{"status":"ok","message-type":"work","message-version":"1.0.0","message":{"indexed":{"date-parts":[[2026,5,13]],"date-time":"2026-05-13T02:06:16Z","timestamp":1778637976730,"version":"3.51.4"},"reference-count":65,"publisher":"Public Library of Science (PLoS)","issue":"3","license":[{"start":{"date-parts":[[2015,3,20]],"date-time":"2015-03-20T00:00:00Z","timestamp":1426809600000},"content-version":"vor","delay-in-days":0,"URL":"http:\/\/creativecommons.org\/licenses\/by\/4.0\/"}],"content-domain":{"domain":["www.ploscompbiol.org"],"crossmark-restriction":false},"short-container-title":["PLoS Comput Biol"],"DOI":"10.1371\/journal.pcbi.1004134","type":"journal-article","created":{"date-parts":[[2015,3,20]],"date-time":"2015-03-20T18:58:49Z","timestamp":1426877929000},"page":"e1004134","update-policy":"https:\/\/doi.org\/10.1371\/journal.pcbi.corrections_policy","source":"Crossref","is-referenced-by-count":22,"title":["When Can Species Abundance Data Reveal Non-neutrality?"],"prefix":"10.1371","volume":"11","author":[{"given":"Omar","family":"Al Hammal","sequence":"first","affiliation":[],"role":[{"role":"author","vocabulary":"crossref"}]},{"given":"David","family":"Alonso","sequence":"additional","affiliation":[],"role":[{"role":"author","vocabulary":"crossref"}]},{"given":"Rampal S.","family":"Etienne","sequence":"additional","affiliation":[],"role":[{"role":"author","vocabulary":"crossref"}]},{"given":"Stephen J.","family":"Cornell","sequence":"additional","affiliation":[],"role":[{"role":"author","vocabulary":"crossref"}]}],"member":"340","published-online":{"date-parts":[[2015,3,20]]},"reference":[{"key":"ref1","doi-asserted-by":"crossref","first-page":"1145","DOI":"10.1126\/science.243.4895.1145","article-title":"Macroecology: The division of food and space among species on continents","volume":"243","author":"JH Brown","year":"1989","journal-title":"Science"},{"key":"ref2","article-title":"Macroecology","author":"JH Brown","year":"1995"},{"key":"ref3","doi-asserted-by":"crossref","first-page":"1","DOI":"10.1086\/324112","article-title":"Comparing classical community models: Theoretical consequences for patterns of diversity","volume":"159","author":"J Chave","year":"2002","journal-title":"Am Nat"},{"key":"ref4","first-page":"107","article-title":"Biotic Interactions in the Tropics: their role in the maintenance of species diversity","author":"D Purves","year":"2005"},{"key":"ref5","first-page":"379","article-title":"A statistical treatment of associations","volume":"44","author":"I Motomura","year":"1932","journal-title":"Jpn J Zool"},{"key":"ref6","doi-asserted-by":"crossref","first-page":"293","DOI":"10.1073\/pnas.43.3.293","article-title":"On the relative abundance of bird species","volume":"43","author":"R Mac Arthur","year":"1957","journal-title":"Proc Natl Acad Sci USA"},{"key":"ref7","doi-asserted-by":"crossref","first-page":"177","DOI":"10.1111\/j.0269-8463.2005.00930.x","article-title":"Species abundance distributions: pattern or process?","volume":"19","author":"A Magurran","year":"2005","journal-title":"Funct Ecol"},{"key":"ref8","doi-asserted-by":"crossref","first-page":"95","DOI":"10.1111\/j.1461-0248.2006.00996.x","article-title":"A niche for neutrality","volume":"2","author":"PB Adler","year":"2007","journal-title":"Ecol Lett"},{"key":"ref9","doi-asserted-by":"crossref","first-page":"318","DOI":"10.1016\/j.tpb.2006.12.008","article-title":"When and why do non-neutral metacommunities appear neutral?","volume":"71","author":"SC Walker","year":"2007","journal-title":"Theor Popul Biol"},{"key":"ref10","doi-asserted-by":"crossref","first-page":"127","DOI":"10.1016\/j.jtbi.2011.03.003","article-title":"Negative density dependence can offset the effect of species competitive asymmetry: A niche-based mechanism for neutral-like patterns","volume":"278","author":"X Du","year":"2011","journal-title":"J Theor Biol"},{"key":"ref11","doi-asserted-by":"crossref","first-page":"995","DOI":"10.1111\/j.1461-0248.2007.01094.x","article-title":"Species abundance distributions: moving beyond single prediction theories to integration within an ecological framework","volume":"10","author":"BJ McGill","year":"2007","journal-title":"Ecol Lett"},{"key":"ref12","doi-asserted-by":"crossref","first-page":"327","DOI":"10.2307\/1942257","article-title":"Community structure: a neutral model analysis","volume":"46","author":"H Caswell","year":"1976","journal-title":"Ecol Monogr"},{"key":"ref13","doi-asserted-by":"crossref","first-page":"1035","DOI":"10.1038\/nature01883","article-title":"Neutral theory and relative species abundance in ecology","volume":"424","author":"I Volkov","year":"2003","journal-title":"Nature"},{"key":"ref14","doi-asserted-by":"crossref","first-page":"901","DOI":"10.1111\/j.1461-0248.2004.00640.x","article-title":"Sampling Hubbell\u2019s neutral theory of biodiversity","volume":"7","author":"D Alonso","year":"2004","journal-title":"Ecol Lett"},{"key":"ref15","doi-asserted-by":"crossref","first-page":"241","DOI":"10.1111\/j.1461-0248.2003.00566.x","article-title":"Neutral theory and community ecology","volume":"7","author":"J Chave","year":"2004","journal-title":"Ecol Lett"},{"issue":"6","key":"ref16","doi-asserted-by":"crossref","first-page":"1411","DOI":"10.1890\/0012-9658(2006)87[1411:EEONT]2.0.CO;2","article-title":"Empirical evaluation of neutral theory","volume":"87","author":"BJ McGill","year":"2006","journal-title":"Ecology"},{"key":"ref17","doi-asserted-by":"crossref","first-page":"485","DOI":"10.1007\/s10955-006-9163-2","article-title":"Neutral community theory: how stochasticity and dispersal-limitation can explain species coexistence","volume":"128","author":"RS Etienne","year":"2006","journal-title":"J Stat Phys"},{"key":"ref18","doi-asserted-by":"crossref","first-page":"1","DOI":"10.1016\/j.jtbi.2010.12.021","article-title":"A sampling theory for asymmetric communities","volume":"273","author":"AE Noble","year":"2011","journal-title":"J Theor Biol"},{"key":"ref19","doi-asserted-by":"crossref","first-page":"961","DOI":"10.1111\/j.1600-0706.2011.19697.x","article-title":"Independent species in independent niches behave neutrally","volume":"120","author":"B Haegeman","year":"2011","journal-title":"Oikos"},{"key":"ref20","doi-asserted-by":"crossref","first-page":"198","DOI":"10.1016\/j.tree.2012.02.001","article-title":"The coherence problem with the unified neutral theory of biodiversity","volume":"27","author":"JS Clark","year":"2012","journal-title":"Trends Ecol Evol"},{"key":"ref21","doi-asserted-by":"crossref","first-page":"1215","DOI":"10.1111\/j.1365-2656.2010.01738.x","article-title":"Different but equal: the implausible assumption at the heart of neutral theory: Implausibility of neutrality","volume":"79","author":"DW Purves","year":"2010","journal-title":"J Anim Ecol"},{"key":"ref22","doi-asserted-by":"crossref","first-page":"451","DOI":"10.1016\/j.tree.2006.03.019","article-title":"The merits of neutral theory","volume":"21","author":"D Alonso","year":"2006","journal-title":"Trends Ecol Evol"},{"key":"ref23","doi-asserted-by":"crossref","first-page":"203","DOI":"10.1016\/j.tree.2012.01.004","article-title":"The case for ecological neutral theory","volume":"27","author":"J Rosindell","year":"2012","journal-title":"Trends Ecol Evol"},{"key":"ref24","doi-asserted-by":"crossref","first-page":"257","DOI":"10.1007\/s10441-012-9144-6","article-title":"The neutral-niche debate: A philosophical perspective","volume":"60","author":"PL Wennekes","year":"2012","journal-title":"Acta Biotheor"},{"key":"ref25","doi-asserted-by":"crossref","first-page":"1","DOI":"10.1111\/j.0269-8463.2005.00948.x","article-title":"Neutrality and the niche","volume":"19","author":"K Gaston","year":"2005","journal-title":"Funct Ecol"},{"key":"ref26","doi-asserted-by":"crossref","first-page":"253","DOI":"10.1111\/j.1461-0248.2004.00717.x","article-title":"A new sampling formula for neutral biodiversity","volume":"8","author":"RS Etienne","year":"2005","journal-title":"Ecol Lett"},{"key":"ref27","doi-asserted-by":"crossref","first-page":"608","DOI":"10.1111\/j.1461-0248.2007.01052.x","article-title":"A neutral sampling formula for multiple samples and an \u201cexact\u201d test of neutrality","volume":"10","author":"RS Etienne","year":"2007","journal-title":"Ecol Lett"},{"key":"ref28","doi-asserted-by":"crossref","first-page":"847","DOI":"10.1890\/08-0750.1","article-title":"Improved estimation of neutral model parameters for multiple samples with different degrees of dispersal limitation","volume":"90","author":"RS Etienne","year":"2009","journal-title":"Ecology"},{"key":"ref29","doi-asserted-by":"crossref","first-page":"510","DOI":"10.1016\/j.jtbi.2008.12.016","article-title":"Maximum likelihood estimation of neutral model parameters for multiple samples with different degrees of dispersal limitation","volume":"257","author":"RS Etienne","year":"2009","journal-title":"J Theor Biol"},{"key":"ref30","doi-asserted-by":"crossref","first-page":"493","DOI":"10.1111\/j.1461-0248.2005.00745.x","article-title":"Confronting different models of community structure to species-abundance data: a bayesian model comparison","volume":"8","author":"RS Etienne","year":"2005","journal-title":"Ecol Lett"},{"key":"ref31","article-title":"The Unified Neutral Theory of Biodiversity and Biogeography","author":"SP Hubbell","year":"2001"},{"key":"ref32","doi-asserted-by":"crossref","first-page":"399","DOI":"10.1111\/j.1461-0248.2006.00884.x","article-title":"Reconciling niche and neutrality: the continuum hypothesis","volume":"9","author":"D Gravel","year":"2006","journal-title":"Ecol Lett"},{"key":"ref33","doi-asserted-by":"crossref","first-page":"150","DOI":"10.1016\/j.jtbi.2010.10.006","article-title":"A mathematical synthesis of niche and neutral theories in community ecology","volume":"169","author":"B Haegeman","year":"2011","journal-title":"J Theor Biol"},{"key":"ref34","doi-asserted-by":"crossref","first-page":"93","DOI":"10.1111\/j.1461-0248.2007.01127.x","article-title":"The implicit assumption of symmetry and the species abundance distribution","volume":"11","author":"D Alonso","year":"2008","journal-title":"Ecol Lett"},{"key":"ref35","doi-asserted-by":"crossref","first-page":"6691","DOI":"10.1073\/pnas.0810096106","article-title":"Species abundance distribution results from a spatial analogy of central limit theorem","volume":"106","author":"AL Sizling","year":"2009","journal-title":"Proc Natl Acad Sci USA"},{"issue":"36","key":"ref36","doi-asserted-by":"crossref","first-page":"15821","DOI":"10.1073\/pnas.1009387107","article-title":"Niche and neutral models predict asymptotically equivalent species abundance distributions in high-diversity ecological communities","volume":"107","author":"R Chisholm","year":"2010","journal-title":"Proc Natl Acad Sci USA"},{"key":"ref37","unstructured":"Hubbell SP, Condit R, Foster RB (2005). Barro colorado forest census plot data. url <ext-link xmlns:xlink=\"http:\/\/www.w3.org\/1999\/xlink\" ext-link-type=\"uri\" xlink:href=\"https:\/\/ctfs.arnarb.harvard.edu\/webatlas\/datasets\/bci\" xlink:type=\"simple\">https:\/\/ctfs.arnarb.harvard.edu\/webatlas\/datasets\/bci<\/ext-link>."},{"key":"ref38","doi-asserted-by":"crossref","DOI":"10.1007\/978-3-662-03664-8","article-title":"Tropical Forest Census Plots","author":"R Condit","year":"1998"},{"key":"ref39","doi-asserted-by":"crossref","first-page":"554","DOI":"10.1126\/science.283.5401.554","article-title":"Light gap disturbances, recruitment limitation, and tree diversity in a neotropical forest","volume":"283","author":"SP Hubbell","year":"1999","journal-title":"Science"},{"key":"ref40","doi-asserted-by":"crossref","first-page":"211","DOI":"10.1006\/tpbi.2000.1486","article-title":"General theory of competitive coexistence in spatially-varying environments","volume":"58","author":"P Chesson","year":"2000","journal-title":"Theor Popul Biol"},{"key":"ref41","doi-asserted-by":"crossref","first-page":"343","DOI":"10.1146\/annurev.ecolsys.31.1.343","article-title":"Mechanisms of maintenance of species diversity","volume":"31","author":"P Chesson","year":"2000","journal-title":"Annu Rev Ecol Syst"},{"key":"ref42","doi-asserted-by":"crossref","first-page":"1","DOI":"10.1016\/j.jtbi.2013.08.024","article-title":"Species abundances and lifetimes: From neutral to niche-stabilized communities","volume":"338","author":"S Pigolotti","year":"2013","journal-title":"J Theor Biol"},{"key":"ref43","doi-asserted-by":"crossref","first-page":"1017","DOI":"10.1111\/j.1461-0248.2007.01096.x","article-title":"The maximum entropy formalism and the idiosyncratic theory of biodiversity","volume":"10","author":"S Pueyo","year":"2007","journal-title":"Ecol Lett"},{"key":"ref44","doi-asserted-by":"crossref","first-page":"248","DOI":"10.1890\/06-1817.1","article-title":"A nearly neutral model of biodiversity","volume":"89","author":"SR Zhou","year":"2008","journal-title":"Ecology"},{"key":"ref45","doi-asserted-by":"crossref","first-page":"143","DOI":"10.1111\/j.2006.0030-1299.15300.x","article-title":"Testing the standard neutral model of biodiversity in lake communities","volume":"116","author":"SC Walker","year":"2007","journal-title":"Oikos"},{"key":"ref46","doi-asserted-by":"crossref","first-page":"45","DOI":"10.1038\/nature06197","article-title":"Patterns of relative species abundance in rainforests and coral reefs","volume":"450","author":"I Volkov","year":"2007","journal-title":"Nature"},{"key":"ref47","doi-asserted-by":"crossref","first-page":"13854","DOI":"10.1073\/pnas.0903244106","article-title":"Inferring species interactions in tropical forests","volume":"106","author":"I Volkov","year":"2009","journal-title":"Proc Natl Acad Sci USA"},{"key":"ref48","doi-asserted-by":"crossref","first-page":"1147","DOI":"10.1111\/j.1461-0248.2005.00817.x","article-title":"A dispersal-limited sampling theory for species and alleles","volume":"8","author":"RS Etienne","year":"2005","journal-title":"Ecol Lett"},{"key":"ref49","doi-asserted-by":"crossref","first-page":"1008","DOI":"10.1111\/j.1461-0248.2008.01208.x","article-title":"Multiple modes in a coral species abundance distribution","volume":"11","author":"M Dornelas","year":"2008","journal-title":"Ecol Lett"},{"key":"ref50","doi-asserted-by":"crossref","unstructured":"Rosindell J, Wong Y, Etienne RS (2008) A coalescence approach to spatial neutral ecology. Ecol Inform: 259\u2013271.","DOI":"10.1016\/j.ecoinf.2008.05.001"},{"key":"ref51","doi-asserted-by":"crossref","first-page":"E1","DOI":"10.1038\/nature04826","article-title":"Theoretical biology\u2014comparing models of species abundance","volume":"441","author":"J Chave","year":"2006","journal-title":"Nature"},{"key":"ref52","doi-asserted-by":"crossref","first-page":"909","DOI":"10.1111\/j.1461-0248.2009.01346.x","article-title":"When can we distinguish between neutral and non-neutral processes in community dynamics under ecological drift?","volume":"12","author":"L Ruokolainen","year":"2009","journal-title":"Ecol Lett"},{"key":"ref53","doi-asserted-by":"crossref","first-page":"E1","DOI":"10.1038\/nature04826","article-title":"Comparing models of species abundance","volume":"441","author":"J Chave","year":"2006","journal-title":"Nature"},{"key":"ref54","doi-asserted-by":"crossref","first-page":"926","DOI":"10.1038\/nature05320","article-title":"Dynamical evolution of ecological systems","volume":"444","author":"S Azaele","year":"2006","journal-title":"Nature"},{"key":"ref55","doi-asserted-by":"crossref","first-page":"71","DOI":"10.1016\/j.tpb.2011.06.002","article-title":"Time-dependent solutions of the spatially implicit neutral model of biodiversity","volume":"80","author":"R Chisholm","year":"2011","journal-title":"J Theor Biol"},{"key":"ref56","doi-asserted-by":"crossref","first-page":"239","DOI":"10.1111\/j.1461-0248.2008.01280.x","article-title":"Inferring the parameters of the neutral theory of biodiversity using phylogrenetic information and implications for tropical forests","volume":"12","author":"F Jabot","year":"2009","journal-title":"Ecol Lett"},{"key":"ref57","doi-asserted-by":"crossref","first-page":"1841","DOI":"10.1111\/j.1558-5646.2011.01265.x","article-title":"Neutral biodiversity can explain the imbalance of phylogenetic trees but not the tempo of their diversification","volume":"65","author":"TJ Davies","year":"2011","journal-title":"Evolution"},{"key":"ref58","doi-asserted-by":"crossref","first-page":"923","DOI":"10.1086\/283778","article-title":"Environmental variability promotes coexistence in lottery competitive systems","volume":"117","author":"PL Chesson","year":"1981","journal-title":"Am Nat"},{"key":"ref59","doi-asserted-by":"crossref","first-page":"392","DOI":"10.1111\/j.0030-1299.2006.14188.x","article-title":"Diversity: between neutrality and structure","volume":"112","author":"S Pueyo","year":"2006","journal-title":"Oikos"},{"key":"ref60","doi-asserted-by":"crossref","first-page":"87","DOI":"10.1016\/0040-5809(72)90035-4","article-title":"The sampling theory of selectively neutral alleles","volume":"3","author":"W Ewens","year":"1972","journal-title":"Theor Popul Biol"},{"key":"ref61","doi-asserted-by":"crossref","first-page":"8466","DOI":"10.1103\/PhysRevE.62.8466","article-title":"A mean field stochastic theory for species rich assembled communities","volume":"62","author":"A McKane","year":"2000","journal-title":"Phys Rev E"},{"key":"ref62","doi-asserted-by":"crossref","first-page":"606","DOI":"10.1086\/303345","article-title":"The distribution of abundances in neutral communities","volume":"155","author":"G Bell","year":"2000","journal-title":"Am Nat"},{"issue":"3","key":"ref63","doi-asserted-by":"crossref","first-page":"e14717","DOI":"10.1371\/journal.pone.0014717","article-title":"The spatial limitations of current neutral models of biodiversity","volume":"6","author":"J Rosindell","year":"2011","journal-title":"PLoS One"},{"key":"ref64","doi-asserted-by":"crossref","first-page":"1308","DOI":"10.1111\/j.0030-1299.2008.16724.x","article-title":"Reconciling neutral community models and environmental filtering: theory and an empirical test","volume":"117","author":"F Jabot","year":"2008","journal-title":"Oikos"},{"key":"ref65","doi-asserted-by":"crossref","first-page":"101","DOI":"10.1214\/ss\/1009213286","article-title":"Interval estimation for a binomial proportion","volume":"16","author":"L Brown","year":"2001","journal-title":"Stat Sci"}],"container-title":["PLOS Computational Biology"],"original-title":[],"language":"en","link":[{"URL":"http:\/\/dx.plos.org\/10.1371\/journal.pcbi.1004134","content-type":"unspecified","content-version":"vor","intended-application":"similarity-checking"}],"deposited":{"date-parts":[[2024,6,7]],"date-time":"2024-06-07T23:22:35Z","timestamp":1717802555000},"score":1,"resource":{"primary":{"URL":"https:\/\/dx.plos.org\/10.1371\/journal.pcbi.1004134"}},"subtitle":[],"editor":[{"given":"Stefano","family":"Allesina","sequence":"first","affiliation":[],"role":[{"role":"editor","vocabulary":"crossref"}]}],"short-title":[],"issued":{"date-parts":[[2015,3,20]]},"references-count":65,"journal-issue":{"issue":"3","published-online":{"date-parts":[[2015,3,20]]}},"URL":"https:\/\/doi.org\/10.1371\/journal.pcbi.1004134","relation":{},"ISSN":["1553-7358"],"issn-type":[{"value":"1553-7358","type":"electronic"}],"subject":[],"published":{"date-parts":[[2015,3,20]]}}}