{"status":"ok","message-type":"work","message-version":"1.0.0","message":{"indexed":{"date-parts":[[2026,5,13]],"date-time":"2026-05-13T03:17:15Z","timestamp":1778642235894,"version":"3.51.4"},"update-to":[{"DOI":"10.1371\/journal.pcbi.1005343","type":"new_version","label":"New version","source":"publisher","updated":{"date-parts":[[2017,2,16]],"date-time":"2017-02-16T00:00:00Z","timestamp":1487203200000}}],"reference-count":64,"publisher":"Public Library of Science (PLoS)","issue":"2","license":[{"start":{"date-parts":[[2017,2,2]],"date-time":"2017-02-02T00:00:00Z","timestamp":1485993600000},"content-version":"vor","delay-in-days":0,"URL":"http:\/\/creativecommons.org\/licenses\/by\/4.0\/"}],"content-domain":{"domain":["www.ploscompbiol.org"],"crossmark-restriction":false},"short-container-title":["PLoS Comput Biol"],"DOI":"10.1371\/journal.pcbi.1005343","type":"journal-article","created":{"date-parts":[[2017,2,2]],"date-time":"2017-02-02T13:40:40Z","timestamp":1486042840000},"page":"e1005343","update-policy":"https:\/\/doi.org\/10.1371\/journal.pcbi.corrections_policy","source":"Crossref","is-referenced-by-count":10,"title":["Correlation-based model of artificially induced plasticity in motor cortex by a bidirectional brain-computer interface"],"prefix":"10.1371","volume":"13","author":[{"given":"Guillaume","family":"Lajoie","sequence":"first","affiliation":[],"role":[{"role":"author","vocabulary":"crossref"}]},{"ORCID":"https:\/\/orcid.org\/0000-0002-7800-0911","authenticated-orcid":true,"given":"Nedialko I.","family":"Krouchev","sequence":"additional","affiliation":[],"role":[{"role":"author","vocabulary":"crossref"}]},{"given":"John F.","family":"Kalaska","sequence":"additional","affiliation":[],"role":[{"role":"author","vocabulary":"crossref"}]},{"given":"Adrienne L.","family":"Fairhall","sequence":"additional","affiliation":[],"role":[{"role":"author","vocabulary":"crossref"}]},{"given":"Eberhard E.","family":"Fetz","sequence":"additional","affiliation":[],"role":[{"role":"author","vocabulary":"crossref"}]}],"member":"340","published-online":{"date-parts":[[2017,2,2]]},"reference":[{"issue":"4","key":"ref1","doi-asserted-by":"crossref","first-page":"751","DOI":"10.1152\/jn.1980.44.4.751","article-title":"Postspike facilitation of forelimb muscle activity by primate corticomotoneuronal cells","volume":"44","author":"PD Cheney","year":"1980","journal-title":"Journal of Neurophysiology"},{"key":"ref2","first-page":"453","article-title":"Dynamic Aspects of Neocortical Function","author":"EE Fetz","year":"1984"},{"issue":"5","key":"ref3","doi-asserted-by":"crossref","first-page":"2125","DOI":"10.1152\/jn.2001.86.5.2125","article-title":"Constraints on Somatotopic Organization in the Primary Motor Cortex","volume":"86","author":"MH Schieber","year":"2001","journal-title":"Journal of Neurophysiology"},{"key":"ref4","article-title":"Neural mechanisms subserving cutaneous sensibility, with special reference to the role of afferent inhibition in sensory perception and discrimination","volume":"vol. 32","author":"VB Mountcastle","year":"1959","journal-title":"Bulletin of the Johns Hopkins Hospital"},{"issue":"1","key":"ref5","doi-asserted-by":"crossref","first-page":"115","DOI":"10.1016\/S0896-6273(03)00162-4","article-title":"Synchrony between neurons with similar muscle fields in monkey motor cortex","volume":"38","author":"A Jackson","year":"2003","journal-title":"Neuron"},{"issue":"5","key":"ref6","doi-asserted-by":"crossref","first-page":"471","DOI":"10.1177\/1073858405278015","article-title":"In search of the motor engram: motor map plasticity as a mechanism for encoding motor experience","volume":"11","author":"MH Monfils","year":"2005","journal-title":"The Neuroscientist: a review journal bringing neurobiology, neurology and psychiatry"},{"issue":"1","key":"ref7","doi-asserted-by":"crossref","first-page":"393","DOI":"10.1146\/annurev.neuro.23.1.393","article-title":"Plasticity and primary motor cortex","volume":"23","author":"JN Sanes","year":"2000","journal-title":"Annual review of neuroscience"},{"issue":"1","key":"ref8","doi-asserted-by":"crossref","first-page":"149","DOI":"10.1146\/annurev.neuro.21.1.149","article-title":"Cortical plasticity: from synapses to maps","volume":"21","author":"DV Buonomano","year":"1998","journal-title":"Annual review of neuroscience"},{"issue":"2","key":"ref9","doi-asserted-by":"crossref","first-page":"456","DOI":"10.1016\/j.cell.2015.09.029","article-title":"Reconstruction and Simulation of Neocortical Microcircuitry","volume":"163","author":"H Markram","year":"2015","journal-title":"Cell"},{"issue":"7115","key":"ref10","doi-asserted-by":"crossref","first-page":"56","DOI":"10.1038\/nature05226","article-title":"Long-term motor cortex plasticity induced by an electronic neural implant","volume":"444","author":"A Jackson","year":"2006","journal-title":"Nature"},{"key":"ref11","doi-asserted-by":"crossref","DOI":"10.3389\/fnsys.2010.00039","article-title":"Rewiring Neural Interactions by Micro-Stimulation","volume":"4","author":"JM Rebesco","year":"2010","journal-title":"Frontiers in Systems Neuroscience"},{"issue":"1","key":"ref12","doi-asserted-by":"crossref","first-page":"139","DOI":"10.1146\/annurev.neuro.24.1.139","article-title":"Synaptic modification by correlated activity: Hebb\u2019s postulate revisited","volume":"24","author":"G Bi","year":"2001","journal-title":"Annual review of neuroscience"},{"issue":"1","key":"ref13","doi-asserted-by":"crossref","first-page":"25","DOI":"10.1146\/annurev.neuro.31.060407.125639","article-title":"Spike timing-dependent plasticity: a Hebbian learning rule","volume":"31","author":"N Caporale","year":"2008","journal-title":"Annual Review of Neuroscience"},{"issue":"2","key":"ref14","doi-asserted-by":"crossref","first-page":"409","DOI":"10.1113\/jphysiol.1997.sp022031","article-title":"Physiology and anatomy of synaptic connections between thick tufted pyramidal neurones in the developing rat neocortex","volume":"500","author":"H Markram","year":"1997","journal-title":"The Journal of Physiology"},{"issue":"5","key":"ref15","doi-asserted-by":"crossref","first-page":"1301","DOI":"10.1016\/j.neuron.2013.08.028","article-title":"Spike-Timing-Dependent Plasticity in Primate Corticospinal Connections Induced during Free Behavior","volume":"80","author":"Y Nishimura","year":"2013","journal-title":"Neuron"},{"issue":"4","key":"ref16","doi-asserted-by":"crossref","first-page":"4498","DOI":"10.1103\/PhysRevE.59.4498","article-title":"Hebbian learning and spiking neurons","volume":"59","author":"R Kempter","year":"1999","journal-title":"Physical Review E"},{"issue":"5","key":"ref17","doi-asserted-by":"crossref","first-page":"533","DOI":"10.1007\/s00422-007-0148-2","article-title":"Spike-timing-dependent plasticity for neurons with recurrent connections","volume":"96","author":"AN Burkitt","year":"2007","journal-title":"Biological Cybernetics"},{"issue":"5","key":"ref18","doi-asserted-by":"crossref","first-page":"365","DOI":"10.1007\/s00422-010-0405-7","article-title":"Emergence of network structure due to spike-timing-dependent plasticity in recurrent neuronal networks V: self-organization schemes and weight dependence","volume":"103","author":"M Gilson","year":"2010","journal-title":"Biological Cybernetics"},{"issue":"5\u20136","key":"ref19","doi-asserted-by":"crossref","first-page":"427","DOI":"10.1007\/s00422-009-0346-1","article-title":"Emergence of network structure due to spike-timing-dependent plasticity in recurrent neuronal networks IV","volume":"101","author":"M Gilson","year":"2009","journal-title":"Biological Cybernetics"},{"issue":"5\u20136","key":"ref20","doi-asserted-by":"crossref","first-page":"411","DOI":"10.1007\/s00422-009-0343-4","article-title":"Emergence of network structure due to spike-timing-dependent plasticity in recurrent neuronal networks III: Partially connected neurons driven by spontaneous activity","volume":"101","author":"M Gilson","year":"2009","journal-title":"Biological Cybernetics"},{"issue":"2","key":"ref21","doi-asserted-by":"crossref","first-page":"103","DOI":"10.1007\/s00422-009-0320-y","article-title":"Emergence of network structure due to spike-timing-dependent plasticity in recurrent neuronal networks. II. Input selectivity\u2014symmetry breaking","volume":"101","author":"M Gilson","year":"2009","journal-title":"Biological Cybernetics"},{"issue":"2","key":"ref22","doi-asserted-by":"crossref","first-page":"81","DOI":"10.1007\/s00422-009-0319-4","article-title":"Emergence of network structure due to spike-timing-dependent plasticity in recurrent neuronal networks. I. Input selectivity\u2013strengthening correlated input pathways","volume":"101","author":"M Gilson","year":"2009","journal-title":"Biological Cybernetics"},{"issue":"6","key":"ref23","doi-asserted-by":"crossref","first-page":"1437","DOI":"10.1162\/neco.2007.19.6.1437","article-title":"Spike-timing-dependent plasticity in balanced random networks","volume":"19","author":"A Morrison","year":"2007","journal-title":"Neural Computation"},{"issue":"9","key":"ref24","doi-asserted-by":"crossref","first-page":"3697","DOI":"10.1523\/JNEUROSCI.23-09-03697.2003","article-title":"Learning input correlations through nonlinear temporally asymmetric Hebbian plasticity","volume":"23","author":"R G\u00fctig","year":"2003","journal-title":"The Journal of neuroscience: the official journal of the Society for Neuroscience"},{"issue":"4","key":"ref25","doi-asserted-by":"crossref","first-page":"041911","DOI":"10.1103\/PhysRevE.73.041911","article-title":"Learning the structure of correlated synaptic subgroups using stable and competitive spike-timing-dependent plasticity","volume":"73","author":"H Meffin","year":"2006","journal-title":"Physical Review E"},{"key":"ref26","doi-asserted-by":"crossref","first-page":"919","DOI":"10.1038\/78829","article-title":"Competitive Hebbian learning through spike-timing-dependent synaptic plasticity","volume":"3","author":"S Song","year":"2000","journal-title":"Nature Neuroscience"},{"issue":"2","key":"ref27","doi-asserted-by":"crossref","first-page":"339","DOI":"10.1016\/S0896-6273(01)00451-2","article-title":"Cortical development and remapping through spike timing-dependent plasticity","volume":"32","author":"S Song","year":"2001","journal-title":"Neuron"},{"key":"ref28","doi-asserted-by":"crossref","first-page":"8812","DOI":"10.1523\/JNEUROSCI.20-23-08812.2000","article-title":"Stable Hebbian learning from spike timing-dependent plasticity","volume":"20","author":"M Van Rossum","year":"2000","journal-title":"The Journal of Neuroscience"},{"issue":"5","key":"ref29","doi-asserted-by":"crossref","first-page":"2600","DOI":"10.1152\/jn.00803.2004","article-title":"Calcium Time Course as a Signal for Spike-Timing\u2013Dependent Plasticity","volume":"93","author":"JE Rubin","year":"2005","journal-title":"Journal of Neurophysiology"},{"issue":"10","key":"ref30","doi-asserted-by":"crossref","first-page":"e25339","DOI":"10.1371\/journal.pone.0025339","article-title":"Stability versus neuronal specialization for STDP: long-tail weight distributions solve the dilemma","volume":"6","author":"M Gilson","year":"2011","journal-title":"PLoS ONE"},{"issue":"6","key":"ref31","doi-asserted-by":"crossref","first-page":"2775","DOI":"10.1152\/jn.91007.2008","article-title":"Memory Retention and Spike-Timing-Dependent Plasticity","volume":"101","author":"G Billings","year":"2009","journal-title":"Journal of Neurophysiology"},{"issue":"5965","key":"ref32","doi-asserted-by":"crossref","first-page":"587","DOI":"10.1126\/science.1179850","article-title":"The Asynchronous State in Cortical Circuits","volume":"327","author":"A Renart","year":"2010","journal-title":"Science (New York, NY)"},{"key":"ref33","doi-asserted-by":"crossref","first-page":"3870","DOI":"10.1523\/JNEUROSCI.18-10-03870.1998","article-title":"The variable discharge of cortical neurons: implications for connectivity, computation, and information coding","volume":"18","author":"MN Shadlen","year":"1998","journal-title":"The Journal of neuroscience: the official journal of the Society for Neuroscience"},{"issue":"5293","key":"ref34","doi-asserted-by":"crossref","first-page":"1724","DOI":"10.1126\/science.274.5293.1724","article-title":"Chaos in neuronal networks with balanced excitatory and inhibitory activity","volume":"274","author":"Cv Vreeswijk","year":"1996","journal-title":"Science (New York, NY)"},{"issue":"6","key":"ref35","doi-asserted-by":"crossref","first-page":"e179","DOI":"10.1371\/journal.pbio.0040179","article-title":"Interacting Adaptive Processes with Different Timescales Underlie Short-Term Motor Learning","volume":"4","author":"MA Smith","year":"2006","journal-title":"PLoS Biology"},{"issue":"1","key":"ref36","first-page":"105","article-title":"Collective dynamics in human and monkey sensorimotor cortex: predicting single neuron spikes","volume":"13","author":"W Truccolo","year":"2010","journal-title":"Nature Publishing Group"},{"issue":"2","key":"ref37","doi-asserted-by":"crossref","first-page":"e42","DOI":"10.1371\/journal.pbio.0000042","article-title":"Learning to Control a Brain\u2013Machine Interface for Reaching and Grasping by Primates","volume":"1","author":"JM Carmena","year":"2003","journal-title":"PLoS Biology"},{"issue":"1","key":"ref38","doi-asserted-by":"crossref","first-page":"515","DOI":"10.1152\/jn.00587.2002","article-title":"Spatiotemporal Tuning of Motor Cortical Neurons for Hand Position and Velocity","volume":"91","author":"L Paninski","year":"2004","journal-title":"Journal of Neurophysiology"},{"issue":"6","key":"ref39","doi-asserted-by":"crossref","first-page":"4235","DOI":"10.1152\/jn.00095.2007","article-title":"Temporal Complexity and Heterogeneity of Single-Neuron Activity in Premotor and Motor Cortex","volume":"97","author":"MM Churchland","year":"2007","journal-title":"Journal of Neurophysiology"},{"issue":"2","key":"ref40","doi-asserted-by":"crossref","first-page":"1040","DOI":"10.1152\/jn.91052.2008","article-title":"Synaptic Linkages Between Corticomotoneuronal Cells Affecting Forelimb Muscles in Behaving Primates","volume":"102","author":"WS Smith","year":"2009","journal-title":"Journal of Neurophysiology"},{"issue":"2","key":"ref41","doi-asserted-by":"crossref","first-page":"1026","DOI":"10.1152\/jn.91051.2008","article-title":"Synaptic interactions between forelimb-related motor cortex neurons in behaving primates","volume":"102","author":"WS Smith","year":"2009","journal-title":"Journal of Neurophysiology"},{"issue":"1","key":"ref42","doi-asserted-by":"crossref","first-page":"360","DOI":"10.1152\/jn.00710.2006","article-title":"Correlations Between the Same Motor Cortex Cells and Arm Muscles During a Trained Task, Free Behavior, and Natural Sleep in the Macaque Monkey","volume":"97","author":"A Jackson","year":"2007","journal-title":"Journal of Neurophysiology"},{"issue":"C","key":"ref43","doi-asserted-by":"crossref","first-page":"220","DOI":"10.1016\/j.jneumeth.2015.07.028","article-title":"Strategies for optical control and simultaneous electrical readout of extended cortical circuits","volume":"256","author":"P Ledochowitsch","year":"2015","journal-title":"Journal of neuroscience methods"},{"issue":"19","key":"ref44","doi-asserted-by":"crossref","first-page":"8691","DOI":"10.1523\/JNEUROSCI.22-19-08691.2002","article-title":"Model of Thalamocortical Slow-Wave Sleep Oscillations and Transitions to Activated States","volume":"22","author":"M Bazhenov","year":"2002","journal-title":"The Journal of neuroscience: the official journal of the Society for Neuroscience"},{"issue":"12","key":"ref45","doi-asserted-by":"crossref","first-page":"5261","DOI":"10.1523\/JNEUROSCI.4683-12.2013","article-title":"Myo-cortical crossed feedback reorganizes primate motor cortex output","volume":"33","author":"T Lucas","year":"2013","journal-title":"The Journal of Neuroscience"},{"key":"ref46","unstructured":"Seeman S, Mogen BJ, Fetz EE, Perlmutter S. Paired stimulation induces spike-timing dependent plasticity of neural connections in primate sensorimotor cortex. In: Society for Neuroscience Annual Meeting proceedings; 2015."},{"key":"ref47","unstructured":"Silversmith DB, Yazdan-Shahmorad A, Sabes PN. Targeted cortical reorganization using optogenetics in non-human primates. In: Society for Neuroscience Annual Meeting proceedings; 2016."},{"issue":"6300","key":"ref48","doi-asserted-by":"crossref","first-page":"691","DOI":"10.1126\/science.aaf7560","article-title":"Imprinting and recalling cortical ensembles","volume":"353","author":"L Carrillo-Reid","year":"2016","journal-title":"Science (New York, NY)"},{"key":"ref49","doi-asserted-by":"crossref","DOI":"10.3389\/fncir.2014.00115","article-title":"Closed-loop neuroscience and neuroengineering","volume":"8","author":"SM Potter","year":"2014","journal-title":"Frontiers in Neural Circuits"},{"issue":"3","key":"ref50","doi-asserted-by":"crossref","first-page":"335","DOI":"10.1007\/s00221-012-3315-1","article-title":"New modalities of brain stimulation for stroke rehabilitation","volume":"224","author":"MA Edwardson","year":"2013","journal-title":"Experimental Brain Research"},{"issue":"52","key":"ref51","doi-asserted-by":"crossref","first-page":"21177","DOI":"10.1073\/pnas.1316885110","article-title":"Restoration of function after brain damage using a neural prosthesis","volume":"110","author":"DJ Guggenmos","year":"2013","journal-title":"Proceedings of the National Academy of Sciences of the United States of America"},{"key":"ref52","article-title":"Spike-timing dependent plasticity in inhibitory circuits","volume":"2","author":"KP Lamsa","year":"2010","journal-title":"Frontiers in Synaptic Neuroscience"},{"issue":"24","key":"ref53","doi-asserted-by":"crossref","first-page":"8870","DOI":"10.1523\/JNEUROSCI.6200-10.2011","article-title":"Ionic Dynamics Mediate Spontaneous Termination of Seizures and Postictal Depression State","volume":"31","author":"GP Krishnan","year":"2011","journal-title":"The Journal of neuroscience: the official journal of the Society for Neuroscience"},{"issue":"6062","key":"ref54","doi-asserted-by":"crossref","first-page":"1569","DOI":"10.1126\/science.1211095","article-title":"Inhibitory plasticity balances excitation and inhibition in sensory pathways and memory networks","volume":"334","author":"TP Vogels","year":"2011","journal-title":"Science (New York, NY)"},{"key":"ref55","doi-asserted-by":"crossref","first-page":"5319","DOI":"10.1038\/ncomms6319","article-title":"Formation and maintenance of neuronal assemblies through synaptic plasticity","volume":"5","author":"A Litwin-Kumar","year":"2014","journal-title":"Nature Communications"},{"issue":"38","key":"ref56","doi-asserted-by":"crossref","first-page":"9673","DOI":"10.1523\/JNEUROSCI.1425-06.2006","article-title":"Triplets of Spikes in a Model of Spike Timing-Dependent Plasticity","volume":"26","author":"JP Pfister","year":"2006","journal-title":"The Journal of neuroscience: the official journal of the Society for Neuroscience"},{"key":"ref57","article-title":"Spike-timing-dependent synaptic modification induced by natural spike trains","author":"R Froemke","year":"2002","journal-title":"Nature"},{"key":"ref58","doi-asserted-by":"crossref","unstructured":"Ocker GK, Shea-Brown E, Buice MA. Linking structure and activity in nonlinear spiking networks. arXivorg. 2016;.","DOI":"10.1101\/080705"},{"issue":"6","key":"ref59","doi-asserted-by":"crossref","first-page":"61919","DOI":"10.1103\/PhysRevE.72.061919","article-title":"Theory of oscillatory firing induced by spatially correlated noise and delayed inhibitory feedback","volume":"72","author":"B Lindner","year":"2005","journal-title":"Physical Review E"},{"issue":"8","key":"ref60","doi-asserted-by":"crossref","first-page":"e1004458","DOI":"10.1371\/journal.pcbi.1004458","article-title":"Self-Organization of Microcircuits in Networks of Spiking Neurons with Plastic Synapses","volume":"11","author":"GK Ocker","year":"2015","journal-title":"PLoS Computational Biology"},{"issue":"2","key":"ref61","first-page":"187","article-title":"Coactivation and timing-dependent integration of synaptic potentiation and depression","volume":"8","author":"HX Wang","year":"2005","journal-title":"Nature Publishing Group"},{"issue":"24","key":"ref62","doi-asserted-by":"crossref","first-page":"10464","DOI":"10.1523\/JNEUROSCI.18-24-10464.1998","article-title":"Synaptic modifications in cultured hippocampal neurons: dependence on spike timing, synaptic strength, and postsynaptic cell type","volume":"18","author":"GQ Bi","year":"1998","journal-title":"The Journal of neuroscience: the official journal of the Society for Neuroscience"},{"issue":"7","key":"ref63","doi-asserted-by":"crossref","first-page":"917","DOI":"10.1016\/j.neunet.2004.03.003","article-title":"Impact of deviation from precise balance of spike-timing-dependent plasticity","volume":"17","author":"N Matsumoto","year":"2004","journal-title":"Neural Networks"},{"issue":"3","key":"ref64","doi-asserted-by":"crossref","first-page":"438","DOI":"10.1111\/j.2517-6161.1971.tb01530.x","article-title":"Point Spectra of Some Mutually Exciting Point Processes","volume":"33","author":"AG Hawkes","year":"1971","journal-title":"Journal of the Royal Statistical Society Series B (Methodological)"}],"updated-by":[{"DOI":"10.1371\/journal.pcbi.1005343","type":"new_version","label":"New version","source":"publisher","updated":{"date-parts":[[2017,2,16]],"date-time":"2017-02-16T00:00:00Z","timestamp":1487203200000}}],"container-title":["PLOS Computational Biology"],"original-title":[],"language":"en","link":[{"URL":"http:\/\/dx.plos.org\/10.1371\/journal.pcbi.1005343","content-type":"unspecified","content-version":"vor","intended-application":"similarity-checking"}],"deposited":{"date-parts":[[2019,9,18]],"date-time":"2019-09-18T02:14:17Z","timestamp":1568772857000},"score":1,"resource":{"primary":{"URL":"https:\/\/dx.plos.org\/10.1371\/journal.pcbi.1005343"}},"subtitle":[],"editor":[{"given":"Matthias","family":"Bethge","sequence":"first","affiliation":[],"role":[{"role":"editor","vocabulary":"crossref"}]}],"short-title":[],"issued":{"date-parts":[[2017,2,2]]},"references-count":64,"journal-issue":{"issue":"2","published-online":{"date-parts":[[2017,2,2]]}},"URL":"https:\/\/doi.org\/10.1371\/journal.pcbi.1005343","relation":{"new_version":[{"id-type":"doi","id":"10.1371\/journal.pcbi.1005343","asserted-by":"object"}]},"ISSN":["1553-7358"],"issn-type":[{"value":"1553-7358","type":"electronic"}],"subject":[],"published":{"date-parts":[[2017,2,2]]}}}