{"status":"ok","message-type":"work","message-version":"1.0.0","message":{"indexed":{"date-parts":[[2026,10,23]],"date-time":"2026-10-23T20:37:45Z","timestamp":1792787865915,"version":"4.5.0"},"reference-count":44,"publisher":"Springer Science and Business Media LLC","issue":"1","license":[{"start":{"date-parts":[[2026,7,21]],"date-time":"2026-07-21T00:00:00Z","timestamp":1784592000000},"content-version":"tdm","delay-in-days":0,"URL":"https:\/\/creativecommons.org\/licenses\/by-nc-nd\/4.0"},{"start":{"date-parts":[[2026,10,16]],"date-time":"2026-10-16T00:00:00Z","timestamp":1792108800000},"content-version":"vor","delay-in-days":87,"URL":"https:\/\/creativecommons.org\/licenses\/by-nc-nd\/4.0"}],"funder":[{"DOI":"10.13039\/501100001809","name":"National Natural Science Foundation of China","doi-asserted-by":"publisher","award":["Nos.82470005"],"award-info":[{"award-number":["Nos.82470005"]}],"id":[{"id":"10.13039\/501100001809","id-type":"DOI","asserted-by":"publisher"}]},{"name":"National Key Research Project of MOST","award":["2023YFA0915000"],"award-info":[{"award-number":["2023YFA0915000"]}]},{"name":"Joint Program of State Key Laboratory of Microbial Technology ,Shandong University","award":["No.M2025-03"],"award-info":[{"award-number":["No.M2025-03"]}],"id":[{"id":"https:\/\/ror.org\/03x08qn04","id-type":"ROR","asserted-by":"crossref"}]}],"content-domain":{"domain":["link.springer.com"],"crossmark-restriction":false},"short-container-title":["BMC Microbiol"],"DOI":"10.1186\/s12866-026-05347-3","type":"journal-article","created":{"date-parts":[[2026,7,21]],"date-time":"2026-07-21T10:25:24Z","timestamp":1784629524000},"update-policy":"https:\/\/doi.org\/10.1007\/springer_crossmark_policy","source":"Crossref","is-referenced-by-count":0,"title":["Ec_Ou_ZC1, a new Tunavirus, reveals Cor superinfection exclusion and differential Avs susceptibility across Drexlerviridae subfamilies"],"prefix":"10.1186","volume":"26","author":[{"given":"Quanrui","family":"Zhang","sequence":"first","affiliation":[],"role":[{"vocabulary":"crossref","role":"author"}]},{"given":"Yingying","family":"Liu","sequence":"additional","affiliation":[],"role":[{"vocabulary":"crossref","role":"author"}]},{"given":"Jiaoxia","family":"Qin","sequence":"additional","affiliation":[],"role":[{"vocabulary":"crossref","role":"author"}]},{"given":"Jixia","family":"Wei","sequence":"additional","affiliation":[],"role":[{"vocabulary":"crossref","role":"author"}]},{"given":"Fan","family":"Yang","sequence":"additional","affiliation":[],"role":[{"vocabulary":"crossref","role":"author"}]},{"given":"Tao","family":"Zhang","sequence":"additional","affiliation":[],"role":[{"vocabulary":"crossref","role":"author"}]},{"given":"Qunxin","family":"She","sequence":"additional","affiliation":[],"role":[{"vocabulary":"crossref","role":"author"}]},{"given":"Chen","family":"Zhang","sequence":"additional","affiliation":[],"role":[{"vocabulary":"crossref","role":"author"}]},{"given":"Min","family":"Wu","sequence":"additional","affiliation":[],"role":[{"vocabulary":"crossref","role":"author"}]}],"member":"297","published-online":{"date-parts":[[2026,7,21]]},"reference":[{"key":"5347_CR1","doi-asserted-by":"publisher","first-page":"754","DOI":"10.1038\/s41564-018-0166-y","volume":"3","author":"M Breitbart","year":"2018","unstructured":"Breitbart M, et al. Phage puppet masters of the marine microbial realm. Nat Microbiol. 2018;3:754\u201366. https:\/\/doi.org\/10.1038\/s41564-018-0166-y.","journal-title":"Nat Microbiol"},{"key":"5347_CR2","doi-asserted-by":"publisher","first-page":"e00052","DOI":"10.1128\/jb.00052-20","volume":"202","author":"AR Mushegian","year":"2020","unstructured":"Mushegian AR. Are there 1031 virus particles on earth, or more, or fewer? J Bacteriol. 2020;202:e00052\u201320. https:\/\/doi.org\/10.1128\/jb.00052-20.","journal-title":"J Bacteriol"},{"key":"5347_CR3","doi-asserted-by":"publisher","first-page":"162","DOI":"10.4292\/wjgpt.v8.i3.162","volume":"8","author":"DM Lin","year":"2017","unstructured":"Lin DM, Koskella B, Lin HC. Phage therapy: an alternative to antibiotics in the age of multi-drug resistance. World J Gastrointest Oncol. 2017;8:162\u201373. https:\/\/doi.org\/10.4292\/wjgpt.v8.i3.162.","journal-title":"World J Gastrointest Oncol"},{"key":"5347_CR4","doi-asserted-by":"publisher","first-page":"fuaa017","DOI":"10.1093\/femsre\/fuaa017","volume":"44","author":"DP Pires","year":"2020","unstructured":"Pires DP, et al. Current challenges and future opportunities of phage therapy. FEMS Microbiol Rev. 2020;44:fuaa017.","journal-title":"FEMS Microbiol Rev"},{"key":"5347_CR5","doi-asserted-by":"publisher","unstructured":"Gordillo Altamirano FL, Barr JJ. Phage therapy in the postantibiotic era. Clin Microbiol Rev. 2019;32. https:\/\/doi.org\/10.1128\/cmr.00066-18.","DOI":"10.1128\/cmr.00066-18"},{"key":"5347_CR6","doi-asserted-by":"publisher","first-page":"17","DOI":"10.1016\/j.cell.2022.11.017","volume":"186","author":"SA Strathdee","year":"2023","unstructured":"Strathdee SA, et al. Phage therapy: from biological mechanisms to future directions. Cell. 2023;186:17\u201331. https:\/\/doi.org\/10.1016\/j.cell.2022.11.017.","journal-title":"Cell"},{"key":"5347_CR7","doi-asserted-by":"publisher","first-page":"1253","DOI":"10.1007\/s00705-020-04577-8","volume":"165","author":"EM Adriaenssens","year":"2020","unstructured":"Adriaenssens EM, et al. Taxonomy of prokaryotic viruses: 2018\u20132019 update from the ICTV Bacterial and Archaeal Viruses Subcommittee. Arch Virol. 2020;165:1253\u201360. https:\/\/doi.org\/10.1007\/s00705-020-04577-8.","journal-title":"Arch Virol"},{"key":"5347_CR8","doi-asserted-by":"publisher","first-page":"74","DOI":"10.1007\/s00705-022-05694-2","volume":"168","author":"D Turner","year":"2023","unstructured":"Turner D, et al. Abolishment of morphology-based taxa and change to binomial species names: 2022 taxonomy update of the ICTV bacterial viruses subcommittee. Arch Virol. 2023;168:74. https:\/\/doi.org\/10.1007\/s00705-022-05694-2.","journal-title":"Arch Virol"},{"key":"5347_CR9","doi-asserted-by":"publisher","first-page":"e00667","DOI":"10.1128\/jvi.00667-23","volume":"97","author":"JM Lewis","year":"2023","unstructured":"Lewis JM, et al. Characterization of the attachment of three new coliphages onto the ferrichrome transporter FhuA. J Virol. 2023;97:e00667\u201323. https:\/\/doi.org\/10.1128\/jvi.00667-23.","journal-title":"J Virol"},{"key":"5347_CR10","doi-asserted-by":"publisher","first-page":"40349","DOI":"10.1038\/srep40349","volume":"7","author":"S Hamdi","year":"2017","unstructured":"Hamdi S, et al. Characterization of two polyvalent phages infecting Enterobacteriaceae. Sci Rep. 2017;7:40349. https:\/\/doi.org\/10.1038\/srep40349.","journal-title":"Sci Rep"},{"key":"5347_CR11","doi-asserted-by":"publisher","first-page":"e0175256","DOI":"10.1371\/journal.pone.0175256","volume":"12","author":"N Soffer","year":"2017","unstructured":"Soffer N, et al. Bacteriophage preparation lytic for Shigella significantly reduces Shigella sonnei contamination in various foods. PLoS ONE. 2017;12:e0175256. https:\/\/doi.org\/10.1371\/journal.pone.0175256.","journal-title":"PLoS ONE"},{"key":"5347_CR12","doi-asserted-by":"publisher","first-page":"2112","DOI":"10.1111\/1462-2920.14590","volume":"21","author":"S Michniewski","year":"2019","unstructured":"Michniewski S, et al. Riding the wave of genomics to investigate aquatic coliphage diversity and activity. Environ Microbiol. 2019;21:2112\u201328. https:\/\/doi.org\/10.1111\/1462-2920.14590.","journal-title":"Environ Microbiol"},{"key":"5347_CR13","doi-asserted-by":"publisher","first-page":"105","DOI":"10.1007\/s00705-017-3589-5","volume":"163","author":"L Kaliniene","year":"2018","unstructured":"Kaliniene L, et al. Molecular analysis of the low-temperature Escherichia coli phage vB_EcoS_NBD2. Arch Virol. 2018;163:105\u201314. https:\/\/doi.org\/10.1007\/s00705-017-3589-5.","journal-title":"Arch Virol"},{"key":"5347_CR14","first-page":"121","volume":"7","author":"S Subramanian","year":"2020","unstructured":"Subramanian S, Parent KN, Doore SM. Ecology, structure, and evolution of Shigella phages. Annu Rev Virol 2020 Annual Reviews. 2020;7:121\u201341.","journal-title":"Annu Rev Virol 2020 Annual Reviews"},{"key":"5347_CR15","doi-asserted-by":"publisher","first-page":"1863","DOI":"10.1016\/j.cell.2023.02.029","volume":"186","author":"A Stokar-Avihail","year":"2023","unstructured":"Stokar-Avihail A, et al. Discovery of phage determinants that confer sensitivity to bacterial immune systems. Cell. 2023;186:1863\u201376. https:\/\/doi.org\/10.1016\/j.cell.2023.02.029. .e16.","journal-title":"Cell"},{"key":"5347_CR16","doi-asserted-by":"publisher","first-page":"352","DOI":"10.1038\/s41586-023-06869-w","volume":"625","author":"E Yirmiya","year":"2024","unstructured":"Yirmiya E, et al. Phages overcome bacterial immunity via diverse anti-defence proteins. Nature. 2024;625:352\u20139. https:\/\/doi.org\/10.1038\/s41586-023-06869-w.","journal-title":"Nature"},{"key":"5347_CR17","doi-asserted-by":"publisher","first-page":"365","DOI":"10.1085\/jgp.22.3.365","volume":"22","author":"EL Ellis","year":"1939","unstructured":"Ellis EL, Delbr\u00fcck M. The growth of bacteriophage. J Gen Physiol. 1939;22:365\u201384. https:\/\/doi.org\/10.1085\/jgp.22.3.365.","journal-title":"J Gen Physiol"},{"key":"5347_CR18","doi-asserted-by":"publisher","first-page":"1695","DOI":"10.1111\/j.1558-5646.2007.00132.x","volume":"61","author":"RH Heineman","year":"2007","unstructured":"Heineman RH, Bull JJ. Testing optimality with experimental evolution: lysis time in a bacteriophage. Evolution. 2007;61:1695\u2013709. https:\/\/doi.org\/10.1111\/j.1558-5646.2007.00132.x.","journal-title":"Evolution"},{"key":"5347_CR19","doi-asserted-by":"publisher","first-page":"e0137624","DOI":"10.1128\/mbio.01376-24","volume":"15","author":"M Dominguez-Mirazo","year":"2024","unstructured":"Dominguez-Mirazo M, et al. Accounting for cellular-level variation in lysis: implications for virus-host dynamics. mBio. 2024;15:e0137624. https:\/\/doi.org\/10.1128\/mbio.01376-24.","journal-title":"mBio"},{"key":"5347_CR20","doi-asserted-by":"publisher","first-page":"eaar4120","DOI":"10.1126\/science.aar4120","volume":"359","author":"S Doron","year":"2018","unstructured":"Doron S, et al. Systematic discovery of antiphage defense systems in the microbial pangenome. Science. 2018;359:eaar4120. https:\/\/doi.org\/10.1126\/science.aar4120.","journal-title":"Science"},{"key":"5347_CR21","doi-asserted-by":"publisher","first-page":"eabm4096","DOI":"10.1126\/science.abm4096","volume":"377","author":"LA Gao","year":"2022","unstructured":"Gao LA, et al. Prokaryotic innate immunity through pattern recognition of conserved viral proteins. Science. 2022;377:eabm4096. https:\/\/doi.org\/10.1126\/science.abm4096.","journal-title":"Science"},{"key":"5347_CR22","doi-asserted-by":"publisher","first-page":"1077","DOI":"10.1126\/science.aba0372","volume":"369","author":"L Gao","year":"2020","unstructured":"Gao L, et al. Diverse enzymatic activities mediate antiviral immunity in prokaryotes. Science. 2020;369:1077\u201384. https:\/\/doi.org\/10.1126\/science.aba0372.","journal-title":"Science"},{"key":"5347_CR23","doi-asserted-by":"publisher","unstructured":"Jako\u010di\u016bn\u0117 D, Moodley A. A rapid bacteriophage DNA extraction method. Methods Protoc. 2018;1. https:\/\/doi.org\/10.3390\/mps1030027.","DOI":"10.3390\/mps1030027"},{"key":"5347_CR24","doi-asserted-by":"publisher","first-page":"lqaf009","DOI":"10.1093\/nargab\/lqaf009","volume":"7","author":"AA Egorov","year":"2025","unstructured":"Egorov AA, Atkinson GC. LoVis4u: a locus visualization tool for comparative genomics and coverage profiles. NAR genom bioinform. 2025;7:lqaf009. https:\/\/doi.org\/10.1093\/nargab\/lqaf009.","journal-title":"NAR genom bioinform"},{"key":"5347_CR25","doi-asserted-by":"publisher","first-page":"23","DOI":"10.1007\/s00705-025-06485-1","volume":"171","author":"P Simmonds","year":"2025","unstructured":"Simmonds P, et al. Changes to virus taxonomy, the international code of virus classification and nomenclature, and the ICTV statutes ratified by the International Committee on Taxonomy of Viruses (2025). Arch Virol. 2025;171:23. https:\/\/doi.org\/10.1007\/s00705-025-06485-1.","journal-title":"Arch Virol"},{"key":"5347_CR26","doi-asserted-by":"publisher","DOI":"10.1101\/2025.07.15.664860","author":"DM Emms","year":"2025","unstructured":"Emms DM, et al. OrthoFinder: scalable phylogenetic orthology inference for comparative genomics. bioRxiv. 2025. https:\/\/doi.org\/10.1101\/2025.07.15.664860.","journal-title":"bioRxiv"},{"key":"5347_CR27","doi-asserted-by":"publisher","first-page":"1","DOI":"10.1186\/s13059-019-1832-y","volume":"20","author":"DM Emms","year":"2019","unstructured":"Emms DM, Kelly S. OrthoFinder: phylogenetic orthology inference for comparative genomics. Genome Biol. 2019;20:1\u201314. https:\/\/doi.org\/10.1186\/s13059-019-1832-y.","journal-title":"Genome Biol"},{"key":"5347_CR28","doi-asserted-by":"publisher","first-page":"33964","DOI":"10.1038\/srep33964","volume":"6","author":"S Deorowicz","year":"2016","unstructured":"Deorowicz S, Debudaj-Grabysz A, Gudy\u015b A. FAMSA: Fast and accurate multiple sequence alignment of huge protein families. Sci Rep. 2016;6:33964. https:\/\/doi.org\/10.1038\/srep33964.","journal-title":"Sci Rep"},{"key":"5347_CR29","doi-asserted-by":"publisher","first-page":"4949","DOI":"10.1093\/bioinformatics\/btac620","volume":"38","author":"C Zhang","year":"2022","unstructured":"Zhang C, Mirarab S. ASTRAL-Pro 2: ultrafast species tree reconstruction from multi-copy gene family trees. Bioinformatics. 2022;38:4949\u201350. https:\/\/doi.org\/10.1093\/bioinformatics\/btac620.","journal-title":"Bioinformatics"},{"key":"5347_CR30","doi-asserted-by":"publisher","first-page":"e9490","DOI":"10.1371\/journal.pone.0009490","volume":"5","author":"MN Price","year":"2010","unstructured":"Price MN, Dehal PS, Arkin AP. FastTree 2 \u2013 Approximately Maximum-Likelihood Trees for Large Alignments. PLoS ONE. 2010;5:e9490. https:\/\/doi.org\/10.1371\/journal.pone.0009490.","journal-title":"PLoS ONE"},{"key":"5347_CR31","doi-asserted-by":"publisher","first-page":"lqab067","DOI":"10.1093\/nargab\/lqab067","volume":"3","author":"P Terzian","year":"2021","unstructured":"Terzian P, et al. PHROG: families of prokaryotic virus proteins clustered using remote homology. NAR Genom Bioinform. 2021;3:lqab067. https:\/\/doi.org\/10.1093\/nargab\/lqab067.","journal-title":"NAR Genom Bioinform"},{"key":"5347_CR32","doi-asserted-by":"publisher","first-page":"7547","DOI":"10.1128\/AEM.02229-13","volume":"79","author":"J Murphy","year":"2013","unstructured":"Murphy J, et al. Bacteriophage orphan DNA methyltransferases: insights from their bacterial origin, function, and occurrence. Appl Environ Microbiol. 2013;79:7547\u201355. https:\/\/doi.org\/10.1128\/AEM.02229-13.","journal-title":"Appl Environ Microbiol"},{"key":"5347_CR33","doi-asserted-by":"publisher","first-page":"425","DOI":"10.1016\/j.virol.2004.09.005","volume":"329","author":"A Uc-Mass","year":"2004","unstructured":"Uc-Mass A, et al. An orthologue of the cor gene is involved in the exclusion of temperate lambdoid phages. Evidence that Cor inactivates FhuA receptor functions. Virology. 2004;329:425\u201333. https:\/\/doi.org\/10.1016\/j.virol.2004.09.005.","journal-title":"Virology"},{"key":"5347_CR34","doi-asserted-by":"publisher","first-page":"1049","DOI":"10.1038\/s41586-025-09260-z","volume":"644","author":"VL Taylor","year":"2025","unstructured":"Taylor VL, et al. Prophages block cell surface receptors to preserve their viral progeny. Nature. 2025;644:1049\u201357. https:\/\/doi.org\/10.1038\/s41586-025-09260-z.","journal-title":"Nature"},{"key":"5347_CR35","doi-asserted-by":"publisher","first-page":"8401","DOI":"10.1093\/nar\/gkac002","volume":"50","author":"KN LeGault","year":"2022","unstructured":"LeGault KN, et al. A phage parasite deploys a nicking nuclease effector to inhibit viral host replication. Nucleic Acids Res. 2022;50:8401\u201317. https:\/\/doi.org\/10.1093\/nar\/gkac002.","journal-title":"Nucleic Acids Res"},{"key":"5347_CR36","doi-asserted-by":"publisher","first-page":"105","DOI":"10.1126\/science.adl1356","volume":"385","author":"EA Birkholz","year":"2024","unstructured":"Birkholz EA, et al. An intron endonuclease facilitates interference competition between coinfecting viruses. Science. 2024;385:105\u201312. https:\/\/doi.org\/10.1126\/science.adl1356.","journal-title":"Science"},{"key":"5347_CR37","doi-asserted-by":"publisher","first-page":"221","DOI":"10.1089\/phage.2022.0003","volume":"3","author":"EW Bumunang","year":"2022","unstructured":"Bumunang EW, et al. Genomic profiling of non-O157 Shiga toxigenic Escherichia coli-infecting bacteriophages from South Africa. Phage. 2022;3:221\u201330. https:\/\/doi.org\/10.1089\/phage.2022.0003.","journal-title":"Phage"},{"key":"5347_CR38","doi-asserted-by":"publisher","first-page":"2379","DOI":"10.1093\/bioinformatics\/btx157","volume":"33","author":"Y Nishimura","year":"2017","unstructured":"Nishimura Y, et al. ViPTree: the viral proteomic tree server. Bioinformatics. 2017;33:2379\u201380. https:\/\/doi.org\/10.1093\/bioinformatics\/btx157.","journal-title":"Bioinformatics"},{"key":"5347_CR39","doi-asserted-by":"publisher","first-page":"e3001424","DOI":"10.1371\/journal.pbio.3001424","volume":"19","author":"E Maffei","year":"2021","unstructured":"Maffei E, et al. Systematic exploration of Escherichia coli phage\u2013host interactions with the BASEL phage collection. PLoS Biol. 2021;19:e3001424. https:\/\/doi.org\/10.1371\/journal.pbio.3001424.","journal-title":"PLoS Biol"},{"key":"5347_CR40","doi-asserted-by":"publisher","first-page":"e3003063","DOI":"10.1371\/journal.pbio.3003063","volume":"23","author":"D Humolli","year":"2025","unstructured":"Humolli D, et al. Completing the BASEL phage collection to unlock hidden diversity for systematic exploration of phage\u2013host interactions. PLoS Biol. 2025;23:e3003063. https:\/\/doi.org\/10.1371\/journal.pbio.3003063.","journal-title":"PLoS Biol"},{"key":"5347_CR41","doi-asserted-by":"publisher","unstructured":"Bucher MJ, Czy\u017c DM. Phage against the Machine: The SIE-ence of Superinfection Exclusion. Viruses. 2024;16. https:\/\/doi.org\/10.3390\/v16091348.","DOI":"10.3390\/v16091348"},{"key":"5347_CR42","doi-asserted-by":"publisher","unstructured":"Villarroel J, et al. HostPhinder: A Phage Host Prediction Tool. Viruses. 2016;8. https:\/\/doi.org\/10.3390\/v8050116.","DOI":"10.3390\/v8050116"},{"key":"5347_CR43","doi-asserted-by":"publisher","unstructured":"Bal\u00e1\u017e A, et al. PHERI-Phage Host ExploRation Pipeline. Microorganisms. 2023;11. https:\/\/doi.org\/10.3390\/microorganisms11061398.","DOI":"10.3390\/microorganisms11061398"},{"key":"5347_CR44","doi-asserted-by":"publisher","unstructured":"Moraru C, Varsani A, Kropinski AM. VIRIDIC\u2014A novel tool to calculate the intergenomic similarities of prokaryote-infecting viruses. Viruses. 2020;12. https:\/\/doi.org\/10.3390\/v12111268.","DOI":"10.3390\/v12111268"}],"container-title":["BMC Microbiology"],"original-title":[],"language":"en","link":[{"URL":"https:\/\/link.springer.com\/article\/10.1186\/s12866-026-05347-3","content-type":"text\/html","content-version":"vor","intended-application":"text-mining"},{"URL":"https:\/\/link.springer.com\/content\/pdf\/10.1186\/s12866-026-05347-3.pdf","content-type":"application\/pdf","content-version":"vor","intended-application":"text-mining"},{"URL":"https:\/\/link.springer.com\/content\/pdf\/10.1186\/s12866-026-05347-3.pdf","content-type":"application\/pdf","content-version":"vor","intended-application":"similarity-checking"}],"deposited":{"date-parts":[[2026,10,16]],"date-time":"2026-10-16T04:10:14Z","timestamp":1792123814000},"score":1,"resource":{"primary":{"URL":"https:\/\/link.springer.com\/article\/10.1186\/s12866-026-05347-3"}},"subtitle":[],"short-title":[],"issued":{"date-parts":[[2026,7,21]]},"references-count":44,"journal-issue":{"issue":"1","published-online":{"date-parts":[[2026,12]]}},"alternative-id":["5347"],"URL":"https:\/\/doi.org\/10.1186\/s12866-026-05347-3","relation":{},"ISSN":["1471-2180"],"issn-type":[{"value":"1471-2180","type":"electronic"}],"subject":[],"published":{"date-parts":[[2026,7,21]]},"assertion":[{"value":"9 December 2025","order":1,"name":"received","label":"Received","group":{"name":"ArticleHistory","label":"Article History"}},{"value":"24 June 2026","order":2,"name":"accepted","label":"Accepted","group":{"name":"ArticleHistory","label":"Article History"}},{"value":"21 July 2026","order":3,"name":"first_online","label":"First Online","group":{"name":"ArticleHistory","label":"Article History"}},{"value":"This study did not involve any experiments with human participants or animals conducted by the authors.","order":1,"name":"Ethics","label":"Ethics approval and consent to participate","group":{"name":"EthicsHeading","label":"Declarations"}},{"value":"Not applicable.","order":2,"name":"Ethics","label":"Consent for publication","group":{"name":"EthicsHeading","label":"Declarations"}},{"value":"The authors declare no competing interests.","order":3,"name":"Ethics","label":"Competing interests","group":{"name":"EthicsHeading","label":"Declarations"}}],"article-number":"891"}}