{"status":"ok","message-type":"work","message-version":"1.0.0","message":{"indexed":{"date-parts":[[2026,3,3]],"date-time":"2026-03-03T07:59:02Z","timestamp":1772524742622,"version":"3.50.1"},"reference-count":61,"publisher":"Public Library of Science (PLoS)","issue":"1","license":[{"start":{"date-parts":[[2015,1,15]],"date-time":"2015-01-15T00:00:00Z","timestamp":1421280000000},"content-version":"vor","delay-in-days":0,"URL":"https:\/\/creativecommons.org\/publicdomain\/zero\/1.0\/"}],"content-domain":{"domain":["www.ploscompbiol.org"],"crossmark-restriction":false},"short-container-title":["PLoS Comput Biol"],"DOI":"10.1371\/journal.pcbi.1004043","type":"journal-article","created":{"date-parts":[[2015,1,15]],"date-time":"2015-01-15T20:47:20Z","timestamp":1421354840000},"page":"e1004043","update-policy":"https:\/\/doi.org\/10.1371\/journal.pcbi.corrections_policy","source":"Crossref","is-referenced-by-count":67,"title":["Synaptic Plasticity Enables Adaptive Self-Tuning Critical Networks"],"prefix":"10.1371","volume":"11","author":[{"given":"Nigel","family":"Stepp","sequence":"first","affiliation":[]},{"given":"Dietmar","family":"Plenz","sequence":"additional","affiliation":[]},{"given":"Narayan","family":"Srinivasa","sequence":"additional","affiliation":[]}],"member":"340","published-online":{"date-parts":[[2015,1,15]]},"reference":[{"key":"ref1","doi-asserted-by":"crossref","first-page":"15921","DOI":"10.1073\/pnas.0904089106","article-title":"Spontaneous cortical activity in awake monkeys composed of neuronal avalanches","volume":"106","author":"T Petermann","year":"2009","journal-title":"Proc Natl Acad Sci"},{"key":"ref2","doi-asserted-by":"crossref","first-page":"123","DOI":"10.1038\/nature09086","article-title":"Sensitivity to perturbations in vivo implies high noise and suggests rate coding in cortex","volume":"466","author":"M London","year":"2010","journal-title":"Nature"},{"key":"ref3","doi-asserted-by":"crossref","first-page":"3870","DOI":"10.1523\/JNEUROSCI.18-10-03870.1998","article-title":"The variable discharge of cortical neurons: Implications for connectivity, computation, and information coding","volume":"18","author":"MN Shadlen","year":"1998","journal-title":"J Neurosci"},{"key":"ref4","doi-asserted-by":"crossref","first-page":"11167","DOI":"10.1523\/JNEUROSCI.23-35-11167.2003","article-title":"Neuronal avalanches in neocortical circuits","volume":"23","author":"JM Beggs","year":"2003","journal-title":"J Neurosci"},{"key":"ref5","doi-asserted-by":"crossref","first-page":"55","DOI":"10.1523\/JNEUROSCI.4637-10.2011","article-title":"Information capacity and transmission are maximized in balanced cortical networks with neuronal avalanches","volume":"31","author":"WL Shew","year":"2011","journal-title":"J Neurosci"},{"key":"ref6","doi-asserted-by":"crossref","first-page":"1061","DOI":"10.1523\/JNEUROSCI.2771-11.2012","article-title":"Maximal variability of phase synchrony in cortical networks with neuronal avalanches","volume":"32","author":"H Yang","year":"2012","journal-title":"J Neurosci"},{"key":"ref7","doi-asserted-by":"crossref","DOI":"10.3389\/fphys.2012.00015","article-title":"Criticality in large-scale brain fMRI dynamics unveiled by a novel point process analysis","volume":"3","author":"E Tagliazucchi","year":"2012","journal-title":"Frontiers in Physiology"},{"key":"ref8","doi-asserted-by":"crossref","first-page":"1370","DOI":"10.1523\/JNEUROSCI.21-04-01370.2001","article-title":"Long-range temporal correlations and scaling behavior in human brain oscillations","volume":"21","author":"K Linkenkaer-Hansen","year":"2001","journal-title":"J Neurosci"},{"key":"ref9","doi-asserted-by":"crossref","first-page":"1413","DOI":"10.1162\/089976604323057443","article-title":"Real-time computation at the edge of chaos in recurrent neural networks","volume":"16","author":"N Bertschinger","year":"2004","journal-title":"Neural Comput"},{"key":"ref10","doi-asserted-by":"crossref","first-page":"58101","DOI":"10.1103\/PhysRevLett.94.058101","article-title":"Critical branching captures activity in living neural networks and maximizes the number of metastable states","volume":"94","author":"C Haldeman","year":"2005","journal-title":"Phys Rev Lett"},{"key":"ref11","doi-asserted-by":"crossref","first-page":"8148","DOI":"10.1523\/JNEUROSCI.0723-06.2006","article-title":"Inverted-u profile of dopamine\u2013nmda-mediated spontaneous avalanche recurrence in superficial layers of rat prefrontal cortex","volume":"26","author":"C Stewart","year":"2006","journal-title":"The Journal of neuroscience"},{"key":"ref12","doi-asserted-by":"crossref","first-page":"348","DOI":"10.1038\/nphys289","article-title":"Optimal dynamical range of excitable networks at criticality","volume":"2","author":"O Kinouchi","year":"2006","journal-title":"Nature Phys"},{"key":"ref13","doi-asserted-by":"crossref","first-page":"15595","DOI":"10.1523\/JNEUROSCI.3864-09.2009","article-title":"Neuronal avalanches imply maximum dynamic range in cortical networks at criticality","volume":"29","author":"WL Shew","year":"2009","journal-title":"J Neurosci"},{"key":"ref14","doi-asserted-by":"crossref","first-page":"e42772","DOI":"10.1371\/journal.pone.0042772","article-title":"Neuromorphic atomic switch networks","volume":"7","author":"AV Avizienis","year":"2012","journal-title":"PloS one"},{"key":"ref15","doi-asserted-by":"crossref","first-page":"51","DOI":"10.1109\/MPUL.2011.2175639","article-title":"Neuromorphic adaptive plastic scalable electronics: Analog learning systems","volume":"3","author":"N Srinivasa","year":"2012","journal-title":"IEEE Pulse"},{"key":"ref16","doi-asserted-by":"crossref","first-page":"857","DOI":"10.1038\/nphys758","article-title":"Dynamical synapses causing self-organized criticality in neural networks","volume":"3","author":"A Levina","year":"2007","journal-title":"Nature Phys"},{"key":"ref17","doi-asserted-by":"crossref","first-page":"801","DOI":"10.1038\/nphys1757","article-title":"Self-organized criticality occurs in nonconservative neuronal networks during \u2018up\u2019 states","volume":"6","author":"D Millman","year":"2010","journal-title":"Nature Phys"},{"key":"ref18","doi-asserted-by":"crossref","first-page":"3977","DOI":"10.1073\/pnas.0912289107","article-title":"Learning as a phenomenon occurring in a critical state","volume":"107","author":"L De Arcangelis","year":"2010","journal-title":"Proc Natl Acad Sci"},{"key":"ref19","doi-asserted-by":"crossref","first-page":"258102","DOI":"10.1103\/PhysRevLett.102.258102","article-title":"Self-tuned critical anti-hebbian networks","volume":"102","author":"MO Magnasco","year":"2009","journal-title":"Phys Rev Lett"},{"key":"ref20","doi-asserted-by":"crossref","first-page":"2758","DOI":"10.1523\/JNEUROSCI.13-07-02758.1993","article-title":"How independent are the messages carried by adjacent inferior temporal cortical neurons?","volume":"13","author":"TJ Gawne","year":"1993","journal-title":"J Neurosci"},{"key":"ref21","doi-asserted-by":"crossref","first-page":"4079","DOI":"10.1152\/jn.01357.2006","article-title":"Self-tuning of neural circuits through short-term synaptic plasticity","volume":"97","author":"D Sussillo","year":"2007","journal-title":"J Neurophysiol"},{"key":"ref22","doi-asserted-by":"crossref","first-page":"1569","DOI":"10.1126\/science.1211095","article-title":"Inhibitory plasticity balances excitation and inhibition in sensory pathways and memory networks","volume":"334","author":"TP Vogels","year":"2011","journal-title":"Science"},{"key":"ref23","doi-asserted-by":"crossref","first-page":"286","DOI":"10.1002\/adma.201103053","article-title":"Emergent criticality in complex turing b-type atomic switch networks","volume":"24","author":"AZ Stieg","year":"2012","journal-title":"Adv Mater"},{"key":"ref24","doi-asserted-by":"crossref","first-page":"389","DOI":"10.1021\/nl203687n","article-title":"A functional hybrid memristor crossbar-Array\/CMOS system for data storage and neuromorphic applications","volume":"12","author":"KH Kim","year":"2012","journal-title":"Nano Lett"},{"key":"ref25","article-title":"HRLSim: A high performance spiking neural network simulator for GPGPU clusters","author":"K Minkovich","year":"2013"},{"key":"ref26","doi-asserted-by":"crossref","first-page":"384011","DOI":"10.1088\/0957-4484\/24\/38\/384011","article-title":"A scalable neural chip with synaptic electronics using CMOS integrated memristors","volume":"24","author":"JM Cruz-Albrecht","year":"2013","journal-title":"Nanotechnology"},{"key":"ref27","doi-asserted-by":"crossref","first-page":"17514","DOI":"10.1523\/JNEUROSCI.3127-11.2011","article-title":"Higher-order interactions characterized in cortical activity","volume":"31","author":"S Yu","year":"2011","journal-title":"J Neurosci"},{"key":"ref28","doi-asserted-by":"crossref","first-page":"587","DOI":"10.1126\/science.1179850","article-title":"The asynchronous state in cortical circuits","volume":"327","author":"A Renart","year":"2010","journal-title":"Science"},{"key":"ref29","doi-asserted-by":"crossref","first-page":"584","DOI":"10.1126\/science.1179867","article-title":"Decorrelated neuronal firing in cortical microcircuits","volume":"327","author":"AS Ecker","year":"2010","journal-title":"Science"},{"key":"ref30","doi-asserted-by":"crossref","first-page":"863","DOI":"10.1103\/PhysRevLett.78.863","article-title":"Description of a turbulent cascade by a fokker-planck equation","volume":"78","author":"R Friedrich","year":"1997","journal-title":"Physical Review Letters"},{"key":"ref31","doi-asserted-by":"crossref","first-page":"190","DOI":"10.1016\/j.physleta.2006.08.003","article-title":"A data analysis method for identifying deterministic components of stable and unstable time-delayed systems with colored noise","volume":"360","author":"K Patanarapeelert","year":"2006","journal-title":"Physics Letters A"},{"key":"ref32","doi-asserted-by":"crossref","first-page":"230","DOI":"10.1037\/a0030970","article-title":"Critical branching neural networks","volume":"120","author":"CT Kello","year":"2013","journal-title":"Psychological review"},{"key":"ref33","doi-asserted-by":"crossref","first-page":"441","DOI":"10.1016\/S0378-4371(01)00144-3","article-title":"Detecting long-range correlations with detrended fluctuation analysis","volume":"295","author":"JW Kantelhardt","year":"2001","journal-title":"Physica A"},{"key":"ref34","doi-asserted-by":"crossref","DOI":"10.3389\/fncir.2013.00119","article-title":"Inhibitory synaptic plasticity: spike timing-dependence and putative network function","volume":"7","author":"TP Vogels","year":"2013","journal-title":"Frontiers in neural circuits"},{"key":"ref35","doi-asserted-by":"crossref","first-page":"594","DOI":"10.1038\/nn.3658","article-title":"Two types of asynchronous activity in networks of excitatory and inhibitory spiking neurons","volume":"17","author":"S Ostojic","year":"2014","journal-title":"Nature neuroscience"},{"key":"ref36","doi-asserted-by":"crossref","first-page":"436","DOI":"10.1037\/a0019098","article-title":"Interaction-dominant dynamics in human cognition: Beyond 1\/\u03b1 fluctuation","volume":"139","author":"EA Ihlen","year":"2010","journal-title":"Journal of Experimental Psychology: General"},{"key":"ref37","doi-asserted-by":"crossref","first-page":"e1003258","DOI":"10.1371\/journal.pcbi.1003258","article-title":"Predictive coding of dynamical variables in balanced spiking networks","volume":"9","author":"M Boerlin","year":"2013","journal-title":"PLoS computational biology"},{"key":"ref38","doi-asserted-by":"crossref","first-page":"1437","DOI":"10.1162\/neco.2007.19.6.1437","article-title":"Spike-timing-dependent plasticity in balanced random networks","volume":"19","author":"A Morrison","year":"2007","journal-title":"Neural computation"},{"key":"ref39","article-title":"Self-organization in nonequilibrium systems","author":"G Nicolis","year":"1977"},{"key":"ref40","doi-asserted-by":"crossref","first-page":"1400","DOI":"10.1103\/PhysRevLett.47.1400","article-title":"Diffusion-limited aggregation, a kinetic critical phenomenon","volume":"47","author":"T Witten Jr","year":"1981","journal-title":"Physical review letters"},{"key":"ref41","doi-asserted-by":"crossref","first-page":"381","DOI":"10.1103\/PhysRevLett.59.381","article-title":"Self-organized criticality: An explanation of the 1\/f noise","volume":"59","author":"P Bak","year":"1987","journal-title":"Phys Rev Lett"},{"key":"ref42","doi-asserted-by":"crossref","DOI":"10.3389\/fncom.2013.00010","article-title":"Stable learning of functional maps in self-organizing spiking neural networks with continuous synaptic plasticity","volume":"7","author":"N Srinivasa","year":"2013","journal-title":"Frontiers in computational neuroscience"},{"key":"ref43","doi-asserted-by":"crossref","first-page":"e1002334","DOI":"10.1371\/journal.pcbi.1002334","article-title":"Balancing feed-forward excitation and inhibition via hebbian inhibitory synaptic plasticity","volume":"8","author":"Y Luz","year":"2012","journal-title":"PLoS computational biology"},{"key":"ref44","doi-asserted-by":"crossref","DOI":"10.3389\/fncom.2014.00053","article-title":"Excitatory and inhibitory stdp jointly tune feedforward neural circuits to selectively propagate correlated spiking activity","volume":"8","author":"FI Kleberg","year":"2014","journal-title":"Frontiers in computational neuroscience"},{"key":"ref45","doi-asserted-by":"crossref","DOI":"10.3389\/fncom.2011.00047","article-title":"Synaptic scaling in combination with many generic plasticity mechanisms stabilizes circuit connectivity","volume":"5","author":"C Tetzlaff","year":"2011","journal-title":"Frontiers in computational neuroscience"},{"key":"ref46","doi-asserted-by":"crossref","DOI":"10.3389\/neuro.10.005.2007","article-title":"Irregular persistent activity induced by synaptic excitatory feedback","volume":"1","author":"F Barbieri","year":"2007","journal-title":"Frontiers in Computational Neuroscience"},{"key":"ref47","doi-asserted-by":"crossref","first-page":"807","DOI":"10.1038\/382807a0","article-title":"Redistribution of synaptic efficacy between neocortical pyramidal neurons","volume":"382","author":"H Markram","year":"1996","journal-title":"Nature"},{"key":"ref48","doi-asserted-by":"crossref","first-page":"821","DOI":"10.1162\/089976698300017502","article-title":"Neural networks with dynamic synapses","volume":"10","author":"M Tsodyks","year":"1998","journal-title":"Neural Comput"},{"key":"ref49","first-page":"464","article-title":"Activity-induced synaptic modifications in hippocampal culture, dependence on spike timing, synaptic strength and cell type","volume":"18","author":"GQ Bi","year":"1998","journal-title":"J Neurosci"},{"key":"ref50","doi-asserted-by":"crossref","first-page":"213","DOI":"10.1126\/science.275.5297.213","article-title":"Regulation of synaptic efficacy by coincidence of postsynaptic APs and EPSPs","volume":"275","author":"H Markram","year":"1997","journal-title":"Science"},{"key":"ref51","doi-asserted-by":"crossref","first-page":"887","DOI":"10.1038\/nn1913","article-title":"Cortical reorganization consistent with spike timing\u2013but not correlation-dependent plasticity","volume":"10","author":"JM Young","year":"2007","journal-title":"Nature Neurosci"},{"key":"ref52","doi-asserted-by":"crossref","first-page":"919","DOI":"10.1038\/78829","article-title":"Competitive hebbian learning through spike-timingdependent synaptic plasticity","volume":"3","author":"S Song","year":"2000","journal-title":"Nature Neurosci"},{"key":"ref53","article-title":"Plasticity of excitatory-inhibitory balance in the auditory cortex","author":"JA D\u2019Amour","year":"2013"},{"key":"ref54","doi-asserted-by":"crossref","first-page":"357","DOI":"10.1146\/annurev.neuro.28.061604.135637","article-title":"Neural network dynamics","volume":"28","author":"TP Vogels","year":"2005","journal-title":"Annu Rev Neurosci"},{"key":"ref55","doi-asserted-by":"crossref","first-page":"483","DOI":"10.1038\/nn.2276","article-title":"Gating multiple signals through detailed balance of excitation and inhibition in spiking networks","volume":"12","author":"TP Vogels","year":"2009","journal-title":"Nature neuroscience"},{"key":"ref56","doi-asserted-by":"crossref","first-page":"3312","DOI":"10.1152\/jn.00953.2009","article-title":"Neuronal avalanches in spontaneous activity in vivo","volume":"104","author":"G Hahn","year":"2010","journal-title":"J Neurophysiol"},{"key":"ref57","doi-asserted-by":"crossref","first-page":"40","DOI":"10.1186\/1471-2202-10-40","article-title":"Subsampling effects in neuronal avalanche distributions recorded in vivo","volume":"10","author":"V Priesemann","year":"2009","journal-title":"BMC Neurosci"},{"key":"ref58","doi-asserted-by":"crossref","first-page":"82","DOI":"10.1063\/1.166141","article-title":"Quantification of scaling exponents and crossover phenomena in nonstationary heartbeat time series","volume":"5","author":"CK Peng","year":"1995","journal-title":"Chaos: An Interdisciplinary Journal of Nonlinear Science"},{"key":"ref59","doi-asserted-by":"crossref","first-page":"785","DOI":"10.1142\/S0218348X95000692","article-title":"Estimators for long-range dependence: An empirical study","volume":"3","author":"MS Taqqu","year":"1995","journal-title":"Fractals"},{"key":"ref60","doi-asserted-by":"crossref","first-page":"11114","DOI":"10.1103\/PhysRevE.64.011114","article-title":"Effect of trends on detrended fluctuation analysis","volume":"64","author":"K Hu","year":"2001","journal-title":"Phys Rev E"},{"key":"ref61","doi-asserted-by":"crossref","first-page":"495","DOI":"10.5194\/npg-11-495-2004","article-title":"Tempting long-memory \u2014 on the interpretation of DFA results","volume":"11","author":"D Maraun","year":"2004","journal-title":"Nonlinear Processes in Geophysics"}],"container-title":["PLOS Computational Biology"],"original-title":[],"language":"en","link":[{"URL":"http:\/\/dx.plos.org\/10.1371\/journal.pcbi.1004043","content-type":"unspecified","content-version":"vor","intended-application":"similarity-checking"}],"deposited":{"date-parts":[[2019,8,19]],"date-time":"2019-08-19T19:20:30Z","timestamp":1566242430000},"score":1,"resource":{"primary":{"URL":"https:\/\/dx.plos.org\/10.1371\/journal.pcbi.1004043"}},"subtitle":[],"editor":[{"given":"Wolfgang","family":"Einh\u00e4user","sequence":"first","affiliation":[]}],"short-title":[],"issued":{"date-parts":[[2015,1,15]]},"references-count":61,"journal-issue":{"issue":"1","published-online":{"date-parts":[[2015,1,15]]}},"URL":"https:\/\/doi.org\/10.1371\/journal.pcbi.1004043","relation":{},"ISSN":["1553-7358"],"issn-type":[{"value":"1553-7358","type":"electronic"}],"subject":[],"published":{"date-parts":[[2015,1,15]]}}}