{"status":"ok","message-type":"work","message-version":"1.0.0","message":{"indexed":{"date-parts":[[2023,5,7]],"date-time":"2023-05-07T23:10:21Z","timestamp":1683501021298},"reference-count":1,"publisher":"Magnolia Press","issue":"1","content-domain":{"domain":[],"crossmark-restriction":false},"short-container-title":["Zootaxa"],"abstract":"<jats:p>Two missing syntypes of the Japanese subterranean amphipod Procrangonyx japonicus (U\u00e9no, 1930), the type species of Procrangonyx Schellenberg, 1934, were rediscovered in the collections of the Kyoto University Museum. The morphology of uropod 3, which has been considered the principal diagnostic character of the genus, is redescribed on the basis of one of the syntypes, and the nomenclatural history of the generic names Procrangonyx and Eocrangonyx Schellenberg, 1937 (corrected from 1936) for some Far-Eastern subterranean amphipod species is reviewed. Owing to confusion between the terms \u201ctype fixation\u201d and \u201ctype designation\u201d\u2014the latter being just one means of accomplishing the former\u2014the view that Procrangonyx is unavailable and invalid has prevailed in recent literature. Procrangonyx was indeed proposed after 1930 with no type species \u201cdesignation\u201d, but under Articles 67.2.1 and 68.3 of the International Code of Zoological Nomenclature, Eucrangonyx japonicus U\u00e9no, 1930 was \u201cfixed\u201d as its type species by monotypy in the original publication. Since a diagnosis of the genus was also provided in the same work, Procrangonyx is available under Article 13.3 of the Code. However, because endopodal segmentation of uropod 3 proves to be variable in P. japonicus, doubt is thrown on the taxonomic distinctness of Procrangonyx vis \u00e0 vis Pseudocrangonyx Akatsuka &amp; Komai, 1922. Additionally, the publication dates of Allocrangonyx Schellenberg, 1937 and Niphargus foreli speziae Schellenberg, 1937 are corrected from 1936.\u00a0<\/jats:p>","DOI":"10.11646\/zootaxa.4532.1.4","type":"journal-article","created":{"date-parts":[[2018,12,17]],"date-time":"2018-12-17T00:15:28Z","timestamp":1545005728000},"source":"Crossref","is-referenced-by-count":1,"title":["Rediscovered syntypes of Procrangonyx japonicus, with nomenclatural consideration of some crangonyctoidean subterranean amphipods (Crustacea: Amphipoda: Allocrangonyctidae, Niphargidae, Pseudocrangonyctidae)"],"prefix":"10.11646","volume":"4532","author":[{"given":"TAKAFUMI","family":"NAKANO","sequence":"first","affiliation":[],"role":[{"role":"author","vocabulary":"crossref"}]},{"given":"KO","family":"TOMIKAWA","sequence":"additional","affiliation":[],"role":[{"role":"author","vocabulary":"crossref"}]},{"given":"MARK J.","family":"GRYGIER","sequence":"additional","affiliation":[],"role":[{"role":"author","vocabulary":"crossref"}]}],"member":"4913","published-online":{"date-parts":[[2018,12,16]]},"reference":[{"key":"44220","unstructured":"<p class=\"HeadingRunIn\">Akatsuka, K. &amp; Komai, T. (1922) <em>Pseudocrangonyx<\/em>, a new genus of subterranean amphipods from Japan. <em>Annotationes Zoologicae Japonenses<\/em>, 10, 119\u2013126.<\/p><p class=\"HeadingRunIn\">Barnard, J.L. &amp; Barnard, C.M. (1983) <em>Freshwater Amphipoda of the World. II. Handbook and Bibliography<\/em>. Hayfield Associations, Virginia, xix + 472 pp. [pp. 359\u2013830]<\/p><p class=\"HeadingRunIn\">Bate, C.S. (1859) On the genus <em>Niphargus<\/em> (Schi\u00f6dte). <em>Proceedings of the Dublin University Zoology &amp; Botanical Association<\/em>, 1, 237\u2013240.<\/p><p class=\"HeadingRunIn\">Bousfield, E.L. (1977) A new look at the systematics of gammaroidean amphipods of the world. <em>Crustaceana Supplement<\/em>, 4, 282\u2013316.<\/p><p class=\"HeadingRunIn\">Bousfield, E.L. (1983) An updated phyletic classification and palaeohistory of the Amphipoda. <em>In<\/em>: Schram, F.R. (Ed.), <em>Crustacean Issues, Volume 1. Crustacean Phylogeny<\/em>. A.A. Balkema, Rotterdam, pp. 257\u2013277.<\/p><p class=\"HeadingRunIn\">Chevreux, \u00c9. &amp; Fage, L. (1925) <em>Faune de France. 9. Amphipodes<\/em>. Paul Lechevalier, Paris, xix, 488 pp.<\/p><p class=\"HeadingRunIn\">Clark, P.F. (2018) Charles Spence Bate: what\u2019s in a name? <em>Zootaxa<\/em>, 4497 (3), 429\u2013438.<br \/> https:\/\/doi.org\/10.11646\/zootaxa.4497.3.7<\/p><p class=\"HeadingRunIn\">Holsinger, J.R. (1971) A new species of the subterranean amphipod genus <em>Allocrangonyx<\/em> (Gammaridae), with a redescription of the genus and remarks on its zoogeography. <em>International Journal of Speleology<\/em>, 3, 317\u2013331.<\/p><p class=\"HeadingRunIn\"> https:\/\/doi.org\/10.5038\/1827-806X.3.3.11<\/p><p class=\"HeadingRunIn\">Holsinger, J.R. (1977) A review of the systematics of the Holarctic amphipod family Crangonyctidae. <em>Crustaceana Supplement<\/em>, 4, 244\u2013281.<\/p><p class=\"HeadingRunIn\">Holsinger, J.R. (1986) Holarctic crangonyctid amphipods. <em>In<\/em>: Botosaneanu, L. (Ed.), <em>Stygofauna Mundi. A Faunistic, Distributional, and Ecological Synthesis of the World Fauna Inhabiting Subterranean Waters (Including the Marine Interstitial)<\/em>. E.J. Brill, Leiden, pp. 535\u2013549.<\/p><p class=\"HeadingRunIn\">Holsinger, J.R. (1989) Allocrangonyctidae and Pseudocrangonyctidae, two new families of Holarctic subterranean amphipod crustaceans (Gammaridea), with comments on their phylogenetic and zoogeographic relationships. <em>Proceedings of the Biological Society of Washington<\/em>, 102, 947\u2013959.<\/p><p class=\"HeadingRunIn\">Horton, T., Gofas, S., Kroh, A., Poore, G.C.B., Read, G., Rosenberg, G., St\u00f6hr, S., Bailly, N., Boury-Esnault, N., Brand\u00e3o, S.N., Costello, M.J., Decock, W., Dekeyzer, S., Hernandez, F., Mees, J., Paulay, G., Vandepitte, L., Vanhoorne, B. &amp; Vranken, S. (2017) Improving nomenclatural consistency: a decade of experience in the World Register of Marine Species. <em>European Journal of Taxonomy<\/em>, 389, 1\u201324.<\/p><p class=\"HeadingRunIn\"> https:\/\/doi.org\/10.5852\/ejt.2017.389<\/p><p class=\"HeadingRunIn\">Horton, T., Lowry, J., De Broyer, C., Bellan-Santini, D., Coleman, C.O., Corbari, L., Costello, M.J., Daneliya, M., Dauvin, J.-C., Fi\u0161er, C., Gasca, R., Grabowski, M., Guerra-Garc\u00eda, J.M., Hendrycks, E., Hughes, L., Jaume, D., Jazdzewski, K., Kim, Y.-H., King, R., Krapp-Schickel, T., LeCroy, S., L\u00f6rz, A.-N., Mamos, T., Senna, A.R., Serejo, C., Sket, B., Souza-Filho, J.F., Tandberg, A.H., Thomas, J., Thurston, M., Vader, W., V\u00e4in\u00f6l\u00e4, R., Vonk, R., White, K. &amp; Zeidler, W. (2018a) World Amphipoda Database. <em>Procrangonyx<\/em> Schellenberg, 1934. Available from: <a href=\"http:\/\/www.marinespecies.org\/amphipoda\/aphia.php?p=taxdetails&amp;id=550654\">http:\/\/<\/a>www.marinespecies.org\/amphipoda\/aphia.php?p=taxdetails&amp;id=550654 (accessed 2 October 2018)<\/p><p class=\"HeadingRunIn\">Horton, T., Lowry, J., De Broyer, C., Bellan-Santini, D., Coleman, C.O., Corbari, L., Costello, M.J., Daneliya, M., Dauvin, J.-C., Fi\u0161er, C., Gasca, R., Grabowski, M., Guerra-Garc\u00eda, J.M., Hendrycks, E., Hughes, L., Jaume, D., Jazdzewski, K., Kim, Y.-H., King, R., Krapp-Schickel, T., LeCroy, S., L\u00f6rz, A.-N., Mamos, T., Senna, A.R., Serejo, C., Sket, B., Souza-Filho, J.F., Tandberg, A.H., Thomas, J., Thurston, M., Vader, W., V\u00e4in\u00f6l\u00e4, R., Vonk, R., White, K. &amp; Zeidler, W. (2018b) World Amphipoda Database. <em>Eocrangonyx<\/em> Schellenberg, 1936. Available from: <a href=\"http:\/\/www.marinespecies.org\/amphipoda\/aphia.php?p=taxdetails&amp;id=876723\">http:\/\/<\/a>www.marinespecies.org\/amphipoda\/aphia.php?p=taxdetails&amp;id=876723 (accessed 2 October 2018)<\/p><p class=\"HeadingRunIn\">Horton, T., Lowry, J., De Broyer, C., Bellan-Santini, D., Coleman, C.O., Corbari, L., Costello, M.J., Daneliya, M., Dauvin, J.-C., Fi\u0161er, C., Gasca, R., Grabowski, M., Guerra-Garc\u00eda, J.M., Hendrycks, E., Hughes, L., Jaume, D., Jazdzewski, K., Kim, Y.-H., King, R., Krapp-Schickel, T., LeCroy, S., L\u00f6rz, A.-N., Mamos, T., Senna, A.R., Serejo, C., Sket, B., Souza-Filho, J.F., Tandberg, A.H., Thomas, J., Thurston, M., Vader, W., V\u00e4in\u00f6l\u00e4, R., Vonk, R., White, K. &amp; Zeidler, W. (2018c) World Amphipoda Database. <em>Allocrangonyx<\/em> Schellenberg, 1936. Available from: <a href=\"http:\/\/www.marinespecies.org\/amphipoda\/aphia.php?p=taxdetails&amp;id=488226\">http:\/\/<\/a>www.marinespecies.org\/amphipoda\/aphia.php?p=taxdetails&amp;id=488226 (accessed 2 October 2018)<\/p><p class=\"HeadingRunIn\">Horton, T., Lowry, J., De Broyer, C., Bellan-Santini, D., Coleman, C.O., Corbari, L., Costello, M.J., Daneliya, M., Dauvin, J.-C., Fi\u0161er, C., Gasca, R., Grabowski, M., Guerra-Garc\u00eda, J.M., Hendrycks, E., Hughes, L., Jaume, D., Jazdzewski, K., Kim, Y.-H., King, R., Krapp-Schickel, T., LeCroy, S., L\u00f6rz, A.-N., Mamos, T., Senna, A.R., Serejo, C., Sket, B., Souza-Filho, J.F., Tandberg, A.H., Thomas, J., Thurston, M., Vader, W., V\u00e4in\u00f6l\u00e4, R., Vonk, R., White, K. &amp; Zeidler, W. (2018d) World Amphipoda Database. <em>Niphargus foreli speziae<\/em> Schellenberg, 1936. Available from: <a href=\"http:\/\/www.marinespecies.org\/amphipoda\/aphia.php?p=taxdetails&amp;id=873577\">http:\/\/<\/a>www.marinespecies.org\/amphipoda\/aphia.php?p=taxdetails&amp;id=873577 (accessed 2 October 2018)<\/p><p class=\"HeadingRunIn\">Hou, Z.-E. &amp; Li, S. (2003) A new subterranean amphipod of the genus <em>Procrangonyx<\/em> from Beijing, China. <em>Crustaceana<\/em>, 76, 1179\u20131188.<\/p><p class=\"HeadingRunIn\"> https:\/\/doi.org\/10.1163\/156854003773123410<\/p><p class=\"HeadingRunIn\">Hubricht, L. (1943) Studies in the Nearctic freshwater Amphipoda, III. Notes on the freshwater Amphipoda of eastern United States, with descriptions of ten new species. <em>The American Midland Naturalist<\/em>, 29, 683\u2013712.<\/p><p class=\"HeadingRunIn\"> https:\/\/doi.org\/10.2307\/2421157<\/p><p class=\"HeadingRunIn\">International Commission on Zoological Nomenclature. (1985) <em>International Code of Zoological Nomenclature<\/em>. <em>3<\/em><sup>rd<\/sup> Edition. International Trust for Zoological Nomenclature, London, xx + 338 pp.<\/p><p class=\"HeadingRunIn\">International Commission on Zoological Nomenclature. (1999) <em>International Code of Zoological Nomenclature<\/em>. <em>4<\/em><sup>th<\/sup> Edition. International Trust for Zoological Nomenclature, London, xxix + 306 pp.<\/p><p class=\"HeadingRunIn\">Ishimaru, S.-i. (1994) A catalogue of gammaridean and ingolfiellidean Amphipoda recorded from the vicinity of Japan. <em>Report of the Sado Marine Biological Station Niigata University<\/em>, 24, 29\u201386.<\/p><p class=\"HeadingRunIn\">Karaman, G.S. &amp; Ruffo, S. (1986) Amphipoda: <em>Niphargus<\/em>-group (Niphargidae sensu Bousfield, 1982). <em>In<\/em>: Botosaneanu, L. (Ed.), <em>Stygofauna Mundi. A Faunistic, Distributional, and Ecological Synthesis of the World Fauna Inhabiting Subterranean Waters (Including the Marine Interstitial)<\/em>. E.J. Brill, Leiden, pp. 514\u2013534.<\/p><p class=\"HeadingRunIn\">Lowry, J.K. &amp; Myers, A.A. (2013) A phylogeny and classification of the Senticaudata subord. nov. Crustacea: Amphipoda). <em>Zootaxa<\/em>, 3610 (1), 1\u201380.<\/p><p class=\"HeadingRunIn\"> https:\/\/doi.org\/10.11646\/zootaxa.3610.1.1<\/p><p class=\"HeadingRunIn\">Mackin, J.G. (1935) Studies on the Crustacea of Oklahoma, III. Subterranean amphipods of the genera <em>Niphargus<\/em> and <em>Boruta<\/em>. <em>Transactions of the American Microscopical Society<\/em>, 54, 41\u201351.<\/p><p class=\"HeadingRunIn\"> https:\/\/doi.org\/10.2307\/3222446<\/p><p class=\"HeadingRunIn\">Nakano, T. &amp; Tomikawa, K. (2018) Reassessment of the groundwater amphipod <em>Paramoera relicta <\/em>synonymizes the genus <em>Relictomoera <\/em>with <em>Paramoera <\/em>(Crustacea: Amphipoda: Pontogeneiidae). <em>Zoological Science<\/em>, 35, 459\u2013467.<\/p><p class=\"HeadingRunIn\"> https:\/\/doi.org\/10.2108\/zs180058<\/p><p class=\"HeadingRunIn\">Schellenberg, A. (1932) Bemerkungen \u00fcber subterrane Amphipoden Gro\u00dfbritanniens. <em>Zoologischer Anzeiger<\/em>, 99, 49\u201358.<\/p><p class=\"HeadingRunIn\">Schellenberg, A. (1934) Eine neue Amphipoden-Gattung aus einer belgischen H\u00f6hle, nebst Bemerkungen \u00fcber die Gattung <em>Crangonyx<\/em>. <em>Zoologischer Anzeiger<\/em>, 106, 215\u2013218.<\/p><p class=\"HeadingRunIn\">Schellenberg, A. (1937a) Die Amphipodengattungen um <em>Crangonyx<\/em>, ihre Verbreitung und ihre Arten. <em>Mitteilungen aus dem Zoologischen Museum in Berlin<\/em>, 22, 31\u201344.<\/p><p class=\"HeadingRunIn\"> https:\/\/doi.org\/10.1002\/mmnz.19370220103<\/p><p class=\"HeadingRunIn\">Schellenberg, A. (1937b) Bemerkungen zu meinem <em>Niphargus<\/em>-Schl\u00fcssel und zur Verbreitung und Variabilit\u00e4t der Arten, nebst Beschreibung neuer <em>Niphargus<\/em>-Formen. <em>Mitteilungen aus dem Zoologischen Museum in Berlin<\/em>, 22, 1\u201330.<\/p><p class=\"HeadingRunIn\"> https:\/\/doi.org\/10.1002\/mmnz.19370220102<\/p><p class=\"HeadingRunIn\">Sidorov, D.A. &amp; Barabanschikov, E.I. (2010) Findings of stygobiotic and amphibiotic Peracarida (Amphipoda) in subsurface waters of the Samarga R. Basin (Northern Primorye) and some considerations about the taxonomic position of \u00ab<em>Orchestia<\/em>\u00bb <em>solifuga<\/em> Iwasa. <em>Bulletin of the North-East Scientific Center, Russian Academy of Sciences Far East Branch<\/em>, 2010, 70\u201375.<\/p><p class=\"HeadingRunIn\">Sidorov, D.[A.] &amp; Holsinger, J.R. (2007) <em>Procrangonyx<\/em> <em>stygoedincus<\/em>, a new species of subterranean amphipod (Pseudocrangonyctidae) from the Far East of Russia, with remarks on biogeographic relationships. <em>Crustaceana<\/em>, 80, 417\u2013430.<\/p><p class=\"HeadingRunIn\"> https:\/\/doi.org\/10.1163\/156854007780440984<\/p><p class=\"HeadingRunIn\">Smith, S.I. &amp; Verrill, A.E. (1871) Notice of the Invertebrata dredged in Lake Superior in 1871, by the U. S. Lake Survey, under the direction of Gen. C. B. Comstock, S. I. Smith, naturalist. <em>The American Journal of Science and Arts<\/em>, Series 3, 2, 448\u2013454.<\/p><p class=\"HeadingRunIn\"> https:\/\/doi.org\/10.2475\/ajs.s3-2.12.448<\/p><p class=\"HeadingRunIn\">Stebbing, T.R.R. (1899) Amphipoda from the Copenhagen Museum and other sources.\u2014Part II. <em>Transactions of the Linnean Society<\/em>, Series 2 (Zoology), 7, 395\u2013432.<\/p><p class=\"HeadingRunIn\"> https:\/\/doi.org\/10.1111\/j.1096-3642.1899.tb00202.x<\/p><p class=\"HeadingRunIn\">Stoch, F. (1998) Revision of the <em>Niphargus<\/em> <em>stygius<\/em> - group in Venetia and Trentino (northeastern Italy), with description of three new species. <em>Bollettino del Museo Civico di Storia Naturale di Verona<\/em>, 22, 229\u2013274.<\/p><p class=\"HeadingRunIn\">Stock, J.H. &amp; Jo, Y.W. (1990) Two Crangonyctidae (Crustacea, Amphipoda) from subterranean waters of the Far East. <em>Stygologia<\/em>, 5, 119\u2013127.<\/p><p class=\"HeadingRunIn\">Tomikawa, K. &amp; Morino, H. (2012) An annotated inventory with a key of freshwater Amphipoda (Crustacea) from Japan. <em>Taxa, Proceedings of the Japanese Society of Systematic Zoology<\/em>, 32, 39\u201351. [in Japanese with English abstract]<\/p><p class=\"HeadingRunIn\">Tomikawa, K., Kobayashi, N., Kyono, M., Ishimaru, S.-i. &amp; Grygier, M.J. (2014) Description of a new species of <em>Sternomoera <\/em>(Crustacea: Amphipoda: Pontogeneiidae) from Japan, with an analysis of the phylogenetic relationships among the Japanese species based on the 28S rRNA gene. <em>Zoological Science<\/em>, 31, 475\u2013490.<\/p><p class=\"HeadingRunIn\"> https:\/\/doi.org\/10.2108\/zs140026<\/p><p class=\"HeadingRunIn\">Tomikawa, K. Kyoto, M., Kuribayashi, K. &amp; Nakano, T. (2017) The enigmatic groundwater amphipod <em>Awacaris<\/em> <em>kawasawai<\/em> revisited: synonymisation of the genus <em>Sternomoera<\/em>, with molecular phylogenetic analyses of <em>Awacaris<\/em> and <em>Sternomoera<\/em> species (Crustacea : Amphipoda : Pontogeneiidae). <em>Invertebrate Systematics<\/em>, 31, 125\u2013140.<\/p><p class=\"HeadingRunIn\"> https:\/\/doi.org\/10.1071\/IS16037<\/p><p class=\"HeadingRunIn\">Tomikawa, K., Nakano, T., Sato, A., Onodera, Y. &amp; Ohtaka, A. (2016) A molecular phylogeny of <em>Pseudocrangonyx<\/em> from Japan, including a new subterranean species (Crustacea, Amphipoda, Pseudocrangonyctidae). <em>Zoosystematics and Evolution<\/em>, 92, 187\u2013202.<\/p><p class=\"HeadingRunIn\"> https:\/\/doi.org\/10.3897\/zse.92.10176<\/p><p class=\"HeadingRunIn\">Tomikawa, K. &amp; Shinoda, S. (2016) Redescription of a subterranean amphipod, <em>Eocrangonyx<\/em> <em>japonicus<\/em> (Peracarida, Amphipoda, Pseudocrangonyctidae) from Japan. <em>Crustaceana<\/em>, 89, 583\u2013594. <\/p><p class=\"HeadingRunIn\"> https:\/\/doi.org\/10.1163\/15685403-00003544<\/p><p class=\"HeadingRunIn\">U\u00e9no, M. (1930) A new subterranean amphipod from Japan. <em>Annotationes Zoologicae Japonenses<\/em>, 13, 21\u201323.<\/p><p> <\/p>"}],"container-title":["Zootaxa"],"original-title":[],"link":[{"URL":"https:\/\/www.mapress.com\/zt\/article\/download\/zootaxa.4532.1.4\/17785","content-type":"application\/pdf","content-version":"vor","intended-application":"text-mining"},{"URL":"https:\/\/www.mapress.com\/zt\/article\/download\/zootaxa.4532.1.4\/17785","content-type":"unspecified","content-version":"vor","intended-application":"similarity-checking"}],"deposited":{"date-parts":[[2023,5,7]],"date-time":"2023-05-07T22:50:26Z","timestamp":1683499826000},"score":1,"resource":{"primary":{"URL":"https:\/\/www.mapress.com\/zt\/article\/view\/zootaxa.4532.1.4"}},"subtitle":[],"short-title":[],"issued":{"date-parts":[[2018,12,16]]},"references-count":1,"journal-issue":{"issue":"1","published-online":{"date-parts":[[2018,12,16]]}},"URL":"https:\/\/doi.org\/10.11646\/zootaxa.4532.1.4","relation":{},"ISSN":["1175-5334","1175-5326"],"issn-type":[{"value":"1175-5334","type":"electronic"},{"value":"1175-5326","type":"print"}],"subject":[],"published":{"date-parts":[[2018,12,16]]}}}