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Coherence","volume":"4","author":"PH Tiesinga","year":"2010","journal-title":"Front Hum Neurosci"},{"key":"ref19","doi-asserted-by":"crossref","first-page":"211","DOI":"10.1016\/S0167-8760(00)00166-5","article-title":"Oscillatory gamma activity in humans: a possible role for object representation","volume":"38","author":"O Bertrand","year":"2000","journal-title":"Int J Psychophysiol"},{"key":"ref20","unstructured":"Brunet N, Bosman C a, Roberts M, Oostenveld R, Womelsdorf T, et al. (2013) Visual Cortical Gamma-Band Activity During Free Viewing of Natural Images. Cereb Cortex: 1\u20139. Available: <ext-link xmlns:xlink=\"http:\/\/www.w3.org\/1999\/xlink\" ext-link-type=\"uri\" xlink:href=\"http:\/\/www.ncbi.nlm.nih.gov\/pubmed\/24108806\" xlink:type=\"simple\">http:\/\/www.ncbi.nlm.nih.gov\/pubmed\/24108806<\/ext-link>. <object-id pub-id-type=\"pmid\">24108806<\/object-id>"},{"key":"ref21","doi-asserted-by":"crossref","first-page":"128","DOI":"10.1006\/ccog.1999.0389","article-title":"Temporal binding, binocular rivalry, and consciousness","volume":"8","author":"AK Engel","year":"1999","journal-title":"Conscious Cogn"},{"key":"ref22","doi-asserted-by":"crossref","first-page":"875","DOI":"10.1016\/j.neuron.2012.06.037","article-title":"Attentional Stimulus Selection through Selective Synchronization between Monkey Visual Areas","volume":"75","author":"CA Bosman","year":"2012","journal-title":"Neuron"},{"key":"ref23","doi-asserted-by":"crossref","first-page":"317","DOI":"10.1016\/j.tins.2007.05.001","article-title":"Human gamma-frequency oscillations associated with 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across visual cortex restrict their possible use in computation","volume":"67","author":"S Ray","year":"2010","journal-title":"Neuron"},{"key":"ref44","doi-asserted-by":"crossref","first-page":"375","DOI":"10.1016\/j.cub.2008.02.023","article-title":"Phase-of-firing coding of natural visual stimuli in primary visual cortex","volume":"18","author":"MA Montemurro","year":"2008","journal-title":"Curr Biol"},{"key":"ref45","doi-asserted-by":"crossref","first-page":"769","DOI":"10.1113\/jphysiol.2008.164111","article-title":"Neuronal oscillations and the rate-to-phase transform: mechanism, model and mutual information","volume":"587","author":"D McLelland","year":"2009","journal-title":"J Physiol"},{"key":"ref46","doi-asserted-by":"crossref","first-page":"293","DOI":"10.1162\/neco.1990.2.3.293","article-title":"Feature Linking via Synchronization among Distributed Assemblies: Simulations of Results from Cat Visual Cortex","volume":"2","author":"R Eckhorn","year":"1990","journal-title":"Neural Comput"},{"key":"ref47","doi-asserted-by":"crossref","first-page":"155","DOI":"10.1162\/neco.1991.3.2.155","article-title":"Stimulus-Dependent Assembly Formation of Oscillatory Responses: I. Synchronization","volume":"3","author":"P K\u00f6nig","year":"1991","journal-title":"Neural Comput"},{"key":"ref48","doi-asserted-by":"crossref","first-page":"333","DOI":"10.1016\/j.neuron.2005.03.002","article-title":"Does neuronal synchrony underlie visual feature grouping?","volume":"46","author":"BJA Palanca","year":"2005","journal-title":"Neuron"},{"key":"ref49","doi-asserted-by":"crossref","first-page":"464","DOI":"10.1109\/72.761705","article-title":"Neural mechanisms of scene segmentation: recordings from the visual cortex suggest basic circuits for linking field models","volume":"10","author":"R Eckhorn","year":"1999","journal-title":"IEEE Trans Neural Netw"},{"key":"ref50","unstructured":"Hermes D, Miller KJ, Wandell BA, Winawer J (2014) Stimulus Dependence of Gamma Oscillations in Human Visual Cortex. Cereb Cortex: bhu091\u2013. Available: <ext-link xmlns:xlink=\"http:\/\/www.w3.org\/1999\/xlink\" ext-link-type=\"uri\" xlink:href=\"http:\/\/cercor.oxfordjournals.org\/content\/early\/2014\/05\/22\/cercor.bhu091.short\" xlink:type=\"simple\">http:\/\/cercor.oxfordjournals.org\/content\/early\/2014\/05\/22\/cercor.bhu091.short<\/ext-link>. Accessed 21 August 2014. <object-id pub-id-type=\"pmid\">24855114<\/object-id>"},{"key":"ref51","doi-asserted-by":"crossref","first-page":"1804","DOI":"10.1103\/PhysRevLett.76.1804","article-title":"Phase synchronization of chaotic oscillators","volume":"76","author":"M Rosenblum","year":"1996","journal-title":"Phys Rev Lett"},{"key":"ref52","doi-asserted-by":"crossref","first-page":"85","DOI":"10.1016\/S0303-2647(98)00053-7","article-title":"Thalamo-cortical interactions modeled by weakly connected oscillators: could the brain use FM radio principles?","volume":"48","author":"FC Hoppensteadt","year":"1998","journal-title":"Biosystems"},{"key":"ref53","doi-asserted-by":"crossref","first-page":"117","DOI":"10.1007\/s004220050279","article-title":"Synaptic organizations and dynamical properties of weakly connected neural oscillators. I. Analysis of a canonical model","volume":"75","author":"FC Hoppensteadt","year":"1996","journal-title":"Biol Cybern"},{"key":"ref54","doi-asserted-by":"crossref","first-page":"33","DOI":"10.1016\/S0896-6273(01)00178-7","article-title":"Traveling electrical waves in cortex: insights from phase dynamics and speculation on a computational role","volume":"29","author":"GB Ermentrout","year":"2001","journal-title":"Neuron"},{"key":"ref55","doi-asserted-by":"crossref","first-page":"655","DOI":"10.1119\/1.1475332","article-title":"Synchronization: A Universal Concept in Nonlinear Science","volume":"70","author":"A Pikovsky","year":"2002","journal-title":"Am J Phys"},{"key":"ref56","doi-asserted-by":"crossref","first-page":"190","DOI":"10.3389\/fnhum.2010.00190","article-title":"Generative models of cortical oscillations: neurobiological implications of the kuramoto model","volume":"4","author":"M Breakspear","year":"2010","journal-title":"Front Hum Neurosci"},{"key":"ref57","doi-asserted-by":"crossref","first-page":"173","DOI":"10.1007\/s10827-006-0006-6","article-title":"Generation of theta oscillations by weakly coupled neural oscillators in the presence of noise","volume":"22","author":"MHK Bendels","year":"2007","journal-title":"J Comput Neurosci"},{"key":"ref58","doi-asserted-by":"crossref","first-page":"158101","DOI":"10.1103\/PhysRevLett.94.158101","article-title":"Efficient Estimation of Phase-Resetting Curves in Real Neurons and its Significance for Neural-Network Modeling","volume":"94","author":"R Gal\u00e1n","year":"2005","journal-title":"Phys Rev Lett"},{"key":"ref59","doi-asserted-by":"crossref","first-page":"2093","DOI":"10.1162\/neco.2007.19.8.2093","article-title":"Oscillatory neural networks with self-organized segmentation of overlapping patterns","volume":"19","author":"T Burwick","year":"2007","journal-title":"Neural Comput"},{"key":"ref60","doi-asserted-by":"crossref","first-page":"1796","DOI":"10.1162\/neco.2008.09-06-342","article-title":"Temporal coding: assembly formation through constructive interference","volume":"20","author":"T Burwick","year":"2008","journal-title":"Neural Comput"},{"key":"ref61","unstructured":"Schwemmer MA, Lewis TJ (2012) Phase Response Curves in Neuroscience. Phase Response Curves in Neuroscience. pp. 3\u201331. Available: <ext-link xmlns:xlink=\"http:\/\/www.w3.org\/1999\/xlink\" ext-link-type=\"uri\" xlink:href=\"http:\/\/www.springerlink.com\/index\/10.1007\/978-1-4614-0739-3\" xlink:type=\"simple\">http:\/\/www.springerlink.com\/index\/10.1007\/978-1-4614-0739-3<\/ext-link>."},{"key":"ref62","first-page":"2086","article-title":"Mode locking and Arnold tongues in integrate-and-fire neural oscillators","volume":"60","author":"S Coombes","year":"1999","journal-title":"Phys Rev E Stat Phys Plasmas Fluids Relat Interdiscip Topics"},{"key":"ref63","doi-asserted-by":"crossref","first-page":"1282","DOI":"10.1073\/pnas.98.3.1282","article-title":"What is a moment? Transient synchrony as a collective mechanism for spatiotemporal integration","volume":"98","author":"JJ Hopfield","year":"2001","journal-title":"Proc Natl Acad Sci U S A"},{"key":"ref64","doi-asserted-by":"crossref","first-page":"8422","DOI":"10.1073\/pnas.0803183105","article-title":"Rate-specific synchrony: using noisy oscillations to detect equally active neurons","volume":"105","author":"DA Markowitz","year":"2008","journal-title":"Proc Natl Acad Sci U S A"},{"key":"ref65","doi-asserted-by":"crossref","first-page":"e1003723","DOI":"10.1371\/journal.pcbi.1003723","article-title":"Phase-Coherence Transitions and Communication in the Gamma Range between Delay-Coupled Neuronal Populations","volume":"10","author":"A Barardi","year":"2014","journal-title":"PLoS Comput Biol"},{"key":"ref66","doi-asserted-by":"crossref","first-page":"427","DOI":"10.1007\/s00422-008-0263-8","article-title":"Minimal Hodgkin-Huxley type models for different classes of cortical and thalamic neurons","volume":"99","author":"M Pospischil","year":"2008","journal-title":"Biol Cybern"},{"key":"ref67","doi-asserted-by":"crossref","first-page":"1569","DOI":"10.1109\/TNN.2003.820440","article-title":"Simple model of spiking neurons","volume":"14","author":"EM Izhikevich","year":"2003","journal-title":"IEEE Trans Neural Netw"},{"key":"ref68","doi-asserted-by":"crossref","unstructured":"Hairer E, Wanner G (1991) Solving ordinary differential equations II: Stiff and differential-algebraic problems. Available: <ext-link xmlns:xlink=\"http:\/\/www.w3.org\/1999\/xlink\" ext-link-type=\"uri\" xlink:href=\"http:\/\/www.mendeley.com\/research\/solving-ordinary-differential-equations-ii-stiff-and-differentialalgebraic-problems-1\/\" xlink:type=\"simple\">http:\/\/www.mendeley.com\/research\/solving-ordinary-differential-equations-ii-stiff-and-differentialalgebraic-problems-1\/<\/ext-link>.","DOI":"10.1007\/978-3-662-09947-6_1"},{"key":"ref69","doi-asserted-by":"crossref","first-page":"347","DOI":"10.1016\/j.neuroimage.2005.02.008","article-title":"On the human sensorimotor-cortex beta rhythm: sources and modeling","volume":"26","author":"O Jensen","year":"2005","journal-title":"Neuroimage"},{"key":"ref70","doi-asserted-by":"crossref","first-page":"3963","DOI":"10.1523\/JNEUROSCI.23-10-03963.2003","article-title":"The Underpinnings of the BOLD Functional Magnetic Resonance Imaging Signal","volume":"23","author":"NK Logothetis","year":"2003","journal-title":"J Neurosci"},{"key":"ref71","doi-asserted-by":"crossref","DOI":"10.1109\/ICCV.2001.937655","article-title":"A database of human segmented natural images and its application to evaluating segmentation algorithms and measuring ecological statistics","volume":"2","author":"D Martin","year":"2001","journal-title":"Proc Eighth IEEE Int Conf Comput Vision ICCV 2001"},{"key":"ref72","doi-asserted-by":"crossref","first-page":"1585","DOI":"10.1016\/j.visres.2005.06.038","article-title":"Local luminance and contrast in natural images","volume":"46","author":"RA Frazor","year":"2006","journal-title":"Vis Res"},{"key":"ref73","doi-asserted-by":"crossref","first-page":"32","DOI":"10.3389\/fnins.2011.00032","article-title":"Beyond the cortical column: abundance and physiology of horizontal connections imply a strong role for inputs from the surround","volume":"5","author":"C Boucsein","year":"2011","journal-title":"Front Neurosci"},{"key":"ref74","doi-asserted-by":"crossref","first-page":"141","DOI":"10.1016\/j.jphysparis.2003.09.001","article-title":"Reaching beyond the classical receptive field of V1 neurons: Horizontal or feedback axons?","volume":"97","author":"A Angelucci","year":"2003","journal-title":"J Physiol Paris"},{"key":"ref75","doi-asserted-by":"crossref","first-page":"552","DOI":"10.1109\/78.558469","article-title":"On instantaneous amplitude and phase of signals","volume":"45","author":"B Picinbono","year":"1997","journal-title":"IEEE Trans Signal Process"},{"key":"ref76","doi-asserted-by":"crossref","first-page":"194","DOI":"10.1002\/(SICI)1097-0193(1999)8:4<194::AID-HBM4>3.0.CO;2-C","article-title":"Measuring phase synchrony in brain signals","volume":"8","author":"JP Lachaux","year":"1999","journal-title":"Hum Brain Mapp"},{"key":"ref77","doi-asserted-by":"crossref","first-page":"177","DOI":"10.1016\/j.jneumeth.2007.03.024","article-title":"Nonparametric statistical testing of EEG- and MEG-data","volume":"164","author":"E Maris","year":"2007","journal-title":"J Neurosci Methods"},{"key":"ref78","doi-asserted-by":"crossref","first-page":"57","DOI":"10.1016\/S0303-2647(98)00050-1","article-title":"Rate coding versus temporal order coding: a theoretical approach","volume":"48","author":"J Gautrais","year":"1998","journal-title":"Biosystems"},{"key":"ref79","article-title":"Estimating and interpreting the instantaneous frequency of a signal","volume":"80","author":"B Boashash","year":"1992","journal-title":"I. Fundamentals. Proc IEEE"},{"key":"ref80","doi-asserted-by":"crossref","first-page":"215","DOI":"10.1137\/0119020","article-title":"On the Calculation of Mutual Information","volume":"19","author":"TE Duncan","year":"1970","journal-title":"SIAM J Appl Math"},{"key":"ref81","doi-asserted-by":"crossref","unstructured":"Jaccard J, Becker MA, Wood G (n.d.) Pairwise multiple comparison procedures: A review.","DOI":"10.1037\/0033-2909.96.3.589"},{"key":"ref82","doi-asserted-by":"crossref","first-page":"392","DOI":"10.1523\/JNEUROSCI.16-01-00392.1996","article-title":"Synchronization of fast (30\u201340 Hz) spontaneous cortical rhythms during brain activation","volume":"16","author":"M Steriade","year":"1996","journal-title":"J Neurosci"},{"key":"ref83","doi-asserted-by":"crossref","first-page":"471","DOI":"10.1113\/jphysiol.1996.sp021397","article-title":"Analysis of gamma rhythms in the rat hippocampus in vitro and in vivo","volume":"493","author":"RD Traub","year":"1996","journal-title":"J Physiol"},{"key":"ref84","doi-asserted-by":"crossref","first-page":"6402","DOI":"10.1523\/JNEUROSCI.16-20-06402.1996","article-title":"Gamma Oscillation by Synaptic Inhibition in a Hippocampal Interneuronal Network Model","volume":"16","author":"X-J Wang","year":"1996","journal-title":"J Neurosci"},{"key":"ref85","doi-asserted-by":"crossref","first-page":"415","DOI":"10.1152\/jn.01095.2002","article-title":"What determines the frequency of fast network oscillations with irregular neural discharges? I. Synaptic dynamics and excitation-inhibition balance","volume":"90","author":"N Brunel","year":"2003","journal-title":"J Neurophysiol"},{"key":"ref86","doi-asserted-by":"crossref","first-page":"247","DOI":"10.1007\/s10827-006-6358-0","article-title":"Attentional modulation of firing rate and synchrony in a model cortical network","volume":"20","author":"C Buia","year":"2006","journal-title":"J Comput Neurosci"},{"key":"ref87","doi-asserted-by":"crossref","first-page":"519","DOI":"10.1080\/13506280143000098","article-title":"Flexible cortical gamma-band correlations suggest neural principles of visual processing","volume":"8","author":"R Eckhorn","year":"2001","journal-title":"Vis cogn"},{"key":"ref88","doi-asserted-by":"crossref","first-page":"283","DOI":"10.1109\/72.363423","article-title":"Locally excitatory globally inhibitory oscillator networks","volume":"6","author":"D Wang","year":"1995","journal-title":"IEEE Trans Neural Netw"},{"key":"ref89","doi-asserted-by":"crossref","first-page":"407","DOI":"10.1038\/nrn3241","article-title":"The origin of extracellular fields and currents\u2014EEG, ECoG, LFP and spikes","volume":"13","author":"G Buzs\u00e1ki","year":"2012","journal-title":"Nat Rev Neurosci"},{"key":"ref90","doi-asserted-by":"crossref","first-page":"1077","DOI":"10.1016\/j.neuron.2013.08.019","article-title":"Attentional Modulation of Cell-Class-Specific Gamma-Band Synchronization in Awake Monkey Area V4","volume":"80","author":"M Vinck","year":"2013","journal-title":"Neuron"},{"key":"ref91","doi-asserted-by":"crossref","first-page":"309","DOI":"10.1016\/j.tins.2007.05.005","article-title":"The gamma cycle","volume":"30","author":"P Fries","year":"2007","journal-title":"Trends Neurosci"},{"key":"ref92","doi-asserted-by":"crossref","first-page":"415","DOI":"10.1152\/jn.01095.2002","article-title":"What determines the frequency of fast network oscillations with irregular neural discharges? I. Synaptic dynamics and excitation-inhibition balance","volume":"90","author":"N Brunel","year":"2003","journal-title":"J Neurophysiol"},{"key":"ref93","first-page":"1285","article-title":"Image segmentation using pulse coupled neural networks. Proceedings of 1994 IEEE International Conference on Neural Networks (ICNN\u201994)","volume":"2","author":"HS Ranganath","year":"1994","journal-title":"IEEE"},{"key":"ref94","doi-asserted-by":"crossref","first-page":"805","DOI":"10.1162\/neco.1997.9.4.805","article-title":"Image Segmentation Based on Oscillatory Correlation","volume":"9","author":"D Wang","year":"1997","journal-title":"Neural Comput"},{"key":"ref95","doi-asserted-by":"crossref","first-page":"1002","DOI":"10.1016\/j.neuron.2013.03.007","article-title":"The theta-gamma neural code","volume":"77","author":"JE Lisman","year":"2013","journal-title":"Neuron"},{"key":"ref96","doi-asserted-by":"crossref","first-page":"7002","DOI":"10.1073\/pnas.0502366102","article-title":"Background gamma rhythmicity and attention in cortical local circuits: a computational study","volume":"102","author":"C B\u00f6rgers","year":"2005","journal-title":"Proc Natl Acad Sci U S A"},{"key":"ref97","doi-asserted-by":"crossref","first-page":"13484","DOI":"10.1523\/JNEUROSCI.2207-09.2009","article-title":"Oscillations, phase-of-firing coding, and spike timing-dependent plasticity: an efficient learning scheme","volume":"29","author":"T Masquelier","year":"2009","journal-title":"J Neurosci"},{"key":"ref98","doi-asserted-by":"crossref","first-page":"683","DOI":"10.1016\/j.neuron.2008.09.014","article-title":"Entrainment of neocortical neurons and gamma oscillations by the hippocampal theta rhythm","volume":"60","author":"A Sirota","year":"2008","journal-title":"Neuron"},{"key":"ref99","doi-asserted-by":"crossref","first-page":"17040","DOI":"10.1523\/JNEUROSCI.2209-11.2011","article-title":"Dual Gamma Rhythm Generators Control Interlaminar Synchrony in Auditory Cortex","volume":"31","author":"M Ainsworth","year":"2011","journal-title":"J Neurosci"},{"key":"ref100","doi-asserted-by":"crossref","first-page":"572","DOI":"10.1016\/j.neuron.2012.08.004","article-title":"Rates and rhythms: a synergistic view of frequency and temporal coding in neuronal networks","volume":"75","author":"M Ainsworth","year":"2012","journal-title":"Neuron"},{"key":"ref101","doi-asserted-by":"crossref","first-page":"947","DOI":"10.1038\/14731","article-title":"Information theory and neural coding","volume":"2","author":"A Borst","year":"1999","journal-title":"Nat Neurosci"},{"key":"ref102","article-title":"Spikes: Exploring the Neural Code","author":"F Rieke","year":"1997"},{"key":"ref103","doi-asserted-by":"crossref","first-page":"13484","DOI":"10.1523\/JNEUROSCI.2207-09.2009","article-title":"Oscillations, phase-of-firing coding, and spike timing-dependent plasticity: an efficient learning scheme","volume":"29","author":"T Masquelier","year":"2009","journal-title":"J Neurosci"},{"key":"ref104","doi-asserted-by":"crossref","first-page":"1","DOI":"10.1016\/j.tins.2004.10.010","article-title":"Spike times make sense","volume":"28","author":"R VanRullen","year":"2005","journal-title":"Trends Neurosci"},{"key":"ref105","doi-asserted-by":"crossref","first-page":"130","DOI":"10.1016\/S0166-2236(96)80019-1","article-title":"Integrator or coincidence detector? The role of the cortical neuron revisited","volume":"19","author":"P K\u00f6nig","year":"1996","journal-title":"Trends Neurosci"},{"key":"ref106","doi-asserted-by":"crossref","first-page":"539","DOI":"10.1038\/35086012","article-title":"Correlated neuronal activity and the flow of neural information","volume":"2","author":"E Salinas","year":"2001","journal-title":"Nat Rev Neurosci"},{"key":"ref107","doi-asserted-by":"crossref","first-page":"125","DOI":"10.1038\/35039062","article-title":"Information processing with population codes","volume":"1","author":"a Pouget","year":"2000","journal-title":"Nat Rev Neurosci"},{"key":"ref108","doi-asserted-by":"crossref","first-page":"529","DOI":"10.1038\/990101","article-title":"Stable propagation of synchronous spiking in cortical neural networks","volume":"402","author":"M Diesmann","year":"1999","journal-title":"Nature"},{"key":"ref109","doi-asserted-by":"crossref","first-page":"762","DOI":"10.1016\/j.neuron.2012.12.036","article-title":"\u03b3 and the coordination of spiking activity in early visual cortex","volume":"77","author":"X Jia","year":"2013","journal-title":"Neuron"},{"key":"ref110","doi-asserted-by":"crossref","first-page":"1","DOI":"10.1016\/0168-0102(96)01075-9","article-title":"Population coding by cell assemblies\u2014what it really is in the brain","volume":"26","author":"Y Sakurai","year":"1996","journal-title":"Neurosci Res"},{"key":"ref111","doi-asserted-by":"crossref","first-page":"325","DOI":"10.1016\/S0896-6273(02)00586-X","article-title":"Theta oscillations in the hippocampus","volume":"33","author":"G Buzs\u00e1ki","year":"2002","journal-title":"Neuron"},{"key":"ref112","unstructured":"Marr D (1982) Vision. book. Available: <ext-link xmlns:xlink=\"http:\/\/www.w3.org\/1999\/xlink\" ext-link-type=\"uri\" xlink:href=\"http:\/\/books.google.com\/books?id=EehUQwAACAAJ&printsec=frontcover\\npapers2:\/\/publication\/uuid\/FBD15E5F-E503-450B-B059-4C15D54099CE\" xlink:type=\"simple\">http:\/\/books.google.com\/books?id=EehUQwAACAAJ&amp;printsec=frontcover\\npapers2:\/\/publication\/uuid\/FBD15E5F-E503-450B-B059-4C15D54099CE<\/ext-link>."},{"key":"ref113","doi-asserted-by":"crossref","first-page":"591","DOI":"10.1109\/72.761716","article-title":"Perfect image segmentation using pulse coupled neural networks","volume":"10","author":"G Kuntimad","year":"1999","journal-title":"IEEE Trans Neural Netw"},{"key":"ref114","doi-asserted-by":"crossref","first-page":"43","DOI":"10.1016\/j.biosystems.2004.05.005","article-title":"Oscillatory network with self-organized dynamical connections for synchronization-based image segmentation","volume":"76","author":"M Kuzmina","year":"2004","journal-title":"Biosystems"},{"key":"ref115","doi-asserted-by":"crossref","first-page":"423","DOI":"10.1016\/S0893-6080(02)00028-X","article-title":"A dynamically coupled neural oscillator network for image segmentation","volume":"15","author":"K Chen","year":"2002","journal-title":"Neural Networks"},{"key":"ref116","doi-asserted-by":"crossref","first-page":"155","DOI":"10.1162\/neco.1991.3.2.155","article-title":"Stimulus-Dependent Assembly Formation of Oscillatory Responses: I. Synchronization","volume":"3","author":"P K\u00f6nig","year":"1991","journal-title":"Neural Comput"},{"key":"ref117","doi-asserted-by":"crossref","first-page":"487","DOI":"10.1016\/j.jphysparis.2005.09.004","article-title":"Temporal codes and sparse representations: a key to understanding rapid processing in the visual system","volume":"98","author":"R Guyonneau","year":"2004","journal-title":"J Physiol Paris"},{"key":"ref118","doi-asserted-by":"crossref","first-page":"663","DOI":"10.1038\/nature08002","article-title":"Driving fast-spiking cells induces gamma rhythm and controls sensory responses","volume":"459","author":"JA Cardin","year":"2009","journal-title":"Nature"}],"container-title":["PLOS Computational Biology"],"original-title":[],"language":"en","link":[{"URL":"http:\/\/dx.plos.org\/10.1371\/journal.pcbi.1004072","content-type":"unspecified","content-version":"vor","intended-application":"similarity-checking"}],"deposited":{"date-parts":[[2024,6,7]],"date-time":"2024-06-07T06:08:21Z","timestamp":1717740501000},"score":1,"resource":{"primary":{"URL":"https:\/\/dx.plos.org\/10.1371\/journal.pcbi.1004072"}},"subtitle":[],"editor":[{"given":"Bard","family":"Ermentrout","sequence":"first","affiliation":[],"role":[{"role":"editor","vocabulary":"crossref"}]}],"short-title":[],"issued":{"date-parts":[[2015,2,13]]},"references-count":118,"journal-issue":{"issue":"2","published-online":{"date-parts":[[2015,2,13]]}},"URL":"https:\/\/doi.org\/10.1371\/journal.pcbi.1004072","relation":{},"ISSN":["1553-7358"],"issn-type":[{"value":"1553-7358","type":"electronic"}],"subject":[],"published":{"date-parts":[[2015,2,13]]}}}