{"status":"ok","message-type":"work","message-version":"1.0.0","message":{"indexed":{"date-parts":[[2026,4,28]],"date-time":"2026-04-28T16:35:06Z","timestamp":1777394106627,"version":"3.51.4"},"reference-count":94,"publisher":"Magnolia Press","issue":"1","content-domain":{"domain":[],"crossmark-restriction":false},"short-container-title":["Zootaxa"],"abstract":"We describe the Cephalozygoptera, a new, extinct suborder of Odonata, composed of the families Dysagrionidae and Sieblosiidae, previously assigned to the Zygoptera, and possibly the Whetwhetaksidae n. fam. The Cephalozygoptera is close to the Zygoptera, but differs most notably by distinctive head morphology. It includes 59 to 64 species in at least 19 genera and one genus-level parataxon. One species is known from the Early Cretaceous (Congqingia rhora Zhang), possibly three from the Paleocene, and the rest from the early Eocene through late Miocene. We describe new taxa from the Ypresian Okanagan Highlands of British Columbia, Canada and Washington, United States of America: 16 new species of Dysagrionidae of the existing genus Dysagrion (D. pruettae); the new genera Okanagrion (O. threadgillae, O. hobani, O. beardi, O. lochmum, O. angustum, O. dorrellae, O. liquetoalatum, O. worleyae, all new species); Okanopteryx (O. jeppesenorum, O. fraseri, O. macabeensis, all new species); Stenodiafanus (S. westersidei, new species); the new genus-level parataxon Dysagrionites (D. delinei new species, D. sp. A, D. sp. B, both new); and one new genus and species of the new family Whetwhetaksidae (Whetwhetaksa millerae).\u00a0<\/jats:p>","DOI":"10.11646\/zootaxa.4934.1.1","type":"journal-article","created":{"date-parts":[[2021,2,24]],"date-time":"2021-02-24T00:40:05Z","timestamp":1614127205000},"source":"Crossref","is-referenced-by-count":17,"title":["The Cephalozygoptera, a new, extinct suborder of Odonata with new taxa from the early Eocene Okanagan Highlands, western North America"],"prefix":"10.11646","volume":"4934","author":[{"ORCID":"https:\/\/orcid.org\/0000-0002-4397-2497","authenticated-orcid":false,"given":"S. BRUCE","family":"ARCHIBALD","sequence":"first","affiliation":[],"role":[{"role":"author","vocabulary":"crossref"}]},{"ORCID":"https:\/\/orcid.org\/0000-0001-9491-8186","authenticated-orcid":false,"given":"ROBERT A.","family":"CANNINGS","sequence":"additional","affiliation":[],"role":[{"role":"author","vocabulary":"crossref"}]},{"ORCID":"https:\/\/orcid.org\/0000-0003-1162-0355","authenticated-orcid":false,"given":"ROBERT J.","family":"ERICKSON","sequence":"additional","affiliation":[],"role":[{"role":"author","vocabulary":"crossref"}]},{"ORCID":"https:\/\/orcid.org\/0000-0001-6224-6303","authenticated-orcid":false,"given":"SETH M.","family":"BYBEE","sequence":"additional","affiliation":[],"role":[{"role":"author","vocabulary":"crossref"}]},{"ORCID":"https:\/\/orcid.org\/0000-0001-7637-199X","authenticated-orcid":false,"given":"ROLF W.","family":"MATHEWES","sequence":"additional","affiliation":[],"role":[{"role":"author","vocabulary":"crossref"}]}],"member":"4913","published-online":{"date-parts":[[2021,2,24]]},"reference":[{"key":"#cr-split#-50561.1","doi-asserted-by":"crossref","unstructured":"Meyer, H.W. (2020) Paleoclimate and paleoecology of the latest Eocene Florissant flora of central Colorado, U.S.A. Palaeogeography, Palaeoclimatology, Palaeoecology<\/em>, 551. [in press]<\/p>

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Blanke, A., Greve, C., Mokso, R., Beckmann, F. &","DOI":"10.1080\/08912963.2012.753884"},{"key":"#cr-split#-50561.13","doi-asserted-by":"crossref","unstructured":"Misof, B. (2013) An updated phylogeny of Anisoptera including formal convergence analysis of morphological characters. Systematic Entomology<\/em>, 38, 474-490.<\/p>

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Fleck, G., Bechly, G., Mart\u00ednez-Delcl\u00f2s, X., Jarzembowski, E.A. &"},{"key":"#cr-split#-50561.23","unstructured":"Nel, A. (2004) A revision of the Mesozoic dragonfly family Tarsophlebiidae, with a discussion on the phylogenetic positions of the Tarsophlebiidae and Sieblosiidae (Odonatoptera: Panodonata). Geodiversitas<\/em>, 26, 33-60.<\/p>

Fleck, G. &"},{"key":"#cr-split#-50561.24","unstructured":"Nel, A. (2003) Revision of the Mesozoic family Aeschnidiidae (Odonata: Anisoptera). Zoologica<\/em>, 153, 1-180.<\/p>

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Garrouste, R. &"},{"key":"#cr-split#-50561.27","doi-asserted-by":"crossref","unstructured":"Nel, A. (2015) New Eocene damselflies and the first Cenozoic damseldragonfly of the isophlebiopteran lineage (Insecta: Odonata). Zootaxa<\/em>, 4028 (3), 354-366.<\/p>

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Garrouste, R. &","DOI":"10.11646\/zootaxa.4028.3.2"},{"key":"#cr-split#-50561.28","doi-asserted-by":"crossref","unstructured":"Nel, A. (2019) Alaskan Palaeogene insects: a challenge for a better knowledge of the Beringian 'route' (Odonata: Aeshnidae, Dysagrionidae). Journal of Systematic Palaeontology<\/em>, 17, 1939-1946.<\/p>

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Goloboff, P.A., Mattoni, C.I. &","DOI":"10.1111\/j.1096-0031.2008.00217.x"},{"key":"#cr-split#-50561.33","unstructured":"Quinteros, A.S. (2006) Continuous characters analyzed as such. Cladistics<\/em>, 22, 589-601.<\/p>

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Huang, D., Azar, D., Cai, C., Maksoud S., Nel, A. &","DOI":"10.1371\/journal.pone.0052594"},{"key":"#cr-split#-50561.38","doi-asserted-by":"crossref","unstructured":"Bechly, G. (2017) Mesomegaloprepidae, a remarkable new damselfly family (Odonata: Zygoptera) from mid-Cretaceous Burmese amber. Cretaceous Research<\/em>, 73, 1-13.<\/p>

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