{"status":"ok","message-type":"work","message-version":"1.0.0","message":{"indexed":{"date-parts":[[2026,9,28]],"date-time":"2026-09-28T17:34:17Z","timestamp":1790616857394,"version":"4.1.0"},"reference-count":258,"publisher":"Magnolia Press","issue":"1","content-domain":{"domain":[],"crossmark-restriction":false},"short-container-title":["Zootaxa"],"abstract":"<jats:p>Exo- and endoskeletal structures of Scydmaeninae are illustrated and described based on exemplary 351 species representing 116 genera and all tribes. Over 1700 illustrations show an unparallel morphological diversity of ant-like stone beetles, and introduce this group to coleopterists, setting standards for morphological descriptions. The morphological terminology is discussed and selected terms are defined or clarified, to ensure unambiguous understanding of terms used by various authors in a different sense. Several selected problems are discussed in detail, as the cephalic structures; a possible hypopharyngeal, and not prelabial, origin of the ligula in some beetles; structures that increase the mechanical robustness of the thorax; the astounding plasticity of evolutionary changes in the ventral prothoracic and pterothoracic pleural structures; novel discoveries in the elytral locking system (incl. elytral-metanepisternal locking device); multiple migrations of the aedeagal membranous area retractor; and a parallelism between Scydmaeninae and Pselaphinae in male secondary sexual characters. Moreover, synapomorphies of Scydmaeninae are discussed, and perspectives for future studies are outlined.\n\u00a0<\/jats:p>","DOI":"10.11646\/zootaxa.5581.1.1","type":"journal-article","created":{"date-parts":[[2025,1,29]],"date-time":"2025-01-29T23:58:51Z","timestamp":1738195131000},"page":"1-390","source":"Crossref","is-referenced-by-count":3,"title":["Introduction to Scydmaeninae (Coleoptera, Staphylinidae): morphology of adults"],"prefix":"10.11646","volume":"5581","author":[{"ORCID":"https:\/\/orcid.org\/0000-0003-2973-1803","authenticated-orcid":false,"given":"PAWE\u0141","family":"JA\u0141OSZY\u0143SKI","sequence":"first","affiliation":[],"role":[{"vocabulary":"crossref","role":"author"}]}],"member":"4913","published-online":{"date-parts":[[2025,1,30]]},"reference":[{"key":"400459","doi-asserted-by":"crossref","unstructured":"Aalbu, R.L., Kanda, K., Merkl, O., Ivie, M.A. & Johnston, M.A. (2023) Reconstitution of some tribes and genera of Lagriinae (Coleoptera, Tenebrionidae). 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(1974) Le syst\u00e8me glandulaire t\u00e9gumentaire du col\u00e9opt\u00e8re myrm\u00e9cophile Claviger testaceus Preyssler, 1790 (Pselaphidae). Zeitschrift f\u00fcr Morphologie und \u00d6kologie der Tiere, 77, 187\u2013219. https:\/\/doi.org\/10.1007\/BF00389904","DOI":"10.1007\/BF00389904"},{"key":"400483","unstructured":"Castellini, G. (1996) Revisione del genere Leptomastax Pirazzoli, 1855 (Coleoptera, Scydmaenidae). Atti del Museo Civico di Storia Naturale di Grosseto, 15, 1\u2013137."},{"key":"400484","unstructured":"Chandler, D.S. (2002) Biology, morphology and systematics of the ant-like litter beetles of Australia (Coleoptera: Staphylinidae: Pselaphinae). Memoirs on Entomology, International, 15, i\u2013vii + 1\u2013560."},{"key":"400485","unstructured":"Chapman, R.F. (2013) The insects. Structure and function. 5th Edition. Cambridge University Press, Cambridge, xxxi + 929 pp."},{"key":"400486","doi-asserted-by":"crossref","unstructured":"Dressler, C. & Beutel, R.G. 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(2009) Good-bye Scydmaenidae, or why the ant-like stone beetles should become megadiverse Staphylinidae sensu latissimo (Coleoptera). European Journal of Entomology, 106 (2), 275\u2013301. https:\/\/doi.org\/10.14411\/eje.2009.035","DOI":"10.14411\/eje.2009.035"},{"key":"400496","doi-asserted-by":"crossref","unstructured":"Hammond, P.M. (1979) 2.22. Wing-folding mechanism of beetles, with special reference to investigations of Adephagan phylogeny (Coleoptera). In: Erwin, T.L., Ball, G.E. & Whitehead, D.R. (Eds.), Carabid beetles: their evolution, natural history, and classification. Proceedings of the first international symposium of carabidology. Dr. W. Junk bv Publishers, Hague, pp. 113\u2013180. https:\/\/doi.org\/10.1007\/978-94-009-9628-1_7","DOI":"10.1007\/978-94-009-9628-1_7"},{"key":"400497","unstructured":"Hlav\u00e1\u010d, P. & Jal\u017ei\u0107, B. (2009) Endogean and cavernicolous Coleoptera of the Balkans. VIII. 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(2011a) Cephazteca, a new Neotropical genus of Cephenniini (Coleoptera: Staphylinidae: Scydmaeninae). Genus, Wroc\u0142aw, 22 (2), 227\u2013240."},{"key":"400512","doi-asserted-by":"crossref","unstructured":"Ja\u0142oszy\u0144ski, P. (2011b) Cephenniini with prothoracic glands and internal cavities: New taxa, enigmatic characters and phylogeny of the Cephennomicrus group of genera (Coleoptera, Staphylinidae, Scydmaeninae). Systematic Entomology, 36, 470\u2013496. https:\/\/doi.org\/10.1111\/j.1365-3113.2011.00574.x","DOI":"10.1111\/j.1365-3113.2011.00574.x"},{"key":"400513","doi-asserted-by":"crossref","unstructured":"Ja\u0142oszy\u0144ski, P. (2011c) Eutheia Stephens (Coleoptera, Staphylinidae, Scydmaeninae) of Central America. Studies on Neotropical Fauna and Environment, 46 (2), 121\u2013130. https:\/\/doi.org\/10.1080\/01650521.2011.575572","DOI":"10.1080\/01650521.2011.575572"},{"key":"400514","doi-asserted-by":"crossref","unstructured":"Ja\u0142oszy\u0144ski, P. 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(2014d) Taxonomic notes on Australian species of Sciacharis (Magellanoconnus) (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa, 3884 (5), 419\u2013428. https:\/\/doi.org\/10.11646\/zootaxa.3884.5.2","DOI":"10.11646\/zootaxa.3884.5.2"},{"key":"400541","doi-asserted-by":"crossref","unstructured":"Ja\u0142oszy\u0144ski, P. (2014e) \u201cThe curse of Horaeomorphus\u201d: Taxonomy of misplaced Australian Cyrtoscydmini (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa, 3828 (1), 1\u201376. https:\/\/doi.org\/10.11646\/zootaxa.3828.1.1","DOI":"10.11646\/zootaxa.3828.1.1"},{"key":"400542","doi-asserted-by":"crossref","unstructured":"Ja\u0142oszy\u0144ski, P. (2014f) The enigmatic genus Stenichnoteras Scott from the Seychelles (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa, 3811 (2), 280\u2013288. https:\/\/doi.org\/10.11646\/zootaxa.3811.2.9","DOI":"10.11646\/zootaxa.3811.2.9"},{"key":"400543","doi-asserted-by":"crossref","unstructured":"Ja\u0142oszy\u0144ski, P. (2014g) The smallest Cyrtoscydmini of Australia: Revision of Microscydmus Saulcy & Croissandeau and Penicillidmus gen. n. (Coleoptera, Staphylinidae, Scydmaeninae). Zootaxa, 3774 (1), 1\u201330. https:\/\/doi.org\/10.11646\/zootaxa.3774.1.1","DOI":"10.11646\/zootaxa.3774.1.1"},{"key":"400544","doi-asserted-by":"crossref","unstructured":"Ja\u0142oszy\u0144ski, P. (2014h) Two new species and new records of Syndicus Motschulsky (Coleoptera, Staphylinidae, Scydmaeninae) from SE Asia and Australia. Zootaxa, 3873 (2), 165\u2013177. https:\/\/doi.org\/10.11646\/zootaxa.3873.2.4","DOI":"10.11646\/zootaxa.3873.2.4"},{"key":"400545","unstructured":"Ja\u0142oszy\u0144ski, P. (2015a) A new genus of Cyrtoscydmini (Coleoptera: Staphylinidae: Scydmaeninae) from Japan. Entomological Science, 18 (3), 330\u2013339."},{"key":"400546","doi-asserted-by":"crossref","unstructured":"Ja\u0142oszy\u0144ski, P. (2015b) Confused identity: subgenera of Pseudoeudesis Binaghi belong in two tribes (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa, 4048 (4), 493\u2013507. https:\/\/doi.org\/10.11646\/zootaxa.4048.4.2","DOI":"10.11646\/zootaxa.4048.4.2"},{"key":"400547","doi-asserted-by":"crossref","unstructured":"Ja\u0142oszy\u0144ski, P. (2015c) Notes on the systematic placement of Sciacharis (Anthicimimus) (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa, 4033 (3), 445\u2013450. https:\/\/doi.org\/10.11646\/zootaxa.4033.3.10","DOI":"10.11646\/zootaxa.4033.3.10"},{"key":"400548","doi-asserted-by":"crossref","unstructured":"Ja\u0142oszy\u0144ski, P. (2015d) Paraneseuthia Franz in the Indian Himalayan Region and Western Palaearctic (Coleoptera, Staphylinidae, Scydmaeninae). Zootaxa, 3948 (2), 287\u2013295. https:\/\/doi.org\/10.11646\/zootaxa.3948.2.9","DOI":"10.11646\/zootaxa.3948.2.9"},{"key":"400549","doi-asserted-by":"crossref","unstructured":"Ja\u0142oszy\u0144ski, P. (2015e) Revision of the genus Stenichnaphes Franz in New Zealand (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa, 3915 (2), 250\u2013262. https:\/\/doi.org\/10.11646\/zootaxa.3915.2.4","DOI":"10.11646\/zootaxa.3915.2.4"},{"key":"400550","doi-asserted-by":"crossref","unstructured":"Ja\u0142oszy\u0144ski, P. (2015f) Schuelkelia gen. n., a new eastern Palaearctic ant-like stone beetle, with synopsis of Eurasian genera of Cyrtoscydmini (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa, 4007 (3), 343\u2013369. https:\/\/doi.org\/10.11646\/zootaxa.4007.3.3","DOI":"10.11646\/zootaxa.4007.3.3"},{"key":"400551","doi-asserted-by":"crossref","unstructured":"Ja\u0142oszy\u0144ski, P. (2015g) Taxonomy of \u201cEuconnus complex\u201d. Part III. Morphology of Euconnus subgenus Napochus and revision of the Australian species (Coleoptera, Staphylinidae, Scydmaeninae). Zootaxa, 3925 (1), 1\u201324. https:\/\/doi.org\/10.11646\/zootaxa.3925.1.1","DOI":"10.11646\/zootaxa.3925.1.1"},{"key":"400552","doi-asserted-by":"crossref","unstructured":"Ja\u0142oszy\u0144ski, P. (2015h) Taxonomy of \u201cEuconnus complex\u201d. Part V. Review of type species of Euconnus subgenera Tetramelus, Paratetramelus and Heteroconnus (Coleoptera, Staphylinidae, Scydmaeninae). Zootaxa, 4040 (3), 280\u2013300. https:\/\/doi.org\/10.11646\/zootaxa.4040.3.2","DOI":"10.11646\/zootaxa.4040.3.2"},{"key":"400553","doi-asserted-by":"crossref","unstructured":"Ja\u0142oszy\u0144ski, P. (2015i) The Cephenniini of China. VII. New species and new records of Cephennodes Reitter of Shaanxi, Gansu and Sichuan (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa, 4033 (3), 393\u2013410. https:\/\/doi.org\/10.11646\/zootaxa.4033.3.5","DOI":"10.11646\/zootaxa.4033.3.5"},{"key":"400554","doi-asserted-by":"crossref","unstructured":"Ja\u0142oszy\u0144ski, P. (2016a) A new species and new records of Pseudocephennium Reitter in Venezuela, with notes on movable endophallic structures (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa, 4088 (4), 563\u2013570. https:\/\/doi.org\/10.11646\/zootaxa.4088.4.7","DOI":"10.11646\/zootaxa.4088.4.7"},{"key":"400555","doi-asserted-by":"crossref","unstructured":"Ja\u0142oszy\u0144ski, P. (2016b) Clavigeroscydmus Franz, a new junior synonym of Scydmaenus Latreille (Coleoptera, Staphylinidae, Scydmaeninae). Zootaxa, 4072 (4), 453\u2013464. https:\/\/doi.org\/10.11646\/zootaxa.4072.4.4","DOI":"10.11646\/zootaxa.4072.4.4"},{"key":"400556","doi-asserted-by":"crossref","unstructured":"Ja\u0142oszy\u0144ski, P. (2016c) Euconnus octosetosus Franz transferred to Plaumanniola Costa Lima: implications for the systematics of Scydmaeninae (Coleoptera: Staphylinidae). 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(2019g) Neotropical Glandulariini with male cephalic modifcations: Protandroconnus Franz and Trichocircus gen. nov. (Coleoptera: Staphylinidae: Scydmaeninae). Annales Zoologici, 69 (3), 515\u2013534. https:\/\/doi.org\/10.3161\/00034541ANZ2019.69.3.003","DOI":"10.3161\/00034541ANZ2019.69.3.003"},{"key":"400595","doi-asserted-by":"crossref","unstructured":"Ja\u0142oszy\u0144ski, P. (2019h) Taxonomy of \u201cEuconnus complex\u201d. Part XIX. Status of subgenus Borneoconnus Franz, with review of species (Coleoptera, Staphylinidae, Scydmaeninae). Zootaxa, 4686 (3), 419\u2013428. https:\/\/doi.org\/10.11646\/zootaxa.4686.3.7","DOI":"10.11646\/zootaxa.4686.3.7"},{"key":"400596","doi-asserted-by":"crossref","unstructured":"Ja\u0142oszy\u0144ski, P. (2019i) Taxonomy of the \u201cAfroeudesis group\u201d of glandulariine ant-like stone beetles (Coleoptera: Staphylinidae: Scydmaeninae). 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(2020a) A new species and new records of Malagasy Megalopinus Eichelbaum (Coleoptera, Staphylinidae, Megalopsidiinae), with notes on the sperm pump in rove beetles. Zootaxa, 4768 (3), 425\u2013434. https:\/\/doi.org\/10.11646\/zootaxa.4768.3.9","DOI":"10.11646\/zootaxa.4768.3.9"},{"key":"400600","doi-asserted-by":"crossref","unstructured":"Ja\u0142oszy\u0144ski, P. (2020b) A new species of Clidicus in East Malaysia (Coleoptera, Staphylinidae, Scydmaeninae). Zootaxa, 4790 (1), 187\u2013192. https:\/\/doi.org\/10.11646\/zootaxa.4790.1.12","DOI":"10.11646\/zootaxa.4790.1.12"},{"key":"400601","doi-asserted-by":"crossref","unstructured":"Ja\u0142oszy\u0144ski, P. (2020c) Aenigmaphes gen. n., a new glandulariine genus of the Australian region (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa, 4731 (3), 439\u2013446. https:\/\/doi.org\/10.11646\/zootaxa.4731.3.12","DOI":"10.11646\/zootaxa.4731.3.12"},{"key":"400602","doi-asserted-by":"crossref","unstructured":"Ja\u0142oszy\u0144ski, P. (2020d) Clidicus minilankanus sp. n., with notes on remaining Sri Lankan Clidicus species (Coleoptera, Staphylinidae, Scydmaeninae). Zootaxa, 4718 (1), 87\u201394. https:\/\/doi.org\/10.11646\/zootaxa.4718.1.7","DOI":"10.11646\/zootaxa.4718.1.7"},{"key":"400603","doi-asserted-by":"crossref","unstructured":"Ja\u0142oszy\u0144ski, P. (2020e) Corrections to the classification of the Neotropical ant-like stone beetles misplaced in Euconnus Thomson (Coleoptera, Staphylinidae, Scydmaeninae). Part 1. Zootaxa, 4755 (2), 251\u2013270. https:\/\/doi.org\/10.11646\/zootaxa.4755.2.3","DOI":"10.11646\/zootaxa.4755.2.3"},{"key":"400604","doi-asserted-by":"crossref","unstructured":"Ja\u0142oszy\u0144ski, P. (2020f) Euconnus (Rhomboconnus): redescriptions of species described by Schaufuss and Sharp, and descriptions of five new Mexican species (Coleoptera, Staphylinidae, Scydmaeninae). 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