{"status":"ok","message-type":"work","message-version":"1.0.0","message":{"indexed":{"date-parts":[[2026,2,19]],"date-time":"2026-02-19T08:23:36Z","timestamp":1771489416362,"version":"3.50.1"},"reference-count":118,"publisher":"Public Library of Science (PLoS)","issue":"5","content-domain":{"domain":["www.ploscompbiol.org"],"crossmark-restriction":false},"short-container-title":["PLoS Comput Biol"],"DOI":"10.1371\/journal.pcbi.1002045","type":"journal-article","created":{"date-parts":[[2011,5,19]],"date-time":"2011-05-19T20:48:49Z","timestamp":1305838129000},"page":"e1002045","update-policy":"https:\/\/doi.org\/10.1371\/journal.pcbi.corrections_policy","source":"Crossref","is-referenced-by-count":66,"title":["Spatial Learning and Action Planning in a Prefrontal Cortical Network Model"],"prefix":"10.1371","volume":"7","author":[{"given":"Louis-Emmanuel","family":"Martinet","sequence":"first","affiliation":[]},{"given":"Denis","family":"Sheynikhovich","sequence":"additional","affiliation":[]},{"given":"Karim","family":"Benchenane","sequence":"additional","affiliation":[]},{"given":"Angelo","family":"Arleo","sequence":"additional","affiliation":[]}],"member":"340","published-online":{"date-parts":[[2011,5,19]]},"reference":[{"key":"ref1","article-title":"The hippocampus as a cognitive map.","author":"J O'Keefe","year":"1978"},{"key":"ref2","doi-asserted-by":"crossref","first-page":"483","DOI":"10.1016\/S0301-0082(96)00060-3","article-title":"Biologically based artificial navigation systems: review and prospects.","volume":"51","author":"O Trullier","year":"1997","journal-title":"Prog Neurobiol"},{"key":"ref3","doi-asserted-by":"crossref","first-page":"327","DOI":"10.1142\/S0219635207001593","article-title":"Multimodal sensory integration and concurrent navigation strategies for spatial cognition in real and artificial organisms.","volume":"6","author":"A Arleo","year":"2007","journal-title":"J Integr Neurosci"},{"key":"ref4","first-page":"215","article-title":"\u201cInsight\u201d in rats.","volume":"4","author":"EC Tolman","year":"1930","journal-title":"Publ Psychol"},{"key":"ref5","doi-asserted-by":"crossref","first-page":"163","DOI":"10.1037\/0033-295X.100.2.163","article-title":"Spatial cognitive maps in animals: new hypotheses on their structure and neural mechanisms.","volume":"100","author":"B Poucet","year":"1993","journal-title":"Psychol Rev"},{"key":"ref6","doi-asserted-by":"crossref","first-page":"283","DOI":"10.1016\/S1389-0417(03)00007-X","article-title":"Map-based navigation in mobile robots: II. a review of map-learning and path-planning strategies.","volume":"4","author":"J Meyer","year":"2003","journal-title":"Cogn Syst Res"},{"key":"ref7","doi-asserted-by":"crossref","first-page":"1658","DOI":"10.1016\/j.imavis.2008.09.001","article-title":"Embodied spatial cognition: Biological and artificial systems.","volume":"27","author":"HA Mallot","year":"2009","journal-title":"Image Vision Comput"},{"key":"ref8","doi-asserted-by":"crossref","first-page":"180","DOI":"10.1002\/hipo.10173","article-title":"Path integration in mammals.","volume":"14","author":"AS Etienne","year":"2004","journal-title":"Hippocampus"},{"key":"ref9","doi-asserted-by":"crossref","first-page":"125","DOI":"10.1006\/nlme.2001.4008","article-title":"Multiple parallel memory systems in the brain of the rat.","volume":"77","author":"NM White","year":"2002","journal-title":"Neurobiol Learn Mem"},{"key":"ref10","doi-asserted-by":"crossref","first-page":"245","DOI":"10.1016\/S0028-3932(02)00157-4","article-title":"Competition among multiple memory systems: converging evidence from animal and human brain studies.","volume":"41","author":"RA Poldrack","year":"2003","journal-title":"Neuropsychologia"},{"key":"ref11","doi-asserted-by":"crossref","first-page":"299","DOI":"10.1007\/s00422-010-0400-z","article-title":"Path planning versus cue responding: a bio-inspired model of switching between navigation strategies.","volume":"103","author":"L Doll\u00e9","year":"2010","journal-title":"Biol Cybern"},{"key":"ref12","doi-asserted-by":"crossref","first-page":"663","DOI":"10.1038\/nrn1932","article-title":"Path integration and the neural basis of the \u2018cognitive map\u2019.","volume":"7","author":"BL McNaughton","year":"2006","journal-title":"Nat Rev Neurosci"},{"key":"ref13","doi-asserted-by":"crossref","first-page":"801","DOI":"10.1038\/nature03721","article-title":"Microstructure of a spatial map in the entorhinal cortex.","volume":"436","author":"T Hafting","year":"2005","journal-title":"Nature"},{"key":"ref14","article-title":"Head Direction Cells and the Neural Mechansims of Spatial Orientation.","author":"SI Wiener","year":"2005"},{"key":"ref15","doi-asserted-by":"crossref","first-page":"89","DOI":"10.1515\/REVNEURO.2004.15.2.89","article-title":"Spatial navigation and hippocampal place cell firing: the problem of goal encoding.","volume":"15","author":"B Poucet","year":"2004","journal-title":"Rev Neurosci"},{"key":"ref16","doi-asserted-by":"crossref","first-page":"663","DOI":"10.1085\/jgp.107.6.663","article-title":"The hippocampus as a cognitive graph.","volume":"107","author":"RU Muller","year":"1996","journal-title":"J Gen Physiol"},{"key":"ref17","doi-asserted-by":"crossref","first-page":"571","DOI":"10.1016\/0306-4522(89)90424-7","article-title":"The three-dimensional organization of the hippocampal formation: a review of anatomical data.","volume":"31","author":"DG Amaral","year":"1989","journal-title":"Neuroscience"},{"key":"ref18","doi-asserted-by":"crossref","first-page":"1055","DOI":"10.1126\/science.8351520","article-title":"Dynamics of the hippocampal ensemble code for space.","volume":"261","author":"MA Wilson","year":"1993","journal-title":"Science"},{"key":"ref19","doi-asserted-by":"crossref","first-page":"747","DOI":"10.1016\/j.neuron.2006.01.037","article-title":"Tracking route progression in the posterior parietal cortex.","volume":"49","author":"DA Nitz","year":"2006","journal-title":"Neuron"},{"key":"ref20","doi-asserted-by":"crossref","first-page":"4602","DOI":"10.1073\/pnas.0407332102","article-title":"Coding for spatial goals in the prelimbic\/infralimbic area of the rat frontal cortex.","volume":"102","author":"V Hok","year":"2005","journal-title":"Proc Natl Acad Sci U S A"},{"key":"ref21","article-title":"The prefrontal cortex: Anatomy, physiology and neurophysiology of the frontal lobe.","author":"JM Fuster","year":"1991"},{"key":"ref22","doi-asserted-by":"crossref","first-page":"405","DOI":"10.1101\/lm.224606","article-title":"Neural representations of location outside the hippocampus.","volume":"13","author":"JJ Knierim","year":"2006","journal-title":"Learn Mem"},{"key":"ref23","doi-asserted-by":"crossref","first-page":"7","DOI":"10.1016\/j.neuroscience.2007.06.056","article-title":"The neuroscience of remote spatial memory: a tale of two cities.","volume":"149","author":"HJ Spiers","year":"2007","journal-title":"Neuroscience"},{"key":"ref24","doi-asserted-by":"crossref","first-page":"727","DOI":"10.1093\/brain\/114.2.727","article-title":"Deficits in strategy application following frontal lobe damage in man.","volume":"114","author":"T Shallice","year":"1991","journal-title":"Brain"},{"key":"ref25","doi-asserted-by":"crossref","first-page":"474","DOI":"10.1037\/0735-7044.109.3.474","article-title":"Medial prefrontal lesions in the rat and spatial navigation: evidence for impaired planning.","volume":"109","author":"S Granon","year":"1995","journal-title":"Behav Neurosci"},{"key":"ref26","doi-asserted-by":"crossref","first-page":"1826","DOI":"10.1016\/j.neuroimage.2006.01.037","article-title":"Thoughts, behaviour, and brain dynamics during navigation in the real world.","volume":"31","author":"HJ Spiers","year":"2006","journal-title":"Neuroimage"},{"key":"ref27","doi-asserted-by":"crossref","first-page":"618","DOI":"10.1002\/hipo.20298","article-title":"A navigational guidance system in the human brain.","volume":"17","author":"HJ Spiers","year":"2007","journal-title":"Hippocampus"},{"key":"ref28","doi-asserted-by":"crossref","first-page":"574","DOI":"10.1002\/cne.903130404","article-title":"Distribution of hippocampal CA1 and subicular efferents in the prefrontal cortex of the rat studied by means of anterograde transport of phaseolus vulgaris-leucoagglutinin.","volume":"313","author":"TM Jay","year":"1991","journal-title":"J Comp Neurol"},{"key":"ref29","doi-asserted-by":"crossref","first-page":"1","DOI":"10.1016\/j.neuroscience.2006.06.027","article-title":"Interactions among the medial prefrontal cortex, hippocampus and midline thalamus in emotional and cognitive processing in the rat.","volume":"142","author":"RP Vertes","year":"2006","journal-title":"Neuroscience"},{"key":"ref30","doi-asserted-by":"crossref","first-page":"40","DOI":"10.1002\/cne.902980104","article-title":"Amygdaloid projections to the frontal cortex and the striatum in the rat.","volume":"298","author":"H Kita","year":"1990","journal-title":"J Comp Neurol"},{"key":"ref31","doi-asserted-by":"crossref","first-page":"499","DOI":"10.1126\/science.182.4111.499","article-title":"Dopaminergic terminals in the rat cortex.","volume":"182","author":"AM Thierry","year":"1973","journal-title":"Science"},{"key":"ref32","doi-asserted-by":"crossref","first-page":"3","DOI":"10.1016\/j.bbr.2003.09.028","article-title":"Do rats have a prefrontal cortex?","volume":"146","author":"HBM Uylings","year":"2003","journal-title":"Behav Brain Res"},{"key":"ref33","article-title":"The amygdala: neurobiological aspects of emotion, memory, and mental dysfunction.","author":"JP Aggleton","year":"1992"},{"key":"ref34","doi-asserted-by":"crossref","first-page":"1","DOI":"10.1152\/jn.1998.80.1.1","article-title":"Predictive reward signal of dopamine neurons.","volume":"80","author":"W Schultz","year":"1998","journal-title":"J Neurophysiol"},{"key":"ref35","doi-asserted-by":"crossref","first-page":"437","DOI":"10.1093\/cercor\/8.5.437","article-title":"Firing characteristics of deep layer neurons in prefrontal cortex in rats performing spatial working memory tasks.","volume":"8","author":"MW Jung","year":"1998","journal-title":"Cereb Cortex"},{"key":"ref36","doi-asserted-by":"crossref","first-page":"423","DOI":"10.1016\/j.jphysparis.2004.01.002","article-title":"Prefrontal cortex function, quasi-physiological stimuli, and synaptic plasticity.","volume":"97","author":"S Otani","year":"2003","journal-title":"J Physiol Paris"},{"key":"ref37","doi-asserted-by":"crossref","first-page":"7208","DOI":"10.1523\/JNEUROSCI.6068-08.2009","article-title":"Rat prefrontal cortical neurons selectively code strategy switches.","volume":"29","author":"EL Rich","year":"2009","journal-title":"J Neurosci"},{"key":"ref38","doi-asserted-by":"crossref","first-page":"319","DOI":"10.1016\/S0896-6273(01)00285-9","article-title":"The prefrontal cortex\u2013an update: time is of the essence.","volume":"30","author":"JM Fuster","year":"2001","journal-title":"Neuron"},{"key":"ref39","doi-asserted-by":"crossref","first-page":"223","DOI":"10.1016\/S0959-4388(02)00314-8","article-title":"Computational perspectives on dopamine function in prefrontal cortex.","volume":"12","author":"JD Cohen","year":"2002","journal-title":"Curr Opin Neurobiol"},{"key":"ref40","doi-asserted-by":"crossref","first-page":"408","DOI":"10.1152\/jn.1957.20.4.408","article-title":"Modality and topographic properties of single neurons of cat&apos;s somatic sensory cortex.","volume":"20","author":"VB Mountcastle","year":"1957","journal-title":"J Neurophysiol"},{"key":"ref41","doi-asserted-by":"crossref","first-page":"540","DOI":"10.1037\/a0016170","article-title":"Is there a geometric module for spatial orientation? insights from a rodent navigation model.","volume":"116","author":"D Sheynikhovich","year":"2009","journal-title":"Psychol Rev"},{"key":"ref42","doi-asserted-by":"crossref","first-page":"61","DOI":"10.1007\/BF00203631","article-title":"A neuron model with fluid properties for solving labyrinthian puzzle.","volume":"64","author":"G Lei","year":"1990","journal-title":"Biol Cybern"},{"key":"ref43","doi-asserted-by":"crossref","first-page":"304","DOI":"10.1109\/70.143349","article-title":"Integration of representation into goal-driven behavior-based robots.","volume":"8","author":"MJ Mataric","year":"1992","journal-title":"IEEE Trans Rob Autom"},{"key":"ref44","doi-asserted-by":"crossref","first-page":"1115","DOI":"10.1162\/0898929054475190","article-title":"A model of prefrontal cortical mechanisms for goal-directed behavior.","volume":"17","author":"ME Hasselmo","year":"2005","journal-title":"J Cogn Neurosci"},{"key":"ref45","doi-asserted-by":"crossref","first-page":"701","DOI":"10.1093\/brain\/120.4.701","article-title":"The columnar organization of the neocortex.","volume":"120","author":"VB Mountcastle","year":"1997","journal-title":"Brain"},{"key":"ref46","doi-asserted-by":"crossref","first-page":"75","DOI":"10.1016\/0006-8993(96)00437-4","article-title":"The organisation of dendritic bundles in the prelimbic cortex (area 32) of the rat.","volume":"730","author":"PL Gabbott","year":"1996","journal-title":"Brain Res"},{"key":"ref47","doi-asserted-by":"crossref","first-page":"145","DOI":"10.1002\/cne.20738","article-title":"Prefrontal cortex in the rat: projections to subcortical autonomic, motor, and limbic centers.","volume":"492","author":"PLA Gabbott","year":"2005","journal-title":"J Comp Neurol"},{"key":"ref48","doi-asserted-by":"crossref","first-page":"935","DOI":"10.1093\/brain\/awf110","article-title":"The minicolumn hypothesis in neuroscience.","volume":"125","author":"DP Buxhoeveden","year":"2002","journal-title":"Brain"},{"key":"ref49","article-title":"An adaptative neural network: the cerebral cortex.","author":"Y Burnod","year":"1988"},{"key":"ref50","doi-asserted-by":"crossref","first-page":"475","DOI":"10.1016\/0006-8993(75)90122-5","article-title":"The \u2018module-concept\u2019 in cerebral cortex architecture.","volume":"95","author":"J Szent\u00e1gothai","year":"1975","journal-title":"Brain Res"},{"key":"ref51","doi-asserted-by":"crossref","first-page":"1839","DOI":"10.1016\/0306-4522(81)90027-0","article-title":"The modular operation of the cerebral neocortex considered as the material basis of mental events.","volume":"6","author":"JC Eccles","year":"1981","journal-title":"Neuroscience"},{"key":"ref52","doi-asserted-by":"crossref","first-page":"351","DOI":"10.1007\/BF00114187","article-title":"Organizational levels of the cerebral cortex: an integrated model.","volume":"39","author":"Y Burnod","year":"1991","journal-title":"Acta Biotheor"},{"key":"ref53","doi-asserted-by":"crossref","first-page":"153","DOI":"10.1038\/nrn1848","article-title":"The blue brain project.","volume":"7","author":"H Markram","year":"2006","journal-title":"Nat Rev Neurosci"},{"key":"ref54","doi-asserted-by":"crossref","first-page":"1903","DOI":"10.1152\/jn.1999.81.4.1903","article-title":"Isodirectional tuning of adjacent interneurons and pyramidal cells during working memory: evidence for microcolumnar organization in PFC.","volume":"81","author":"SG Rao","year":"1999","journal-title":"J Neurophysiol"},{"key":"ref55","doi-asserted-by":"crossref","first-page":"1399","DOI":"10.1016\/S0896-6273(00)80658-3","article-title":"Neural activity in the primate prefrontal cortex during associative learning.","volume":"21","author":"WF Asaad","year":"1998","journal-title":"Neuron"},{"key":"ref56","doi-asserted-by":"crossref","first-page":"265","DOI":"10.1023\/A:1024174026286","article-title":"Specificity in the functional architecture of primate prefrontal cortex.","volume":"31","author":"DA Lewis","year":"2002","journal-title":"J Neurocytol"},{"key":"ref57","doi-asserted-by":"crossref","first-page":"287","DOI":"10.1007\/s004220000171","article-title":"Spatial cognition and neuro-mimetic navigation: a model of hippocampal place cell activity.","volume":"83","author":"A Arleo","year":"2000","journal-title":"Biol Cybern"},{"key":"ref58","doi-asserted-by":"crossref","first-page":"639","DOI":"10.1109\/TNN.2004.826221","article-title":"Cognitive navigation based on nonuniform gabor space sampling, unsupervised growing networks, and reinforcement learning.","volume":"15","author":"A Arleo","year":"2004","journal-title":"IEEE Trans Neural Netw"},{"key":"ref59","doi-asserted-by":"crossref","first-page":"919","DOI":"10.1038\/nn.2337","article-title":"Replay of rule-learning related neural patterns in the prefrontal cortex during sleep.","volume":"12","author":"A Peyrache","year":"2009","journal-title":"Nat Neurosci"},{"key":"ref60","doi-asserted-by":"crossref","first-page":"921","DOI":"10.1016\/j.neuron.2010.05.013","article-title":"Coherent theta oscillations and reorganization of spike timing in the hippocampal- prefrontal network upon learning.","volume":"66","author":"K Benchenane","year":"2010","journal-title":"Neuron"},{"key":"ref61","doi-asserted-by":"crossref","first-page":"629","DOI":"10.1038\/382629a0","article-title":"Reward expectancy in primate prefrontal neurons.","volume":"382","author":"M Watanabe","year":"1996","journal-title":"Nature"},{"key":"ref62","doi-asserted-by":"crossref","first-page":"5493","DOI":"10.1523\/JNEUROSCI.19-13-05493.1999","article-title":"Prospective coding for objects in primate prefrontal cortex.","volume":"19","author":"G Rainer","year":"1999","journal-title":"J Neurosci"},{"key":"ref63","doi-asserted-by":"crossref","first-page":"704","DOI":"10.1038\/19525","article-title":"Relative reward preference in primate orbitofrontal cortex.","volume":"398","author":"L Tremblay","year":"1999","journal-title":"Nature"},{"key":"ref64","doi-asserted-by":"crossref","first-page":"167","DOI":"10.1146\/annurev.neuro.24.1.167","article-title":"An integrative theory of prefrontal function.","volume":"24","author":"EK Miller","year":"2001","journal-title":"Annu Rev Neurosci"},{"key":"ref65","doi-asserted-by":"crossref","first-page":"13172","DOI":"10.1073\/pnas.162485599","article-title":"Parallel processing of serial movements in prefrontal cortex.","volume":"99","author":"BB Averbeck","year":"2002","journal-title":"Proc Natl Acad Sci U S A"},{"key":"ref66","doi-asserted-by":"crossref","first-page":"77","DOI":"10.1016\/j.bbr.2003.09.016","article-title":"Learning-related changes in response patterns of prefrontal neurons during instrumental conditioning.","volume":"146","author":"AB Mulder","year":"2003","journal-title":"Behav Brain Res"},{"key":"ref67","first-page":"1030","article-title":"An information-theoretic approach to deciphering the hippocampal code.","author":"WE Skaggs","year":"1992"},{"key":"ref68","doi-asserted-by":"crossref","first-page":"405","DOI":"10.1162\/08997660252741176","article-title":"Measuring information spatial densities.","volume":"14","author":"M Bezzi","year":"2001","journal-title":"Neural Comput"},{"key":"ref69","doi-asserted-by":"crossref","first-page":"837","DOI":"10.1098\/rstb.2005.1623","article-title":"The cortical column: a structure without a function.","volume":"360","author":"JC Horton","year":"2005","journal-title":"Philos Trans R Soc Lond B Biol Sci"},{"key":"ref70","doi-asserted-by":"crossref","first-page":"12099","DOI":"10.1073\/pnas.0807271105","article-title":"Confusing cortical columns.","volume":"105","author":"P Rakic","year":"2008","journal-title":"Proc Natl Acad Sci U S A"},{"key":"ref71","doi-asserted-by":"crossref","first-page":"357","DOI":"10.1146\/annurev.neuro.28.061604.135637","article-title":"Neural network dynamics.","volume":"28","author":"TP Vogels","year":"2005","journal-title":"Annu Rev Neurosci"},{"key":"ref72","doi-asserted-by":"crossref","first-page":"487","DOI":"10.1177\/1073858408317066","article-title":"Propagating waves of activity in the neocortex: what they are, what they do.","volume":"14","author":"J Wu","year":"2008","journal-title":"Neuroscientist"},{"key":"ref73","doi-asserted-by":"crossref","first-page":"1549","DOI":"10.1038\/nn1802","article-title":"Propagating waves mediate information transfer in the motor cortex.","volume":"9","author":"D Rubino","year":"2006","journal-title":"Nat Neurosci"},{"key":"ref74","doi-asserted-by":"crossref","first-page":"359","DOI":"10.1146\/annurev.neuro.29.051605.112851","article-title":"Habits, rituals, and the evaluative brain.","volume":"31","author":"AM Graybiel","year":"2008","journal-title":"Annu Rev Neurosci"},{"key":"ref75","doi-asserted-by":"crossref","first-page":"65","DOI":"10.1006\/nlme.1996.0007","article-title":"Inactivation of hippocampus or caudate nucleus with lidocaine differentially affects expression of place and response learning.","volume":"65","author":"MG Packard","year":"1996","journal-title":"Neurobiol Learn Mem"},{"key":"ref76","article-title":"Exploration and navigation using hierarchical cognitive maps.","author":"N Schmajuk","year":"2006"},{"key":"ref77","first-page":"1313","article-title":"Neurosolver: a step toward a neuromorphic general problem solver.","volume":"volume 3","author":"A Bieszczad","year":"1994"},{"key":"ref78","first-page":"252","article-title":"Modeling prefrontal functions for robot navigation.","volume":"volume 1","author":"H Frezza-Buet","year":"1999"},{"key":"ref79","doi-asserted-by":"crossref","first-page":"627","DOI":"10.1017\/S0140525X00013959","article-title":"Multiple representations of space underlying behavior.","volume":"5","author":"I Lieblich","year":"1982","journal-title":"Behav Brain Sci"},{"key":"ref80","doi-asserted-by":"crossref","first-page":"165","DOI":"10.1007\/BF00201023","article-title":"Purposive behavior and cognitive mapping: a neural network model.","volume":"67","author":"NA Schmajuk","year":"1992","journal-title":"Biol Cybern"},{"key":"ref81","doi-asserted-by":"crossref","first-page":"111","DOI":"10.1023\/A:1008821210922","article-title":"Learning view graphs for robot navigation.","volume":"5","author":"MO Franz","year":"1998","journal-title":"Auton Robots"},{"key":"ref82","doi-asserted-by":"crossref","first-page":"13293","DOI":"10.1073\/pnas.94.24.13293","article-title":"A hierarchical neuronal network for planning behavior.","volume":"94","author":"S Dehaene","year":"1997","journal-title":"Proc Natl Acad Sci U S A"},{"key":"ref83","doi-asserted-by":"crossref","first-page":"569","DOI":"10.1016\/S0893-6080(03)00095-9","article-title":"Hierarchical cognitive maps.","volume":"16","author":"H Voicu","year":"2003","journal-title":"Neural Netw"},{"key":"ref84","doi-asserted-by":"crossref","first-page":"1339","DOI":"10.1162\/0899766053630369","article-title":"A hierarchy of associations in hippocampo-cortical systems: cognitive maps and navigation strategies.","volume":"17","author":"JP Banquet","year":"2005","journal-title":"Neural Comput"},{"key":"ref85","doi-asserted-by":"crossref","first-page":"1987","DOI":"10.1162\/089976600300015024","article-title":"DEA: an architecture for goal planning and classification.","volume":"12","author":"F Fleuret","year":"2000","journal-title":"Neural Comput"},{"key":"ref86","doi-asserted-by":"crossref","first-page":"1964","DOI":"10.1093\/cercor\/bhi072","article-title":"An integrate-and-fire model of prefrontal cortex neuronal activity during performance of goal-directed decision making.","volume":"15","author":"RA Koene","year":"2005","journal-title":"Cereb Cortex"},{"key":"ref87","doi-asserted-by":"crossref","first-page":"211","DOI":"10.1037\/0278-7393.15.2.211","article-title":"Subjective hierarchies in spatial memory.","volume":"15","author":"TP McNamara","year":"1989","journal-title":"J Exp Psychol Learn Mem Cogn"},{"key":"ref88","doi-asserted-by":"crossref","first-page":"117","DOI":"10.1016\/0023-9690(79)90040-7","article-title":"Spatial memory in the rat on a hierarchical maze.","volume":"10","author":"WA Roberts","year":"1979","journal-title":"Learn Motiv"},{"key":"ref89","doi-asserted-by":"crossref","first-page":"69","DOI":"10.1037\/0097-7403.16.1.69","article-title":"Hierarchical structures: chunking by food type facilitates spatial memory.","volume":"16","author":"NL Dallal","year":"1990","journal-title":"J Exp Psychol Anim Behav Process"},{"key":"ref90","doi-asserted-by":"crossref","first-page":"187","DOI":"10.1037\/0097-7403.21.3.187","article-title":"Coding of hierarchical versus linear pattern structure in rats and humans.","volume":"21","author":"SB Fountain","year":"1995","journal-title":"J Exp Psychol Anim Behav Process"},{"key":"ref91","doi-asserted-by":"crossref","first-page":"20","DOI":"10.1037\/0097-7403.21.1.20","article-title":"Further evidence for hierarchical chunking in rat spatial memory.","volume":"21","author":"T Macuda","year":"1995","journal-title":"J Exp Psychol Anim Behav Process"},{"key":"ref92","doi-asserted-by":"crossref","first-page":"3053","DOI":"10.1097\/00001756-199709290-00010","article-title":"Perinatal choline supplementation increases the threshold for chunking in spatial memory.","volume":"8","author":"WH Meck","year":"1997","journal-title":"Neuroreport"},{"key":"ref93","doi-asserted-by":"crossref","first-page":"140","DOI":"10.1126\/science.1157086","article-title":"Finite scale of spatial representation in the hippocampus.","volume":"321","author":"KB Kjelstrup","year":"2008","journal-title":"Science"},{"key":"ref94","doi-asserted-by":"crossref","first-page":"262","DOI":"10.1016\/j.cognition.2008.08.011","article-title":"Hierarchically organized behavior and its neural foundations: a reinforcement learning perspective.","volume":"113","author":"MM Botvinick","year":"2009","journal-title":"Cognition"},{"key":"ref95","doi-asserted-by":"crossref","first-page":"181","DOI":"10.1016\/S0004-3702(99)00052-1","article-title":"Between MDPs and semi-MDPs: a framework for temporal abstraction in reinforcement learning.","volume":"112","author":"RS Sutton","year":"1999","journal-title":"Artif Intell"},{"key":"ref96","doi-asserted-by":"crossref","first-page":"1181","DOI":"10.1126\/science.1088545","article-title":"The architecture of cognitive control in the human prefrontal cortex.","volume":"302","author":"E Koechlin","year":"2003","journal-title":"Science"},{"key":"ref97","doi-asserted-by":"crossref","first-page":"145","DOI":"10.1016\/j.neuron.2006.02.023","article-title":"Temporal encoding of place sequences by hippocampal cell assemblies.","volume":"50","author":"G Dragoi","year":"2006","journal-title":"Neuron"},{"key":"ref98","doi-asserted-by":"crossref","first-page":"4071","DOI":"10.1523\/JNEUROSCI.3408-05.2006","article-title":"Impaired sequential egocentric and allocentric memories in forebrain-specific-NMDA receptor knock-out mice during a new task dissociating strategies of navigation.","volume":"26","author":"L Rondi-Reig","year":"2006","journal-title":"J Neurosci"},{"key":"ref99","doi-asserted-by":"crossref","first-page":"1123","DOI":"10.1093\/cercor\/bhh211","article-title":"Prefrontal-hippocampal dynamics involved in learning regularities across episodes.","volume":"15","author":"CF Doeller","year":"2005","journal-title":"Cereb Cortex"},{"key":"ref100","doi-asserted-by":"crossref","first-page":"1802","DOI":"10.1016\/j.neuroimage.2006.02.006","article-title":"Differential hippocampal and prefrontal-striatal contributions to instance-based and rule-based learning.","volume":"31","author":"CF Doeller","year":"2006","journal-title":"Neuroimage"},{"key":"ref101","doi-asserted-by":"crossref","first-page":"555","DOI":"10.1038\/nn1880","article-title":"Memory consolidation or transformation: context manipulation and hippocampal representations of memory.","volume":"10","author":"G Winocur","year":"2007","journal-title":"Nat Neurosci"},{"key":"ref102","doi-asserted-by":"crossref","first-page":"1407","DOI":"10.1093\/cercor\/bhm172","article-title":"Dopamine modulation of hippocampal-prefrontal cortical interaction drives memory-guided behavior.","volume":"18","author":"Y Goto","year":"2008","journal-title":"Cereb Cortex"},{"key":"ref103","doi-asserted-by":"crossref","first-page":"39","DOI":"10.1196\/annals.1440.001","article-title":"Episodic simulation of future events: concepts, data, and applications.","volume":"1124","author":"DL Schacter","year":"2008","journal-title":"Ann N Y Acad Sci"},{"key":"ref104","doi-asserted-by":"crossref","first-page":"631","DOI":"10.1016\/j.neuron.2006.03.045","article-title":"Activity in the lateral prefrontal cortex reflects multiple steps of future events in action plans.","volume":"50","author":"H Mushiake","year":"2006","journal-title":"Neuron"},{"key":"ref105","doi-asserted-by":"crossref","first-page":"707","DOI":"10.1016\/S0896-6273(00)81072-7","article-title":"Experience-Dependent asymmetric shape of hippocampal receptive fields.","volume":"25","author":"M Mehta","year":"2000","journal-title":"Neuron"},{"key":"ref106","doi-asserted-by":"crossref","first-page":"1227","DOI":"10.1016\/S0896-6273(03)00752-9","article-title":"Prospective and retrospective memory coding in the hippocampus.","volume":"40","author":"J Ferbinteanu","year":"2003","journal-title":"Neuron"},{"key":"ref107","doi-asserted-by":"crossref","first-page":"821","DOI":"10.1162\/089892903322370744","article-title":"A computational model of prefrontal control in free recall: strategic memory use in the california verbal learning task.","volume":"15","author":"S Becker","year":"2003","journal-title":"J Cogn Neurosci"},{"key":"ref108","doi-asserted-by":"crossref","first-page":"340","DOI":"10.1037\/0033-295X.114.2.340","article-title":"Remembering the past and imagining the future: a neural model of spatial memory and imagery.","volume":"114","author":"P Byrne","year":"2007","journal-title":"Psychol Rev"},{"key":"ref109","doi-asserted-by":"crossref","first-page":"189","DOI":"10.1037\/h0061626","article-title":"Cognitive maps in rats and men.","volume":"55","author":"EC Tolman","year":"1948","journal-title":"Psychol Rev"},{"key":"ref110","doi-asserted-by":"crossref","first-page":"9","DOI":"10.1085\/jgp.200409015","article-title":"Representation of objects in space by two classes of hippocampal pyramidal cells.","volume":"124","author":"B Rivard","year":"2004","journal-title":"J Gen Physiol"},{"key":"ref111","doi-asserted-by":"crossref","first-page":"7324","DOI":"10.1523\/JNEUROSCI.1909-08.2008","article-title":"Different CA1 and CA3 representations of novel routes in a shortcut situation.","volume":"28","author":"A Alvernhe","year":"2008","journal-title":"J Neurosci"},{"key":"ref112","doi-asserted-by":"crossref","first-page":"12176","DOI":"10.1523\/JNEUROSCI.3761-07.2007","article-title":"Neural ensembles in CA3 transiently encode paths forward of the animal at a decision point.","volume":"27","author":"A Johnson","year":"2007","journal-title":"J Neurosci"},{"key":"ref113","doi-asserted-by":"crossref","first-page":"317","DOI":"10.1002\/hipo.450030307","article-title":"Phase relationship between hippocampal place units and the EEG theta rhythm.","volume":"3","author":"J O'Keefe","year":"1993","journal-title":"Hippocampus"},{"key":"ref114","doi-asserted-by":"crossref","first-page":"e402","DOI":"10.1371\/journal.pbio.0030402","article-title":"Theta rhythms coordinate hippocampal-prefrontal interactions in a spatial memory task.","volume":"3","author":"MW Jones","year":"2005","journal-title":"PLoS Biol"},{"key":"ref115","doi-asserted-by":"crossref","first-page":"1161","DOI":"10.1038\/nn1756","article-title":"Separate neural pathways process different decision costs.","volume":"9","author":"PH Rudebeck","year":"2006","journal-title":"Nat Neurosci"},{"key":"ref116","doi-asserted-by":"crossref","first-page":"1065","DOI":"10.1016\/S0893-6080(05)80159-5","article-title":"A model of hippocampal function.","volume":"7","author":"N Burgess","year":"1994","journal-title":"Neural Netw"},{"key":"ref117","doi-asserted-by":"crossref","first-page":"1","DOI":"10.1002\/(SICI)1098-1063(2000)10:1<1::AID-HIPO1>3.0.CO;2-1","article-title":"A model of hippocampally dependent navigation, using the temporal difference learning rule.","volume":"10","author":"DJ Foster","year":"2000","journal-title":"Hippocampus"},{"key":"ref118","doi-asserted-by":"crossref","first-page":"1222","DOI":"10.1038\/nn.2384","article-title":"Selective suppression of hippocampal ripples impairs spatial memory.","volume":"12","author":"G Girardeau","year":"2009","journal-title":"Nat Neurosci"}],"container-title":["PLoS Computational Biology"],"original-title":[],"language":"en","link":[{"URL":"http:\/\/dx.plos.org\/10.1371\/journal.pcbi.1002045","content-type":"unspecified","content-version":"vor","intended-application":"similarity-checking"}],"deposited":{"date-parts":[[2019,6,10]],"date-time":"2019-06-10T23:53:41Z","timestamp":1560210821000},"score":1,"resource":{"primary":{"URL":"https:\/\/dx.plos.org\/10.1371\/journal.pcbi.1002045"}},"subtitle":[],"editor":[{"given":"Olaf","family":"Sporns","sequence":"first","affiliation":[]}],"short-title":[],"issued":{"date-parts":[[2011,5,19]]},"references-count":118,"journal-issue":{"issue":"5","published-online":{"date-parts":[[2011,5,19]]}},"URL":"https:\/\/doi.org\/10.1371\/journal.pcbi.1002045","relation":{},"ISSN":["1553-7358"],"issn-type":[{"value":"1553-7358","type":"electronic"}],"subject":[],"published":{"date-parts":[[2011,5,19]]}}}