen, J. (1999) The spiders of China.<\/em> Hebei Science and Technology Publishing House, Shijiazhuang, 640 pp.<\/p>

Star\u0119ga, W. (1974) Baldachinspinnen (Aranei: Linyphiidae) aus der Mongolei. Annales Zoologici<\/em>, 32, 19\u201327.<\/p>

Strand, E. (1901) Theridiiden aus dem n\u00f6rdlichen Norwegen. Archiv for Mathematik og Naturvidenskad Christiania<\/em>, 24 (2), 1\u201366.<\/p>

Strand, E. (1906) Die arktischen Araneae, Opiliones und Chernetes. In<\/em>: R\u00f6mer, F. & Schaudinn, F. (Eds.), Fauna Arctica. Eine Zusammenstellung der arktischen Tierformen, mit besonderer Ber\u00fccksichtigung des Spitzbergen-Gebietes auf Grund der Ergebnisse der Deutschen Expedition in das N\u00f6rdliche Eissmeer im Jahre 1898. Vierter Band<\/em>. Gustav Fischer, Jena, pp. 431\u2013478.<\/p>

Tanasevitch, A. (2009) The linyphiid spiders of Iran (Arachnida, Araneae, Linyphiidae). Revue suisse de Zoologie<\/em>, 116 (3\u20134), 379\u2013420.<\/p>

Tanasevitch, A. (2011) Linyphiid spiders (Araneae: Linyphiidae) from Pakistan and India. Revue suisse de Zoologie<\/em>, 118 (3), 561\u2013598.<\/p>

Tanasevitch, A.V. (2012) A new species of Porrhomma<\/em> Simon, 1884 from the mountains of southern Siberia, Russia (Aranei: Linyphiidae: Linyphiinae). Arthropoda Selecta<\/em>, 21, 369\u2013374.<\/p>

Tanasevitch, A.V. (2013a) On synonymy of linyphiid spiders of the Russian fauna. 3 (Arachnida: Aranei: Linyphiidae). Arthropoda Selecta<\/em>, 22, 171\u2013187.<\/p>

Tanasevitch, A.V. (2013b) The linyphiid spiders of the Altais, southern Siberia (Aranei: Linyphiidae). Arthropoda Selecta<\/em>, 22, 267\u2013306.<\/p>

Tanasevitch, A.V. (2014) New species and records of linyphiid spiders from Laos (Araneae, Linyphiidae). Zootaxa<\/em>, 3841 (1), 67\u201389.<\/p>

Thaler, K. (1967) Zum Vorkommen von Troglohyphantes<\/em>-Arten in Tirol und dem Trentino (Arachn., Araneae, Linyphiidae). Berichte des Naturwissenschaftlich-Medizinischen Vereins in Innsbruck<\/em>, 55, 155\u2013173.<\/p>

Thaler, K. (1968) Zum Vorkommen von Porrhomma<\/em>-Arten in Tirol und anderen Alpenl\u00e4ndern (Arachn., Araneae, Linyphiidae). Berichte des Naturwissenschaftlich-Medizinischen Vereins in Innsbruck<\/em>, 56, 361\u2013388.<\/p>

Thaler, K. (1975) The genus Opistoxys<\/em> Simon\u2014a new synonym in Linyphiid spiders. Bulletin of the British Arachnological Society<\/em>, 3, 142\u2013144.<\/p>

Thaler, K. (1983) Bemerkenswerte Spinnenfunde in Nordtirol (\u00d6sterreich) und Nachbarl\u00e4ndern: Deckennetzspinnen, Linyphiidae (Arachnida: Aranei). Ver\u00f6ffentlichungen des Tiroler Landesmuseums Ferdinandeum<\/em>, 63, 135\u2013167.<\/p>

Thaler, K. (1991) Porrhomma<\/em>. In<\/em>: Heimer, S. & Nentwig, W., Spinnen Mitteleuropas: ein Bestimmungsbuch<\/em>. Paul Parey, Berlin & Hamburg, pp. 236\u2013239.<\/p>

Thaler, K., Buchar, J. & Kubcov\u00e1, L. (2003) Neue Funde von zwei Porrhomma<\/em>-Arten in Zentraleuropa (Araneae, Linyphiidae). Entomologische Nachrichten und Berichte<\/em>, 46, 173\u2013176.<\/p>

Thaler, K. & Plachter, H. (1983) Spinnen aus H\u00f6hlen der Fr\u00e4nkischen Alb, Deutschland (Arachnida: Araneae: Erigonidae, Linyphiidae). Senckenbergiana biologica<\/em>, 63, 249\u2013263.<\/p>

Thorell, T. (1875) Diagnoses Aranearum Europaearum aliquot novarum. Tijdschrift voor Entomologie<\/em>, 18, 81\u2013108.<\/p>

Tretzel, E. (1956) Ein Beitrag zur Systematik und Verbreitung mitteleurop\u00e4ischer Arten aus der Gattung Porrhomma<\/em> (Araneae, Linyphiidae). Zoologischer Anzeiger<\/em>, 157 (3\/4), 42\u201356.<\/p>

Tu, L. & Hormiga, G. (2010) The female genitalic morphology of \u201cmicronetine\u201d spiders (Araneae, Linyphiidae). Genetica<\/em>, 138, 59\u201373.
https:\/\/doi.org\/10.1007\/s10709-009-9368-9<\/p>

Wanless, F.R. (1973) The female genitalia of British spiders of the genus Lepthyphantes<\/em> (Linyphiidae) II. Bulletin of the British Arachnological Society<\/em>, 2 (7), 127\u2013142.<\/p>

Westring, N. (1851) F\u00f6rteckning \u00f6fver de till n\u00e4rvarande tid k\u00e4nde, i Sverige f\u00f6rekommande spindelarter, utg\u00f6rande ett antal af 253, deraf 132 \u00e4ro nya f\u00f6r svenska Faunan. G\u00f6teborgs Kungliga Vetenskaps- och Vitterhets-Samh\u00e4lles Handlingar<\/em>, 2, 25\u201362.<\/p>

Weiss, I. (1997) Ein weiterer Nachweis von Porrhomma myops<\/em> f\u00fcr Deutschland (Araneae: Linyphiidae). Arachnologische Mitteilungen<\/em>, 13, 41\u201342.
https:\/\/doi.org\/10.5431\/aramit1304<\/p>

Weiss, I. & Petrisor, A. (1999) List of the spiders (Arachnida: Araneae) from Romania. Travaux du Museum National d\u2019Histoire Naturelle \u00abGrigore Antipa\u00bb<\/em>, 41, 79\u2013107.<\/p>

Wiehle, H. (1956) Spinnentiere oder Arachnoidea (Araneae). 28. Familie: Linyphiidae \u2012 Baldachinspinnen. Die Tierwelt Deutschlands. 44 <\/em>Teil. Gustav Fischer, Jena, 337 pp.<\/p>

World Spider Catalog (2017) World Spider Catalog. Version 18.0. Natural History Museum Bern. Available from: http:\/\/www.wsc.nmbe.ch\/ (accessed 1 May 2017)<\/p>

Wunderlich, J. (1983) Linyphiidae aus Nepal, IV. Bisher unbekannte und f\u00fcr Nepal neue Arten (Arachnida: Araneae). Senckenbergiana biologica<\/em>, 63 (1982), 219\u2013248.<\/p>

Wunderlich, J. (1990) Porrhomma microcavense<\/em> n. sp. aus Deutschland (Arachnida: Araneae: Linyphiidae). Entomologische Zeitschrift<\/em>, 100, 164\u2013168.<\/p>

Wunderlich, J. (2008) Differing views of the taxonomy of spiders (Araneae), and on spiders\u2019 intraspecific variability. Beitr\u00e4ge zur Araneologie<\/em>, 5, 756\u2013781.<\/p>

Yaginuma, T. & Saito, H. (1981) A new cave spider of the genus Porrhomma<\/em> (Araneae, Linyphiidae) found in a limestone cave of Shikoku, southwest Japan. Journal of the Speleological Society of Japan<\/em>, 6, 33\u201336.<\/p>

\u017d\u00e1k, K., Hercman, H., Orvo\u0161v\u00e1, M. & Ja\u010dov\u00e1, I. (2009) Cryogenic cave carbonates from the Cold Wind Cave, N\u00edzke Tatry Mountains, Slovakia: Extending the age range of cryogenic cave carbonate formation to the Saalian. International Journal of Speleology<\/em>, 38 (2), 139\u2013152.
https:\/\/doi.org\/10.5038\/1827-806X.38.2.5<\/p>

\u017d\u00e1k, K., Urban, J., C\u00edlek, V. & Hercman, H. (2004) Cryogenic cave calcite from several Central European caves: age, carbon and oxygen isotopes and a genetic model. Chemical Geology<\/em>, 206, 119\u2013136.
https:\/\/doi.org\/10.1016\/j.chemgeo.2004.01.012<\/p>

Zhu, C. & Wang, H. (1983) [A new species of spider of the genus Porrhomma<\/em> (Araneae: Linyphiidae)]. Journal of the Bethune Medical University<\/em>, 9, 148\u2013149. [in Chinese]<\/p>

Zhu, C., Wen, Z. & Sun, X. (1986) Description of two new species and the male spider of one species of Linyphiidae (Arachnida: Araneae). Journal of the Norman Bethune University of the Medical Sciences<\/em>, 12, 205\u2013208. [in Chinese, English summary]<\/p>

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