{"status":"ok","message-type":"work","message-version":"1.0.0","message":{"indexed":{"date-parts":[[2026,7,31]],"date-time":"2026-07-31T22:11:39Z","timestamp":1785535899962,"version":"3.56.0"},"reference-count":44,"publisher":"Magnolia Press","issue":"4","content-domain":{"domain":[],"crossmark-restriction":false},"short-container-title":["Zootaxa"],"abstract":"<jats:p>Damarchus pylorus sp. nov., D. lanna sp. nov. and D. dao sp. nov. are described from males and females collected in western and northern Thailand. Damarchus pylorus sp. nov. and D. lanna sp. nov. are morphologically unique within the genus by males having a para-embolic apophysis and a keel composed of coarse laminae on the lower (posterior) side of the palpal organ, and by females having sperm receptacles, composed of a strongly sclerotized base and an unsclerotized head, situated on the ventral side of a relatively long genital atrium. The distantly related D. dao sp. nov. is distinguished from all other congeners by males having a tibia I coupling spur with three megaspines. Presumed relationships of these three species and of the genus Damarchus are discussed. Information on biology and burrow structures of the new species is provided. Damarchus pylorus sp. nov. stands out by plugging the side shaft of its burrow with a soil pellet; similar plugging devices in unbranched burrows are known from other spider families. \n\u00a0<\/jats:p>","DOI":"10.11646\/zootaxa.5336.4.2","type":"journal-article","created":{"date-parts":[[2023,8,24]],"date-time":"2023-08-24T00:17:36Z","timestamp":1692836256000},"page":"481-508","source":"Crossref","is-referenced-by-count":4,"title":["Three new and exceptional Damarchus species from Thailand (Araneae: Bemmeridae)"],"prefix":"10.11646","volume":"5336","author":[{"ORCID":"https:\/\/orcid.org\/0000-0002-4010-6024","authenticated-orcid":false,"given":"PETER J.","family":"SCHWENDINGER","sequence":"first","affiliation":[],"role":[{"vocabulary":"crossref","role":"author"}]},{"ORCID":"https:\/\/orcid.org\/0000-0001-6432-1039","authenticated-orcid":false,"given":"KOMSAN","family":"HONGPADHARAKIREE","sequence":"additional","affiliation":[],"role":[{"vocabulary":"crossref","role":"author"}]}],"member":"4913","published-online":{"date-parts":[[2023,8,24]]},"reference":[{"key":"230504","doi-asserted-by":"crossref","unstructured":"Abraham, H.C. (1924) Some mygalomorph spiders from the Malay Peninsula. Proceedings of the Zoological Society of London, 1924, 1091\u20131124. https:\/\/doi.org\/10.1111\/j.1096-3642.1924.tb03332.x","DOI":"10.1111\/j.1096-3642.1924.tb03332.x"},{"key":"230505","unstructured":"Bacelar, A. (1937) Trap-door spiders from Algarve (south of Portugal). In: Jorge, A.R. (Ed.), Comptes rendus du 12. Congr\u00e8s International de Zoologie tenu \u00e0 Lisbonne du 15 au 21 Septembre 1935. Vol. 3. Casa Portuguese, Lisboa, pp. 1567\u20131577, pls. 78\u201379."},{"key":"230506","doi-asserted-by":"crossref","unstructured":"Basumatary, P. & Brahma, D. (2021) One new burrow spider of the genus Gravelyia Mirza & Mondal 2018 (Araneae: Nemesiidae) from north-east India. Acta Arachnologica, 70 (1), 39\u201346. https:\/\/doi.org\/10.2476\/asjaa.70.39","DOI":"10.2476\/asjaa.70.39"},{"key":"230507","doi-asserted-by":"crossref","unstructured":"Decae, A.E., Schwendinger, P.J. & Hongpadharakiree, K. (2021) Descriptions of four new trapdoor spider species in the subfamily Ummidiinae from Thailand (Araneae, Mygalomorphae, Halonoproctidae). Zootaxa, 4984 (1), 300\u2013323. https:\/\/doi.org\/10.11646\/zootaxa.4984.1.22","DOI":"10.11646\/zootaxa.4984.1.22"},{"key":"230508","unstructured":"Frade, M.A. & Bacelar, A. (1931) R\u00e9vision des Nemesia de la faune ib\u00e9rique et description d\u2018esp\u00e8ces nouvelles de ce genre. Bulletin du Mus\u00e9um National d\u2018Histoire de Paris, S\u00e9rie 2, 3 (2), 222\u2013238."},{"key":"230509","doi-asserted-by":"crossref","unstructured":"Gravely, F.H. (1921) The spiders and scorpions of Barkuda Island. Records of the Indian Museum, Calcutta, 22, 399\u2013421. https:\/\/doi.org\/10.5962\/bhl.part.1478","DOI":"10.5962\/bhl.part.1478"},{"key":"230510","doi-asserted-by":"crossref","unstructured":"Gravely, F.H. (1935) Notes on Indian mygalomorph spiders. II. 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Zootaxa, 4819 (3), 521\u2013543. https:\/\/doi.org\/10.11646\/zootaxa.4819.3.5","DOI":"10.11646\/zootaxa.4819.3.5"},{"key":"230513","doi-asserted-by":"crossref","unstructured":"Leroy, A. & Leroy, J. (2005) Notes on the natural history of a trapdoor spider Ancylotrypa Simon (Araneae, Cyrtaucheniidae) that constructs a spherical burrow plug. The Journal of Arachnology, 33, 558\u2013561. https:\/\/doi.org\/10.1636\/CS04-119.1","DOI":"10.1636\/CS04-119.1"},{"key":"230514","doi-asserted-by":"crossref","unstructured":"Li, S. & Zonstein, S. (2015) Eight new species of the spider genera Raveniola and Sinopesa from China and Vietnam (Araneae, Nemesiidae). ZooKeys, 519, 1\u201332. https:\/\/doi.org\/10.3897\/zookeys.519.8784","DOI":"10.3897\/zookeys.519.8784"},{"key":"230515","unstructured":"Main, B.Y. (1964) Spiders of Australia: a guide to their identification with brief notes on the natural history of common forms. Jacaranda Press, Brisbane, 124 pp."},{"key":"230516","unstructured":"Main, B.Y. (1972) The mygalomorph spider genus Stanwellia Rainbow & Pulleine (Dipluridae) and its relationships to Aname Koch and certain other diplurine genera. Journal and Proceedings of the Royal Society of Western Australia, 55 (4), 100\u2013114."},{"key":"230517","unstructured":"Main, B.Y. (1982) Notes on the revised taxonomic position of the Black Wishbone spider Dekana diversicolor Hogg (Mygalomorphae: Dipluridae). Journal of the Royal Society of Western Australia, 65 (1), 25\u201329."},{"key":"230518","unstructured":"Main, B.Y. (1984) Spiders. 2nd Edition. Collins, Sydney, 296 pp."},{"key":"230519","doi-asserted-by":"crossref","unstructured":"Mirza, Z.A. & Mondal, A. 2018. A new genus Gravelyia with two species of the family Nemesiidae (Araneae: Mygalomorphae) from India. 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(1985) The spider infraorder Mygalomorphae (Araneae): cladistics and systematics. Bulletin of the American Museum of Natural History, 182 (1), 1\u2013180."},{"key":"230527","unstructured":"Raven, R.J. & Schwendinger, P. (1995) Three new mygalomorph spider genera from Thailand and China (Araneae). Memoirs of the Queensland Museum, 38 (2), 623\u2013641."},{"key":"230528","doi-asserted-by":"crossref","unstructured":"Raven, R.J., Jell, P.A. & Knezour, R.A. (2015) Edwa maryae gen. et sp. nov. in the Norian Blackstone Formation of the Ipswich Basin\u2014the first Triassic spider (Mygalomorphae) from Australia. Alcheringa, 39, 1\u20135. https:\/\/doi.org\/10.1080\/03115518.2015.993300","DOI":"10.1080\/03115518.2015.993300"},{"key":"230529","unstructured":"Roewer, C.F. (1962) Einige Arachniden aus den Batu-Caves in Malaya. Pacific Insects, 4 (2), 517\u2013520."},{"key":"230530","unstructured":"Schwendinger, P.J. (1988) Biological observations on orthognathous spiders in northern Thailand (Araneae: Mesothelae, Mygalomorphae). In: Haupt, J. (Ed.), XI. Europ\u00e4isches Arachnologisches Colloquium. TUB-Dokumentationen, Kongresse und Tagungen, Berlin, 38, 231\u2013236."},{"key":"230531","unstructured":"Schwendinger, P.J. (1991) Two new trap-door spiders from Thailand (Araneae, Mygalomorphae, Idiopidae). Bulletin of the British arachnological Society, 8 (8), 233\u2013240."},{"key":"230532","unstructured":"Schwendinger, P.J. (1996) The fauna of orthognathous spiders (Araneae: Mesothelae, Mygalomorphae) in Thailand. In: Mahnert, V. (Ed.), Proceedings of the 13th International Congress of Arachnology, Geneva, 3\u20138 September 1995. Revue suisse de Zoologie, Special Edition 2, pp. 577\u2013584."},{"key":"230533","doi-asserted-by":"crossref","unstructured":"Schwendinger, P.J. & Hongpadharakiree, K. (2014) Three new Prothemenops species (Araneae: Idiopidae) from central Thailand. 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Zootaxa, 2971 (1), 33\u201339. https:\/\/doi.org\/10.11646\/zootaxa.2971.1.3","DOI":"10.11646\/zootaxa.2971.1.3"},{"key":"230537","unstructured":"Thorell, T. (1890) Studi sui ragni Malesi e Papuani. IV, 1. Annali del Museo Civico di Storia Naturale di Genova, 28, 5\u2013421."},{"key":"230538","unstructured":"Thorell, T. (1891) Spindlar fr\u00e5n Nikobarerna och andra delar af s\u00f6dra Asien. Kongliga Svenska Vetenskaps-Akademiens Handlingar, 24 (2), 1\u2013149."},{"key":"230539","doi-asserted-by":"crossref","unstructured":"Thorell, T. (1895) Descriptive catalogue of the spiders of Burma, based upon the collection made by Eugene W. Oates and preserved in the British Museum. British Museum (Natural History), London, 406 pp. https:\/\/doi.org\/10.5962\/bhl.title.17492","DOI":"10.5962\/bhl.title.17492"},{"key":"230540","unstructured":"World Spider Catalog (2023) World Spider Catalog. Version 24.5. Natural History Museum Bern, Bern. Available from: http:\/\/wsc.nmbe.ch (accessed 11August 2023)"},{"key":"230541","unstructured":"Xu, X. & Yin, C.-M. (2002) A new species of the genus Raveniola from Baiyungdong Cave, Hunan Province (Araneae: Nemesiidae). Acta Zootaxonomica Sinica, 27 (3), 474\u2013476."},{"key":"230542","unstructured":"Zhu, M. & Zang, B. (2011) Spider fauna of Henan, Arachnida: Araneae. Science Press, Beijing, 557 pp., 2 pls."},{"key":"230543","unstructured":"Zonstein, S.L. (1987) A new genus of mygalomorph spiders of the subfamily Nemesiinae (Aranei, Nemesiidae) in the Palearctic fauna. Zoologicheskii Zhurnal, 66 (10), 1013\u20131019. [in Russian]"},{"key":"230544","doi-asserted-by":"crossref","unstructured":"Zonstein, S. & Marusik, Y.M. (2012) A review of the genus Raveniola (Araneae, Nemesiidae) in China, with notes on allied genera and description of four new species from Yunnan. 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