{"status":"ok","message-type":"work","message-version":"1.0.0","message":{"indexed":{"date-parts":[[2026,5,14]],"date-time":"2026-05-14T17:38:52Z","timestamp":1778780332879,"version":"3.51.4"},"reference-count":1,"publisher":"Magnolia Press","issue":"1","content-domain":{"domain":[],"crossmark-restriction":false},"short-container-title":["Zootaxa"],"abstract":"This paper provides formal descriptions of some of the unnamed taxa that were included in the most recent molecular phylogeny of pholcid spiders (Eberle et al. 2018, BMC Evolutionary Biology, 18, 141). The focus is on new genera and on species that belong to recently revised genera. Eight new genera and 25 new species are formally described. In Arteminae, three new species are described in Artema Walckenaer, 1837: A. bahla sp. n., A. ghubrat sp. n., and A. dhofar sp. n. (all from Oman); five new species in Arnapa gen. n. (eastern Indonesia and New Guinea): A. arfak sp. n., A. tinoor sp. n., A. manokwari sp. n., A. meja sp. n., A. tolire sp. n.; and one new species in Chisosa Huber, 2000: C. caquetio sp. n. (Netherlands Antilles). In Ninetinae, three new monotypic genera are described: Pemona gen. n., with the type species P. sapo sp. n. (Venezuela); Pinocchio gen. n., with the type species P. barauna sp. n. (Brazil); and Magana gen. n., with the type species M. velox sp. n. (Oman). In Modisiminae, three new species are described in Chibchea Huber, 2000 (all from Brazil): C. amapa sp. n., C. santosi sp. n., and C. hamadae sp. n.; one new species in Psilochorus Simon, 1893: P. bromelicolus sp. n. (Brazil); and three new monotypic genera, all from Brazil: Arenita gen. n., with the type species A. fazendinha sp. n.; Kairona gen. n., with the type species K. selva sp. n.; and Saciperere gen. n., with the type species S. catuaba sp. n. In Pholcinae, a new monotypic genus is described: Giloloa gen. n., with the type species G. sofifi sp. n. (Indonesia); three new species in the genus Aetana Huber, 2005 (all from Indonesia): A. ternate sp. n.; A. mokwam sp. n.; A. ondawamei sp. n.; and two new species in the genus Panjange Deeleman-Reinhold & Deeleman, 1983 (both from Indonesia): P. thomi sp. n., and P. togutil sp. n.\u00a0\u00a0\u00a0\u00a0\u00a0\u00a0\u00a0 Artema ghubrat is a cave-dwelling species and the only (slightly) troglomorphic representative of Arteminae; A. dhofar is presumably the closest known relative of the pantropical and synanthropic A. atlanta. The new genus Arnapa is probably species rich in eastern Indonesia and New Guinea but poorly collected; its morphological delimitation from other Australasian Arteminae (Wugigarra Huber, 2001; Holocneminus Berland, 1942; Trichocyclus Simon, 1908) needs further study. Arnapa nigromaculatus (Kulczy\u0144ski, 1911) comb. n. is newly transferred from Psilochorus. Pemona sapo is the first representative of Ninetinae from Venezuela. The genus Chibchea, previously known from the Andes only, is for the first time recorded from Brazil\/lowland Amazonia. Arenita fazendinha is among the few species in Pholcidae with extremely reduced procursus and barely modified male chelicerae. Kairona selva is unique among Pholcidae for its brush of strong hairs on a median horn anteriorly on the ocular area. Saciperere catuaba is one of only four pholcid species currently known to occur both in the Amazon and in the Atlantic Forest; however, variation indicates that more than one species might be included. It is among the few spiders known to have asymmetric genitalia (antisymmetric female internal genitalia). The Brazilian Psilochorus bromelicolus is the first South American Psilochorus of which both sexes are adequately described; however, the assignment to Psilochorus is tentative. Aetana ternate has extremely elongated procursi and accordingly elongated female internal genitalia. Aetana ondawamei and A. mokwam have almost identical male pedipalps and chelicerae (except for size) but differ clearly in the female genitalia.\u00a0<\/jats:p>","DOI":"10.11646\/zootaxa.4546.1.1","type":"journal-article","created":{"date-parts":[[2019,1,22]],"date-time":"2019-01-22T00:34:48Z","timestamp":1548117288000},"source":"Crossref","is-referenced-by-count":17,"title":["Filling the gaps: descriptions of unnamed species included in the latest molecular phylogeny of Pholcidae (Araneae)"],"prefix":"10.11646","volume":"4546","author":[{"given":"BERNHARD A.","family":"HUBER","sequence":"first","affiliation":[],"role":[{"role":"author","vocabulary":"crossref"}]},{"given":"LEONARDO S.","family":"CARVALHO","sequence":"additional","affiliation":[],"role":[{"role":"author","vocabulary":"crossref"}]}],"member":"4913","published-online":{"date-parts":[[2019,1,22]]},"reference":[{"key":"44433","unstructured":"

Aharon, S., Huber, B.A. & Gavish-Regev, E. (2017) Daddy-long-leg giants: revision of the spider genus Artema<\/em> Walckenaer, 1837 (Araneae, Pholcidae). European Journal of Taxonomy<\/em>, 376, 1\u201357.<\/p>

https:\/\/doi.org\/10.5852\/ejt.2017.376<\/p>

Beatty, J.A., Berry, J.W. & Huber, B.A. (2008) The pholcid spiders of Micronesia and Polynesia (Araneae, Pholcidae). Journal of Arachnology<\/em>, 36, 1\u201325.<\/p>

https:\/\/doi.org\/10.1636\/H05-66.1<\/p>

Brignoli, P.M. (1981) Studies on the Pholcidae, I. Notes on the genera Artema<\/em> and Physocyclus<\/em> (Araneae). Bulletin of the American Museum of Natural History<\/em>, 170 (1), 90\u2013100.<\/p>

Brown, B.V. (1993) A further chemical alternative to critical-point-drying for preparing small (or large) flies. Fly Times<\/em>, 11, 10.<\/p>

Busschere, C. de, Fannes, W., Henrard, A., Gaublomme, E., Jocqu\u00e9, R. & Baert, L. (2014) Unravelling the goblin spiders puzzle: rDNA phylogeny of the family Oonopidae (Araneae). Arthropod Systematics & Phylogeny<\/em>, 72 (2), 177\u2013192.<\/p>

Chrysanthus, O.F.M. (1967) Spiders from South New Guinea IX. Tijdschrift voor Entomologie<\/em>, 110 (5), 89\u2013105.<\/p>

Deeleman-Reinhold, C.L. (1986a) Studies on tropical Pholcidae II. Redescription of Micromerys gracilis<\/em> Bradley and Calapnita vermiformis<\/em> Simon (Araneae, Pholcidae) and description of some related new species. Memoirs of the Queensland Museum<\/em>, 22, 205\u2013224.<\/p>

Deeleman-Reinhold, C.L. (1986b) Leaf-dwelling Pholcidae in Indo-Australian rain forests. In<\/em>: Eberhard, W.G., Lubin, Y.D. & Robinson, B.C. (Eds.), Proceedings of the Ninth International Congress of Arachnology, Panama 1983<\/em>. Smithsonian Institution Press, Washington, D.C., pp. 45\u201348.<\/p>

Deeleman-Reinhold, C.L. (1995) Redescription of Holocneminus multiguttatus<\/em> Simon and description of two new species of pholcid spiders from Australia. Beitr\u00e4ge zur Araneologie<\/em>, 4, 31\u201341.<\/p>

Deeleman-Reinhold, C.L. & Deeleman, P.R. (1983) Studies on tropical Pholcidae I. Panjange<\/em>, a new genus of Indo-Australian leaf- and rock-dwelling pholcid spiders (Araneae). Zoologische Medelelingen<\/em>, 57, 121\u2013130.<\/p>

Eberle, J., Dimitrov, D., Valdez-Mondrag\u00f3n, A. & Huber, B.A. (2018) Microhabitat change drives diversification in pholcid spiders. BMC Evolutionary Biology<\/em>, 18, 141.<\/p>

https:\/\/doi.org\/10.1186\/s12862-018-1244-8<\/p>

Gertsch, W.J. (1971) A report on some Mexican cave spiders. Association for Mexican Cave Studies Bulletin<\/em>, 4, 47\u2013111.<\/p>

Gonz\u00e1lez-Sponga, M.A. (2005) Ar\u00e1cnidos de Venezuela. Tres nuevos g\u00e9neros y cuatro nuevas especies de la familia Pholcidae (Araneae). Saber, Universidad de Oriente, Venezuela<\/em>, 17, 99\u2013109.<\/p>

Huber, B.A. (1998) Genital mechanics in some Neotropical pholcid spiders (Araneae: Pholcidae), with implications for systematics. Journal of Zoology, London<\/em>, 244, 587\u2013599.<\/p>

https:\/\/doi.org\/10.1111\/j.1469-7998.1998.tb00063.x<\/p>

Huber, B.A. (2000) New World pholcid spiders (Araneae: Pholcidae): a revision at generic level. Bulletin of the American Museum of Natural History<\/em>, 254, 1\u2013348.<\/p>

https:\/\/doi.org\/10.1206\/0003-0090(2000)254<0001:nwpsap>2.0.co;2<\/a><\/p>

Huber, B.A. (2001) The pholcids of Australia (Araneae; Pholcidae): taxonomy, biogeography, and relationships. Bulletin of the American Museum of Natural History<\/em>, 260, 1\u2013144.<\/p>

https:\/\/doi.org\/10.1206\/0003-0090(2001)260%3C0001:TPOAAP%3E2.0.CO;2<\/a><\/p>

Huber, B.A. (2005a) High species diversity, male-female coevolution, and metaphyly in Southeast Asian pholcid spiders: the case of Belisana<\/em> Thorell 1898 (Araneae, Pholcidae). Zoologica<\/em>, 155, 1\u2013126.<\/p>

Huber, B.A. (2005b) Revision of the genus Spermophora<\/em> Hentz in Southeast Asia and on the Pacific Islands, with descriptions of three new genera (Araneae: Pholcidae). Zoologische Mededelingen<\/em>, 79-2 (4), 61\u2013172.<\/p>

Huber, B.A. (2009) Four new generic and 14 new specific synonymies in Pholcidae, and transfer of Pholcoides<\/em> Roewer to Filistatidae (Araneae). Zootaxa<\/em>, 1970, 64\u201368.<\/p>

Huber, B.A. (2011a) Revision and cladistic analysis of Pholcus<\/em> and closely related taxa (Araneae, Pholcidae). Bonner zoologische Monographien<\/em>, 58, 1\u2013509.<\/p>

Huber, B.A. (2011b) Phylogeny and classification of Pholcidae (Araneae): an update. Journal of Arachnology<\/em>, 39, 211\u2013222.<\/p>

https:\/\/doi.org\/10.1636\/CA10-57.1<\/p>

Huber, B.A. (2015) Small scale endemism in Brazil\u2019s Atlantic Forest: 14 new species of Mesabolivar<\/em> (Araneae, Pholcidae), each known from a single locality. Zootaxa<\/em>, 3942 (1), 1\u201360.<\/p>

https:\/\/doi.org\/10.11646\/zootaxa.3942.1.1<\/p>

Huber, B.A. (2016a) Spider diversity and endemism in a South American hotspot: 20 new species of Carapoia<\/em> (Araneae: Pholcidae) from Brazil\u2019s Atlantic Forest. Zootaxa<\/em>, 4177 (1), 1\u201369.<\/p>

https:\/\/doi.org\/10.11646\/zootaxa.4177.1.1<\/p>

Huber, B.A. (2016b) A new genus of ground and litter-dwelling pholcine spiders from Sarawak (Araneae, Pholcidae). European Journal of Taxonomy<\/em>, 186, 1\u201315.
https:\/\/doi.org\/10.5852\/ejt.2016.186<\/p>

Huber, B.A. (2017) Revision and cladistic analysis of the Southeast Asian leaf-dwelling spider genus Calapnita<\/em> Simon (Araneae, Pholcidae). Zootaxa<\/em>, 4219 (1), 1\u201363.<\/p>

https:\/\/doi.org\/10.11646\/zootaxa.4219.1.1<\/p>

Huber, B.A. (2018a) The South American spider genera Mesabolivar<\/em> and Carapoia<\/em> (Araneae, Pholcidae): new species and a framework for redrawing generic limits. Zootaxa<\/em>, 4395 (1), 1\u2013178.<\/p>

https:\/\/doi.org\/10.11646\/zootaxa.4395.1.1<\/p>

Huber, B.A. (2018b) Cave-dwelling pholcid spiders (Araneae, Pholcidae): a review. Subterranean Biology<\/em>, 26, 1\u201318.<\/p>

https:\/\/doi.org\/10.3897\/subtbiol.26.26430<\/p>

Huber, B.A. (in press) The pholcid spiders of Sri Lanka (Araneae: Pholcidae). Zootaxa<\/em>.<\/p>

Huber, B.A. & Leh Moi Ung, C. (2016) The Panjange nigrifrons<\/em> group in Borneo (Araneae: Pholcidae): high diversity in Sarawak, apparent absence in Sabah. European Journal of Taxonomy<\/em>, 184, 1\u201332.<\/p>

https:\/\/doi.org\/10.5852\/ejt.2016.184<\/p>

Huber, B.A. & Nu\u00f1eza, O.M. (2015) Evolution of genital asymmetry, exaggerated eye stalks, and extreme palpal elongation in Panjange<\/em> spiders (Araneae: Pholcidae). European Journal of Taxonomy<\/em>, 169, 1\u201346.<\/p>

https:\/\/doi.org\/10.5852\/ejt.2015.169<\/p>

Huber, B.A., Rheims, C.A. & Brescovit, A.D. (2005) Speciation without changes in genital shape: a case study on Brazilian pholcid spiders (Araneae: Pholcidae). Zoologischer Anzeiger<\/em>, 243, 273\u2013279.<\/p>

https:\/\/doi.org\/10.1016\/j.jcz.2004.12.001<\/p>

Huber, B.A., Sinclair, B. & Schmitt, M. (2007) The evolution of asymmetric genitalia in spiders and insects. Biological Reviews<\/em>, 82, 647\u2013698.<\/p>

https:\/\/doi.org\/10.1111\/j.1469-185X.2007.00029.x<\/p>

Huber, B.A., Colmenares, P.A. & Ramirez, M.J. (2014) Fourteen new generic and ten new specific synonymies in Pholcidae (Araneae), and transfer of Mystes<\/em> Bristowe to Filistatidae. Zootaxa<\/em>, 3847 (3), 413\u2013422.<\/p>

https:\/\/doi.org\/10.11646\/zootaxa.3847.3.5<\/p>

Huber, B.A., Nu\u00f1eza, O.M. & Leh Moi Ung, C. (2015a) Revision, phylogeny, and microhabitat shifts in the Southeast Asian spider genus Aetana<\/em> (Araneae, Pholcidae). European Journal of Taxonomy<\/em>, 162, 1\u201378.<\/p>

https:\/\/doi.org\/10.5852\/ejt.2015.162<\/p>

Huber, B.A., Petcharad, B. & Bumrungsri, S. (2015b) Revision of the enigmatic Southeast Asian spider genus Savarna<\/em> (Araneae, Pholcidae). European Journal of Taxonomy<\/em>, 160, 1\u201323.<\/p>

https:\/\/doi.org\/10.5852\/ejt.2015.160<\/p>

Huber, B.A., Koh, J.K.H., Ghazali, A.R.M., Nu\u00f1eza, O., Leh Moi Ung, C. & Petcharad, B. (2016a) New leaf- and litter-dwelling species of the genus Pholcus<\/em> from Southeast Asia (Araneae, Pholcidae). European Journal of Taxonomy<\/em>, 200, 1\u201345.<\/p>

https:\/\/doi.org\/10.5852\/ejt.2016.200<\/p>

Huber, B.A., Petcharad, B., Leh Moi Ung, C., Koh, J.K.H. & Ghazali, A.R.M. (2016b) The Southeast Asian Pholcus halabala<\/em> species group (Araneae, Pholcidae), new data from field observations and ultrastructure. European Journal of Taxonomy<\/em>, 190, 1\u201355.<\/p>

https:\/\/doi.org\/10.5852\/ejt.2016.190<\/p>

Huber, B.A., Nu\u00f1eza, O.M. & Leh Moi Ung, C. (2016c) The Philippine hair wax spiders and their relatives: revision of the Pholcus bicornutus<\/em> species group (Araneae, Pholcidae). European Journal of Taxonomy<\/em>, 225, 1\u201334.<\/p>

https:\/\/doi.org\/10.5852\/ejt.2016.225<\/p>

Huber, B.A., Eberle, J. & Dimitrov, D. (2018) The phylogeny of pholcid spiders (Araneae, Pholcidae): a critical evaluation of relationships suggested by molecular data. ZooKeys<\/em>, 789, 51\u2013101.<\/p>

https:\/\/doi.org\/10.3897\/zookeys.789.22781<\/p>

Huber, B.A., Caspar, K. & Eberle, J. (in press) New species reveal unexpected interspecific microhabitat diversity in the genus Uthina<\/em> Simon, 1893 (Araneae: Pholcidae). Invertebrate Systematics.<\/em><\/p>

Koch, C.L. (1850) \u00dcbersicht des Arachnidensystems<\/em>. Heft 5<\/em>. C.H. Zeh, N\u00fcrnberg, 77 pp.<\/p>

Kulczy\u0144ski, W. (1911) Spinnen aus Nord-Neu-Guinea. In: Nova Guinea. R\u00e9sultats de l'exp\u00e9dition scientifique n\u00e9erlandaise \u00e0 la Nouvelle-Guin\u00e9e en 1903 sous les auspices de Arthur Wichmann, <\/em>Leiden, Zoologie, 5 (4), pp. 423\u2013518.<\/p>

Lipscomb, D., Platnick, N. & Wheeler, Q. (2003) The intellectual content of taxonomy: a comment on DNA taxonomy. Trends in Ecology and Evolution<\/em>, 18 (2), 65\u201366.<\/p>

h<\/a>ttps:\/\/doi.org\/10.1016\/S0169-5347(02)00060-5<\/a><\/p>

Mayr E. & Ashlock, P.D. (1991) Principles of Systematic Zoology. McGraw-Hill, New York, xx + 475 pp.<\/p>

Roewer, C.F. (1960) Solifugen und Opilionoiden. Araneae Orthognathae, Haplogynae und Entelegynae. G\u00f6teborgs Kungliga Vetenskaps och Vitterhets Samh\u00e4lles Handlingar<\/em>, 8 (7), 3\u201353.<\/p>

Samyn, Y. & De Clerck, O. (2012) No name, no game. European Journal of Taxonomy<\/em>, 10, 1\u20133.<\/p>

https:\/\/doi.org\/10.5852\/ejt.2012.10<\/p>

Simon, E. (1890) \u00c9tude sur les arachnides de l'Yemen. Annales de la Soci\u00e9t\u00e9 Entomologique de France<\/em>, Series 6, 10, 77\u2013124.<\/p>

Simon, E. (1893) Histoire Naturelle des Araign\u00e9es. <\/em>1 (2). 2nd<\/sup> edition. Roret, Paris, 233 pp. [pp. 256\u2013488]<\/p>

Slowik, J. (2009) A review of the cellar spider genus Psilochorus<\/em> Simon 1893 in America north of Mexico (Araneae: Pholcidae). Zootaxa<\/em>, 2144, 1\u201353.<\/p>

Walckenaer, C.A. (1837) Histoire Naturelle des Insectes. Apt\u00e8res. I<\/em>. Libraire Encylop\u00e9dique de Roret, Paris, pp. 1\u2013682.<\/p>

Wheeler, Q.D. (2004) Taxonomic triage and the poverty of phylogeny. Philosophical Transactions of the Royal Society, London, Series B, Biological Sciences<\/em>, 359 (1444), 571\u2013583.<\/p>

https:\/\/doi.org\/10.1098\/rstb.2003.1452<\/p>

Wheeler, W.C., Coddington, J.A., Crowley, L.M., Dimitrov, D., Goloboff, P.A., Griswold, C.E., Hormiga, G., Prendini, L., Ram\u00edrez, M.J., Sierwald, P., Almeida-Silva, L., Alvarez-Padilla, F., Arnedo, M.A., Benavides, L.R., Benjamin, S.P., Bond, J.E., Grismado, C.J., Hasanf, E., Hedin, M., Izquierdo, M.A., Labarque, F.M., Ledford, J., Lopardo, L., Maddison, W.P., Miller, J.A., Piacentini, L.N., Platnick, N.I., Polotow, D., Silva-D\u00e1vila, D., Scharff, N., Sz\u0171ts, T., Ubick, D., Vink, C.J., Wood, H.M. & Zhang, J. (2017) The spider tree of life: phylogeny of Araneae based on target-gene analyses from an extensive taxon sampling. Cladistics<\/em>, 33 (6), 574\u2013616.<\/p>

https:\/\/doi.org\/10.1111\/cla.12182<\/p>

Zhang, J. & Maddison, W.P. (2013) Molecular phylogeny, divergence times and biogeography of spiders of the subfamily Euophryinae (Araneae: Salticidae). Molecular Phylogenetics and Evolution<\/em>, 68, 81\u201392.<\/p>

https:\/\/doi.org\/10.1016\/j.ympev.2013.03.017<\/p>

<\/p>

<\/p>"}],"container-title":["Zootaxa"],"original-title":[],"link":[{"URL":"https:\/\/www.mapress.com\/zt\/article\/download\/zootaxa.4546.1.1\/18808","content-type":"application\/pdf","content-version":"vor","intended-application":"text-mining"},{"URL":"https:\/\/www.mapress.com\/zt\/article\/download\/zootaxa.4546.1.1\/18808","content-type":"unspecified","content-version":"vor","intended-application":"similarity-checking"}],"deposited":{"date-parts":[[2023,5,7]],"date-time":"2023-05-07T22:16:52Z","timestamp":1683497812000},"score":1,"resource":{"primary":{"URL":"https:\/\/www.mapress.com\/zt\/article\/view\/zootaxa.4546.1.1"}},"subtitle":[],"short-title":[],"issued":{"date-parts":[[2019,1,22]]},"references-count":1,"journal-issue":{"issue":"1","published-online":{"date-parts":[[2019,1,22]]}},"URL":"https:\/\/doi.org\/10.11646\/zootaxa.4546.1.1","relation":{},"ISSN":["1175-5334","1175-5326"],"issn-type":[{"value":"1175-5334","type":"electronic"},{"value":"1175-5326","type":"print"}],"subject":[],"published":{"date-parts":[[2019,1,22]]}}}