{"status":"ok","message-type":"work","message-version":"1.0.0","message":{"indexed":{"date-parts":[[2026,7,24]],"date-time":"2026-07-24T07:19:32Z","timestamp":1784877572775,"version":"3.55.0"},"reference-count":82,"publisher":"Public Library of Science (PLoS)","issue":"7","license":[{"start":{"date-parts":[[2009,7,17]],"date-time":"2009-07-17T00:00:00Z","timestamp":1247788800000},"content-version":"unspecified","delay-in-days":0,"URL":"http:\/\/creativecommons.org\/licenses\/by\/4.0\/"}],"content-domain":{"domain":["www.ploscompbiol.org"],"crossmark-restriction":false},"short-container-title":["PLoS Comput Biol"],"DOI":"10.1371\/journal.pcbi.1000436","type":"journal-article","created":{"date-parts":[[2009,7,16]],"date-time":"2009-07-16T21:55:13Z","timestamp":1247781313000},"page":"e1000436","update-policy":"https:\/\/doi.org\/10.1371\/journal.pcbi.corrections_policy","source":"Crossref","is-referenced-by-count":578,"title":["The Natural Statistics of Audiovisual Speech"],"prefix":"10.1371","volume":"5","author":[{"given":"Chandramouli","family":"Chandrasekaran","sequence":"first","affiliation":[],"role":[{"vocabulary":"crossref","role":"author"}]},{"given":"Andrea","family":"Trubanova","sequence":"additional","affiliation":[],"role":[{"vocabulary":"crossref","role":"author"}]},{"given":"S\u00e9bastien","family":"Stillittano","sequence":"additional","affiliation":[],"role":[{"vocabulary":"crossref","role":"author"}]},{"given":"Alice","family":"Caplier","sequence":"additional","affiliation":[],"role":[{"vocabulary":"crossref","role":"author"}]},{"given":"Asif A.","family":"Ghazanfar","sequence":"additional","affiliation":[],"role":[{"vocabulary":"crossref","role":"author"}]}],"member":"340","published-online":{"date-parts":[[2009,7,17]]},"reference":[{"key":"ref1","doi-asserted-by":"crossref","first-page":"734","DOI":"10.1016\/S0959-4388(01)00277-X","article-title":"Analysis of temporal patterns of communication signals.","volume":"11","author":"GS Pollack","year":"2001","journal-title":"Curr Opin Neurobiol"},{"key":"ref2","doi-asserted-by":"crossref","first-page":"e1000209","DOI":"10.1371\/journal.pcbi.1000209","article-title":"A hierarchy of time-scales and the brain.","volume":"4","author":"SJ Kiebel","year":"2008","journal-title":"PLoS Comput Biol"},{"key":"ref3","first-page":"217","article-title":"Possible principles underlying the transformation of sensory messages.","author":"H Barlow","year":"1961"},{"key":"ref4","doi-asserted-by":"crossref","first-page":"e144","DOI":"10.1371\/journal.pcbi.0020144","article-title":"Mapping Information Flow in Sensorimotor Networks.","volume":"2","author":"M Lungarella","year":"2006","journal-title":"PLoS Comput Biol"},{"key":"ref5","doi-asserted-by":"crossref","first-page":"13","DOI":"10.1162\/1064546053278973","article-title":"The development of embodied cognition: six lessons from babies.","volume":"11","author":"L Smith","year":"2005","journal-title":"Artificial Life"},{"key":"ref6","doi-asserted-by":"crossref","DOI":"10.1002\/9780470757024.ch3","article-title":"Primacy of Multimodal Speech Perception.","author":"LD Rosenblum","year":"2005"},{"key":"ref7","doi-asserted-by":"crossref","first-page":"1001","DOI":"10.1098\/rstb.2007.2155","article-title":"The processing of audio-visual speech: empirical and neural bases.","volume":"363","author":"R Campbell","year":"2008","journal-title":"Philos Trans R Soc Lond B Biol Sci"},{"key":"ref8","doi-asserted-by":"crossref","first-page":"1193","DOI":"10.1146\/annurev.neuro.24.1.1193","article-title":"Natural Image Statistics and Neural Representation.","volume":"24","author":"EP Simoncelli","year":"2001","journal-title":"Annu Rev Neurosci"},{"key":"ref9","doi-asserted-by":"crossref","first-page":"3394","DOI":"10.1121\/1.1624067","article-title":"Modulation spectra of natural sounds and ethological theories of auditory processing.","volume":"114","author":"NC Singh","year":"2003","journal-title":"J Acoust Soc Am"},{"key":"ref10","doi-asserted-by":"crossref","first-page":"691","DOI":"10.1016\/S0006-3495(99)77236-X","article-title":"Analysis of Dynamic Brain Imaging Data.","volume":"76","author":"PP Mitra","year":"1999","journal-title":"Biophys J"},{"key":"ref11","doi-asserted-by":"crossref","first-page":"245","DOI":"10.1016\/S0167-6393(02)00107-3","article-title":"The analysis of speech in different temporal integration windows: cerebral lateralization as \u2018asymmetric sampling in time\u2019.","volume":"41","author":"D Poeppel","year":"2003","journal-title":"Speech Comm"},{"key":"ref12","doi-asserted-by":"crossref","first-page":"106","DOI":"10.1016\/j.tics.2008.01.002","article-title":"Neuronal oscillations and visual amplification of speech.","volume":"12","author":"CE Schroeder","year":"2008","journal-title":"Trends Cogn Sci"},{"key":"ref13","doi-asserted-by":"crossref","first-page":"71","DOI":"10.1098\/rstb.1992.0009","article-title":"Lipreading and Audio-Visual Speech Perception.","volume":"335","author":"Q Summerfield","year":"1992","journal-title":"Philos Trans R Soc Lond B Biol Sci"},{"key":"ref14","doi-asserted-by":"crossref","first-page":"465","DOI":"10.1016\/j.wocn.2003.09.005","article-title":"Temporal properties of spontaneous speech\u2013a syllable-centric perspective.","volume":"31","author":"S Greenberg","year":"2003","journal-title":"J Phon"},{"key":"ref15","doi-asserted-by":"crossref","first-page":"405","DOI":"10.1111\/j.1467-8721.2008.00615.x","article-title":"Speech Perception as a Multimodal Phenomenon.","volume":"17","author":"LD Rosenblum","year":"2008","journal-title":"Curr Dir Psychol Sci"},{"key":"ref16","doi-asserted-by":"crossref","first-page":"2421","DOI":"10.1121\/1.2229005","article-title":"An audio-visual corpus for speech perception and automatic speech recognition.","volume":"120","author":"M Cooke","year":"2006","journal-title":"J Acoust Soc Am"},{"key":"ref17","article-title":"The DARPA TIMIT Acoustic Phonetic Continuous Speech Corpus CDROM.","author":"JS Garofolo","year":"1993"},{"key":"ref18","article-title":"X Ray Microbeam Speech Production Database","author":"J Westbury","year":"1994"},{"key":"ref19","doi-asserted-by":"crossref","first-page":"1184","DOI":"10.1121\/1.3050257","article-title":"A study of lip movements during spontaneous dialog and its application to voice activity detection.","volume":"125","author":"D Sodoyer","year":"2009","journal-title":"J Acoust Soc Am"},{"key":"ref20","doi-asserted-by":"crossref","first-page":"706","DOI":"10.1109\/TCSVT.2004.826754","article-title":"Accurate and Quasi-Automatic Lip Tracking.","volume":"14","author":"N Eveno","year":"2004","journal-title":"IEEE T Circ Syst Vid"},{"key":"ref21","article-title":"Inner Lip Segmentation by Combining Active Contours and Parametric Models.","author":"S Stillittano","year":"2008"},{"key":"ref22","first-page":"282","article-title":"Un poste visage-parole. Acquisition et traitement des contours labiaux (A device for the capture and processing of lip contours); 1990;","author":"T Lallouache"},{"key":"ref23","doi-asserted-by":"crossref","first-page":"87","DOI":"10.1038\/416087a","article-title":"Chimaeric sounds reveal dichotomies in auditory perception.","volume":"416","author":"ZM Smith","year":"2002","journal-title":"Nature"},{"key":"ref24","doi-asserted-by":"crossref","first-page":"585","DOI":"10.1121\/1.413112","article-title":"Temporal envelope and fine structure cues for speech intelligibility.","volume":"97","author":"R Drullman","year":"1995","journal-title":"J Acoust Soc Am"},{"key":"ref25","doi-asserted-by":"crossref","first-page":"773","DOI":"10.1152\/jn.90843.2008","article-title":"Different Neural Frequency Bands Integrate Faces and Voices Differently in the Superior Temporal Sulcus.","volume":"101","author":"C Chandrasekaran","year":"2009","journal-title":"J Neurophysiol"},{"key":"ref26","doi-asserted-by":"crossref","first-page":"279","DOI":"10.1016\/j.neuron.2006.12.011","article-title":"Neuronal Oscillations and Multisensory Interaction in Primary Auditory Cortex.","volume":"53","author":"P Lakatos","year":"2007","journal-title":"Neuron"},{"key":"ref27","doi-asserted-by":"crossref","first-page":"963","DOI":"10.1016\/j.cub.2008.05.043","article-title":"Integration of Bimodal Looming Signals through Neuronal Coherence in the Temporal Lobe.","volume":"18","author":"JX Maier","year":"2008","journal-title":"Curr Biol"},{"key":"ref28","doi-asserted-by":"crossref","first-page":"4093","DOI":"10.1523\/JNEUROSCI.0330-07.2007","article-title":"Looming biases in monkey auditory cortex.","volume":"27","author":"JX Maier","year":"2007","journal-title":"J Neurosci"},{"key":"ref29","doi-asserted-by":"crossref","first-page":"717","DOI":"10.1162\/089976601300014312","article-title":"Sampling Properties of the Spectrum and Coherency of Sequences of Action Potentials.","volume":"13","author":"MR Jarvis","year":"2001","journal-title":"Neural Computation"},{"key":"ref30","doi-asserted-by":"crossref","first-page":"5004","DOI":"10.1523\/JNEUROSCI.0799-05.2005","article-title":"Multisensory Integration of Dynamic Faces and Voices in Rhesus Monkey Auditory Cortex.","volume":"25","author":"AA Ghazanfar","year":"2005","journal-title":"J Neurosci"},{"key":"ref31","doi-asserted-by":"crossref","first-page":"1181","DOI":"10.1073\/pnas.0408949102","article-title":"Visual speech speeds up the neural processing of auditory speech.","volume":"102","author":"V van Wassenhove","year":"2005","journal-title":"Proc Natl Acad Sci U S A"},{"key":"ref32","doi-asserted-by":"crossref","first-page":"598","DOI":"10.1016\/j.neuropsychologia.2006.01.001","article-title":"Temporal window of integration in auditory-visual speech perception.","volume":"45","author":"V van Wassenhove","year":"2007","journal-title":"Neuropsychologia"},{"key":"ref33","doi-asserted-by":"crossref","first-page":"1197","DOI":"10.1121\/1.1288668","article-title":"The use of visible speech cues for improving auditory detection of spoken sentences.","volume":"108","author":"KW Grant","year":"2000","journal-title":"J Acoust Soc Am"},{"key":"ref34","first-page":"61","article-title":"Time-varying information for visual speech perception.","author":"LD Rosenblum","year":"1998"},{"key":"ref35","first-page":"123","article-title":"The moving face during speech communication.","author":"K Munhall","year":"1998"},{"key":"ref36","doi-asserted-by":"crossref","first-page":"141","DOI":"10.1016\/0304-3940(91)90914-F","article-title":"Seeing speech: visual information from lip movements modifies activity in the human auditory cortex.","volume":"127","author":"M Sams","year":"1991","journal-title":"Neurosci Lett"},{"key":"ref37","doi-asserted-by":"crossref","first-page":"746","DOI":"10.1038\/264746a0","article-title":"Hearing lips and seeing voices.","volume":"264","author":"H McGurk","year":"1976","journal-title":"Nature"},{"key":"ref38","doi-asserted-by":"crossref","first-page":"515","DOI":"10.1016\/j.cognition.2005.12.007","article-title":"Eye movements of monkey observers viewing vocalizing conspecifics.","volume":"101","author":"AA Ghazanfar","year":"2006","journal-title":"Cognition"},{"key":"ref39","doi-asserted-by":"crossref","first-page":"536","DOI":"10.3758\/BF03194581","article-title":"Word identification and eye fixation locations in visual and visual-plus-auditory presentations of spoken sentences.","volume":"65","author":"CR Lansing","year":"2003","journal-title":"Percept Psychophys"},{"key":"ref40","doi-asserted-by":"crossref","first-page":"19","DOI":"10.1016\/j.specom.2004.09.008","article-title":"Investigating the audio-visual speech detection advantage.","volume":"44","author":"J Kim","year":"2004","journal-title":"Speech Comm"},{"key":"ref41","doi-asserted-by":"crossref","first-page":"1159","DOI":"10.1044\/jshr.3906.1159","article-title":"Point-Light Facial Displays Enhance Comprehension of Speech in Noise.","volume":"39","author":"LD Rosenblum","year":"1996","journal-title":"J Speech Hear Res"},{"key":"ref42","doi-asserted-by":"crossref","first-page":"318","DOI":"10.1037\/0096-1523.22.2.318","article-title":"An audiovisual test of kinematic primitives for visual speech perception.","volume":"22","author":"LD Rosenblum","year":"1996","journal-title":"J Exp Psychol Hum Percept Perform"},{"key":"ref43","doi-asserted-by":"crossref","DOI":"10.7551\/mitpress\/3422.003.0015","article-title":"Spatial and Temporal Constraints on Audiovisual Speech Perception.","author":"K Munhall","year":"2004"},{"key":"ref44","doi-asserted-by":"crossref","first-page":"1165","DOI":"10.1155\/S1110865702207015","article-title":"Separation of audio-visual speech sources: a new approach exploiting the audio-visual coherence of speech stimuli.","author":"D Sodoyer","year":"2002","journal-title":"EURASIP J Appl Signal Process"},{"key":"ref45","doi-asserted-by":"crossref","DOI":"10.1109\/CVPRW.2006.34","article-title":"Audiovisual Gestalts","author":"G Monaci","year":"2006"},{"key":"ref46","doi-asserted-by":"crossref","first-page":"R922","DOI":"10.1016\/j.cub.2006.09.049","article-title":"Speech Production: The Force of Your Words.","volume":"16","author":"K Munhall","year":"2006","journal-title":"Curr Biol"},{"key":"ref47","first-page":"1","article-title":"Speech physiology, speech perception, and acoustic phonetics","author":"P Lieberman","year":"1988"},{"key":"ref48","doi-asserted-by":"crossref","first-page":"23","DOI":"10.1016\/S0167-6393(98)00048-X","article-title":"Quantitative association of vocal-tract and facial behavior.","volume":"26","author":"H Yehia","year":"1998","journal-title":"Speech Comm"},{"key":"ref49","doi-asserted-by":"crossref","first-page":"133","DOI":"10.1111\/j.0963-7214.2004.01502010.x","article-title":"Visual prosody and speech intelligibility - Head movement improves auditory speech perception.","volume":"15","author":"KG Munhall","year":"2004","journal-title":"Psych Sci"},{"key":"ref50","doi-asserted-by":"crossref","first-page":"555","DOI":"10.1006\/jpho.2002.0165","article-title":"Linking facial animation, head motion and speech acoustics.","volume":"30","author":"HC Yehia","year":"2002","journal-title":"J Phon"},{"key":"ref51","first-page":"1709","article-title":"\u201cPutting the face to the voice\u201d: matching identity across modality.","volume":"13","author":"M Kamachi","year":"2003","journal-title":"Curr Biol"},{"key":"ref52","doi-asserted-by":"crossref","first-page":"1809","DOI":"10.1371\/journal.pbio.0040326","article-title":"Implicit multisensory associations influence voice recognition.","volume":"4","author":"K von Kriegstein","year":"2006","journal-title":"PLoS Biol"},{"key":"ref53","first-page":"3","article-title":"Some preliminaries to a comprehensive account of audio-visual speech perception.","author":"Q Summerfield","year":"1987"},{"key":"ref54","doi-asserted-by":"crossref","first-page":"2670","DOI":"10.1121\/1.409836","article-title":"Effect of reducing slow temporal modulations on speech reception.","volume":"95","author":"R Drullman","year":"1994","journal-title":"J Acoust Soc Am"},{"key":"ref55","doi-asserted-by":"crossref","first-page":"303","DOI":"10.1126\/science.270.5234.303","article-title":"Speech Recognition with Primarily Temporal Cues.","volume":"270","author":"RV Shannon","year":"1995","journal-title":"Science"},{"key":"ref56","doi-asserted-by":"crossref","first-page":"13367","DOI":"10.1073\/pnas.201400998","article-title":"Speech comprehension is correlated with temporal response patterns recorded from auditory cortex.","volume":"98","author":"E Ahissar","year":"2001","journal-title":"Proc Natl Acad Sci U S A"},{"key":"ref57","doi-asserted-by":"crossref","first-page":"8738","DOI":"10.1073\/pnas.140228397","article-title":"Brain-wave representation of words by superposition of a few sine waves.","volume":"97","author":"P Suppes","year":"2000","journal-title":"Proc Natl Acad Sci U S A"},{"key":"ref58","doi-asserted-by":"crossref","first-page":"1001","DOI":"10.1016\/j.neuron.2007.06.004","article-title":"Phase Patterns of Neuronal Responses Reliably Discriminate Speech in Human Auditory Cortex.","volume":"54","author":"H Luo","year":"2007","journal-title":"Neuron"},{"key":"ref59","doi-asserted-by":"crossref","DOI":"10.1016\/B978-0-12-248550-3.50032-5","article-title":"Temporal Regulation of Speech.","author":"J Ohala","year":"1975"},{"key":"ref60","doi-asserted-by":"crossref","first-page":"1204","DOI":"10.1044\/jshr.3705.1204","article-title":"Effect of Video Frame Rate on Subjects&apos; Ability to Shadow One of Two Competing Verbal Passages.","volume":"37","author":"M Vitkovitch","year":"1994","journal-title":"J Speech Hear Res"},{"key":"ref61","doi-asserted-by":"crossref","first-page":"121","DOI":"10.1002\/(SICI)1099-0720(199604)10:2<121::AID-ACP371>3.0.CO;2-V","article-title":"Visible Speech as a Function of Image Quality: Effects of Display Parameters on Lipreading Ability.","volume":"10","author":"M Vitkovitch","year":"1996","journal-title":"Appl Cogn Psychol"},{"key":"ref62","article-title":"The multisensory roles for auditory cortex in primate vocal communication.","author":"AA Ghazanfar","year":"2009","journal-title":"Hear Res"},{"key":"ref63","doi-asserted-by":"crossref","first-page":"278","DOI":"10.1016\/j.tics.2006.04.008","article-title":"Is neocortex essentially multisensory?","volume":"10","author":"AA Ghazanfar","year":"2006","journal-title":"Trends Cogn Sci"},{"key":"ref64","doi-asserted-by":"crossref","first-page":"1904","DOI":"10.1152\/jn.00263.2005","article-title":"An oscillatory hierarchy controlling neuronal excitability and stimulus processing in the auditory cortex.","volume":"94","author":"P Lakatos","year":"2005","journal-title":"J Neurophysiol"},{"key":"ref65","doi-asserted-by":"crossref","first-page":"1626","DOI":"10.1126\/science.1128115","article-title":"High gamma power is phase-locked to theta oscillations in human neocortex.","volume":"313","author":"RT Canolty","year":"2006","journal-title":"Science"},{"key":"ref66","doi-asserted-by":"crossref","first-page":"1127","DOI":"10.1016\/j.neuron.2007.09.038","article-title":"Endogenous cortical rhythms determine cerebral specialization for speech perception and production.","volume":"56","author":"AL Giraud","year":"2007","journal-title":"Neuron"},{"key":"ref67","doi-asserted-by":"crossref","first-page":"760","DOI":"10.1038\/19652","article-title":"Cognitive restoration of reversed speech.","volume":"398","author":"K Saberi","year":"1999","journal-title":"Nature"},{"key":"ref68","doi-asserted-by":"crossref","first-page":"719","DOI":"10.1068\/p090719","article-title":"The Detection of Auditory Visual Desynchrony.","author":"NF Dixon","year":"1980","journal-title":"Perception"},{"key":"ref69","doi-asserted-by":"crossref","first-page":"423","DOI":"10.1037\/h0045283","article-title":"Perceived order in different sense modalities.","volume":"62","author":"IJ Hirsh","year":"1961","journal-title":"J Exp Psychol"},{"key":"ref70","doi-asserted-by":"crossref","first-page":"351","DOI":"10.3758\/BF03206811","article-title":"Temporal constraints on the McGurk effect.","volume":"58","author":"KG Munhall","year":"1996","journal-title":"Percept Psychophys"},{"key":"ref71","doi-asserted-by":"crossref","first-page":"1560","DOI":"10.1093\/cercor\/bhm187","article-title":"Visual Modulation of Neurons in Auditory Cortex.","volume":"18","author":"C Kayser","year":"2008","journal-title":"Cereb Cortex"},{"key":"ref72","doi-asserted-by":"crossref","first-page":"926","DOI":"10.3758\/BF03211929","article-title":"Eye movement of perceivers during audiovisual speech perception.","volume":"60","author":"E Vatikiotis-Bateson","year":"1998","journal-title":"Percept Psychophys"},{"key":"ref73","doi-asserted-by":"crossref","first-page":"162","DOI":"10.1016\/j.neuron.2007.01.003","article-title":"Paving the Way Forward: Integrating the Senses through Phase-Resetting of Cortical Oscillations.","volume":"53","author":"AA Ghazanfar","year":"2007","journal-title":"Neuron"},{"key":"ref74","doi-asserted-by":"crossref","first-page":"554","DOI":"10.1016\/S0960-9822(03)00168-4","article-title":"Eye position affects activity in primary auditory cortex of primates.","volume":"13","author":"U Werner-Reiss","year":"2003","journal-title":"Curr Biol"},{"key":"ref75","doi-asserted-by":"crossref","first-page":"3522","DOI":"10.1152\/jn.01228.2003","article-title":"Timing and laminar profile of eye-position effects on auditory responses in primate auditory cortex.","volume":"92","author":"KMG Fu","year":"2004","journal-title":"J Neurophysiol"},{"key":"ref76","doi-asserted-by":"crossref","first-page":"200","DOI":"10.1093\/cercor\/bhm046","article-title":"Transient Cortical Excitation at the Onset of Visual Fixation.","volume":"18","author":"C Rajkai","year":"2008","journal-title":"Cereb Cortex"},{"key":"ref77","first-page":"247","article-title":"How can coarticulation models account for speech sensitivity in audio-visual desynchronization?","author":"C Abry","year":"1996"},{"key":"ref78","first-page":"291","article-title":"Lips and Jaw Movements for Vowels and Consonants: Spatio-Temporal Characteristics and Bimodal Recognition Applications.","author":"P Cosi","year":"1995"},{"key":"ref79","doi-asserted-by":"crossref","DOI":"10.7551\/mitpress\/6979.001.0001","article-title":"Understanding intelligence","author":"R Pfeifer","year":"1999"},{"key":"ref80","doi-asserted-by":"crossref","first-page":"4457","DOI":"10.1523\/JNEUROSCI.0541-08.2008","article-title":"Interactions between the Superior Temporal Sulcus and Auditory Cortex Mediate Dynamic Face\/Voice Integration in Rhesus Monkeys.","volume":"28","author":"AA Ghazanfar","year":"2008","journal-title":"J Neurosci"},{"key":"ref81","doi-asserted-by":"crossref","first-page":"6747","DOI":"10.1073\/pnas.0710826105","article-title":"Simulation of talking faces in the human brain improves auditory speech recognition.","volume":"105","author":"K von Kriegstein","year":"2008","journal-title":"Proc Natl Acad Sci U S A"},{"key":"ref82","doi-asserted-by":"crossref","first-page":"1926","DOI":"10.1126\/science.1099745","article-title":"Neuronal oscillations in cortical networks.","volume":"304","author":"G Buzsaki","year":"2004","journal-title":"Science"}],"container-title":["PLoS Computational Biology"],"original-title":[],"language":"en","link":[{"URL":"http:\/\/dx.plos.org\/10.1371\/journal.pcbi.1000436","content-type":"unspecified","content-version":"vor","intended-application":"similarity-checking"}],"deposited":{"date-parts":[[2025,2,11]],"date-time":"2025-02-11T03:04:36Z","timestamp":1739243076000},"score":1,"resource":{"primary":{"URL":"https:\/\/dx.plos.org\/10.1371\/journal.pcbi.1000436"}},"subtitle":[],"editor":[{"given":"Karl J.","family":"Friston","sequence":"first","affiliation":[],"role":[{"vocabulary":"crossref","role":"editor"}]}],"short-title":[],"issued":{"date-parts":[[2009,7,17]]},"references-count":82,"journal-issue":{"issue":"7","published-online":{"date-parts":[[2009,7,17]]}},"URL":"https:\/\/doi.org\/10.1371\/journal.pcbi.1000436","relation":{},"ISSN":["1553-7358"],"issn-type":[{"value":"1553-7358","type":"electronic"}],"subject":[],"published":{"date-parts":[[2009,7,17]]}}}