{"status":"ok","message-type":"work","message-version":"1.0.0","message":{"indexed":{"date-parts":[[2026,5,21]],"date-time":"2026-05-21T12:44:07Z","timestamp":1779367447815,"version":"3.53.0"},"update-to":[{"DOI":"10.1371\/journal.pcbi.1006029","type":"new_version","label":"New version","source":"publisher","updated":{"date-parts":[[2018,3,13]],"date-time":"2018-03-13T00:00:00Z","timestamp":1520899200000}}],"reference-count":41,"publisher":"Public Library of Science (PLoS)","issue":"3","license":[{"start":{"date-parts":[[2018,3,1]],"date-time":"2018-03-01T00:00:00Z","timestamp":1519862400000},"content-version":"vor","delay-in-days":0,"URL":"http:\/\/creativecommons.org\/licenses\/by\/4.0\/"}],"funder":[{"DOI":"10.13039\/100009619","name":"Japan Agency for Medical Research and Development","doi-asserted-by":"publisher","award":["Platform Project for Supporting in Drug Discovery and Life Science Research"],"award-info":[{"award-number":["Platform Project for Supporting in Drug Discovery and Life Science Research"]}],"id":[{"id":"10.13039\/100009619","id-type":"DOI","asserted-by":"publisher"}]},{"DOI":"10.13039\/501100001691","name":"Japan Society for the Promotion of Science","doi-asserted-by":"publisher","award":["26840077"],"award-info":[{"award-number":["26840077"]}],"id":[{"id":"10.13039\/501100001691","id-type":"DOI","asserted-by":"publisher"}]}],"content-domain":{"domain":["www.ploscompbiol.org"],"crossmark-restriction":false},"short-container-title":["PLoS Comput Biol"],"DOI":"10.1371\/journal.pcbi.1006029","type":"journal-article","created":{"date-parts":[[2018,3,1]],"date-time":"2018-03-01T18:28:35Z","timestamp":1519928915000},"page":"e1006029","update-policy":"https:\/\/doi.org\/10.1371\/journal.pcbi.corrections_policy","source":"Crossref","is-referenced-by-count":12,"title":["Inverse tissue mechanics of cell monolayer expansion"],"prefix":"10.1371","volume":"14","author":[{"ORCID":"https:\/\/orcid.org\/0000-0002-2801-4522","authenticated-orcid":true,"given":"Yohei","family":"Kondo","sequence":"first","affiliation":[],"role":[{"vocabulary":"crossref","role":"author"}]},{"ORCID":"https:\/\/orcid.org\/0000-0001-7263-1555","authenticated-orcid":true,"given":"Kazuhiro","family":"Aoki","sequence":"additional","affiliation":[],"role":[{"vocabulary":"crossref","role":"author"}]},{"given":"Shin","family":"Ishii","sequence":"additional","affiliation":[],"role":[{"vocabulary":"crossref","role":"author"}]}],"member":"340","published-online":{"date-parts":[[2018,3,1]]},"reference":[{"issue":"5","key":"ref1","doi-asserted-by":"crossref","first-page":"948","DOI":"10.1016\/j.cell.2013.05.008","article-title":"Forces in Tissue Morphogenesis and Patterning","volume":"153","author":"CP Heisenberg","year":"2013","journal-title":"Cell"},{"issue":"8","key":"ref2","doi-asserted-by":"crossref","first-page":"633","DOI":"10.1038\/nrm2222","article-title":"Cell surface mechanics and the control of cell shape, tissue patterns and morphogenesis","volume":"8","author":"T Lecuit","year":"2007","journal-title":"Nature reviews Molecular cell biology"},{"issue":"2","key":"ref3","doi-asserted-by":"crossref","first-page":"186","DOI":"10.1242\/dev.119776","article-title":"Measuring forces and stresses in situ in living tissues","volume":"143","author":"K Sugimura","year":"2016","journal-title":"Development"},{"issue":"32","key":"ref4","doi-asserted-by":"crossref","first-page":"12974","DOI":"10.1073\/pnas.1119313109","article-title":"Emerging modes of collective cell migration induced by geometrical constraints","volume":"109","author":"SRK Vedula","year":"2012","journal-title":"Proceedings of the National Academy of Sciences"},{"issue":"9","key":"ref5","doi-asserted-by":"crossref","first-page":"856","DOI":"10.1038\/nmat3689","article-title":"Propulsion and navigation within the advancing monolayer sheet","volume":"12","author":"JH Kim","year":"2013","journal-title":"Nature materials"},{"issue":"3","key":"ref6","doi-asserted-by":"crossref","first-page":"217","DOI":"10.1038\/ncb2917","article-title":"Interplay of RhoA and mechanical forces in collective cell migration driven by leader cells","volume":"16","author":"M Reffay","year":"2014","journal-title":"Nat Cell Biol"},{"issue":"2","key":"ref7","doi-asserted-by":"crossref","first-page":"79","DOI":"10.1016\/S0962-8924(01)02205-X","article-title":"Flexible substrata for the detection of cellular traction forces","volume":"12","author":"KA Beningo","year":"2002","journal-title":"Trends in Cell Biology"},{"issue":"6","key":"ref8","doi-asserted-by":"crossref","first-page":"426","DOI":"10.1038\/nphys1269","article-title":"Physical forces during collective cell migration","volume":"5","author":"X Trepat","year":"2009","journal-title":"Nature Physics"},{"issue":"6","key":"ref9","doi-asserted-by":"crossref","first-page":"469","DOI":"10.1038\/nmat3025","article-title":"Collective cell guidance by cooperative intercellular forces","volume":"10","author":"DT Tambe","year":"2011","journal-title":"Nature materials"},{"issue":"7","key":"ref10","doi-asserted-by":"crossref","first-page":"1625","DOI":"10.1016\/j.bpj.2016.03.002","article-title":"Inference of internal stress in a cell monolayer","volume":"110","author":"V Nier","year":"2016","journal-title":"Biophysical Journal"},{"issue":"41","key":"ref11","doi-asserted-by":"crossref","first-page":"17271","DOI":"10.1073\/pnas.0902085106","article-title":"The role of fluctuations and stress on the effective viscosity of cell aggregates","volume":"106","author":"P Marmottant","year":"2009","journal-title":"Proceedings of the National Academy of Sciences"},{"key":"ref12","doi-asserted-by":"crossref","first-page":"218101","DOI":"10.1103\/PhysRevLett.104.218101","article-title":"Aspiration of Biological Viscoelastic Drops","volume":"104","author":"K Guevorkian","year":"2010","journal-title":"Phys Rev Lett"},{"issue":"75","key":"ref13","doi-asserted-by":"crossref","first-page":"2614","DOI":"10.1098\/rsif.2012.0263","article-title":"Mechanical state, material properties and continuous description of an epithelial tissue","volume":"9","author":"I Bonnet","year":"2012","journal-title":"Journal of The Royal Society Interface"},{"issue":"41","key":"ref14","doi-asserted-by":"crossref","first-page":"16449","DOI":"10.1073\/pnas.1213301109","article-title":"Characterizing the mechanics of cultured cell monolayers","volume":"109","author":"aR Harris","year":"2012","journal-title":"Proceedings of the National Academy of Sciences"},{"issue":"5","key":"ref15","doi-asserted-by":"crossref","first-page":"33","DOI":"10.1140\/epje\/i2015-15033-4","article-title":"Colloquium: Mechanical formalisms for tissue dynamics","volume":"38","author":"S Tlili","year":"2015","journal-title":"The European Physical Journal E"},{"key":"ref16","first-page":"47","article-title":"Cell Mechanics. vol. 83 of Methods in Cell Biology","author":"MT Frey","year":"2007"},{"issue":"11","key":"ref17","doi-asserted-by":"crossref","first-page":"1463","DOI":"10.1093\/bioinformatics\/btp184","article-title":"Globally optimal stitching of tiled 3D microscopic image acquisitions","volume":"25","author":"S Preibisch","year":"2009","journal-title":"Bioinformatics"},{"issue":"2","key":"ref18","doi-asserted-by":"crossref","first-page":"156","DOI":"10.1364\/OL.33.000156","article-title":"Efficient subpixel image registration algorithms","volume":"33","author":"M Guizar-Sicairos","year":"2008","journal-title":"Opt Lett"},{"issue":"3","key":"ref19","doi-asserted-by":"crossref","first-page":"500","DOI":"10.1109\/TPAMI.2010.143","article-title":"Large displacement optical flow: descriptor matching in variational motion estimation","volume":"33","author":"T Brox","year":"2011","journal-title":"IEEE Transactions on Pattern Analysis and Machine Intelligence"},{"issue":"10","key":"ref20","doi-asserted-by":"crossref","first-page":"3835","DOI":"10.1073\/pnas.0607134104","article-title":"On the mechanism of wing size determination in fly development","volume":"104","author":"L Hufnagel","year":"2007","journal-title":"Proceedings of the National Academy of Sciences"},{"issue":"1","key":"ref21","doi-asserted-by":"crossref","first-page":"65","DOI":"10.1016\/j.bpj.2013.11.015","article-title":"Border Forces and Friction Control Epithelial Closure Dynamics","volume":"106","author":"O Cochet-Escartin","year":"2014","journal-title":"Biophysical Journal"},{"issue":"49","key":"ref22","doi-asserted-by":"crossref","first-page":"20863","DOI":"10.1073\/pnas.1011086107","article-title":"Fluidization of tissues by cell division and apoptosis","volume":"107","author":"J Ranft","year":"2010","journal-title":"Proceedings of the National Academy of Sciences"},{"key":"ref23","doi-asserted-by":"crossref","first-page":"7328","DOI":"10.1039\/C5SM01382H","article-title":"Celebrating Soft Matter\u2019s 10th Anniversary: Cell division: a source of active stress in cellular monolayers","volume":"11","author":"A Doostmohammadi","year":"2015","journal-title":"Soft Matter"},{"issue":"41","key":"ref24","doi-asserted-by":"crossref","first-page":"15988","DOI":"10.1073\/pnas.0705062104","article-title":"Collective migration of an epithelial monolayer in response to a model wound","volume":"104","author":"M Poujade","year":"2007","journal-title":"Proceedings of the National Academy of Sciences"},{"issue":"2","key":"ref25","doi-asserted-by":"crossref","first-page":"1","DOI":"10.1371\/journal.pone.0055172","article-title":"Monolayer Stress Microscopy: Limitations, Artifacts, and Accuracy of Recovered Intercellular Stresses","volume":"8","author":"DT Tambe","year":"2013","journal-title":"PLoS ONE"},{"issue":"1","key":"ref26","doi-asserted-by":"crossref","first-page":"1","DOI":"10.1111\/j.2517-6161.1977.tb01600.x","article-title":"Maximum likelihood from incomplete data via the EM algorithm","volume":"39","author":"AP Dempster","year":"1977","journal-title":"Journal of the Royal Statistical Society, Series B"},{"issue":"8","key":"ref27","doi-asserted-by":"crossref","first-page":"1445","DOI":"10.2514\/3.3166","article-title":"Maximum likelihood estimates of linear dynamic systems","volume":"3","author":"HE Rauch","year":"1965","journal-title":"AIAA Journal"},{"key":"ref28","doi-asserted-by":"crossref","first-page":"2381","DOI":"10.1039\/C5SM02857D","article-title":"One-dimensional collective migration of a proliferating cell monolayer","volume":"12","author":"P Recho","year":"2016","journal-title":"Soft Matter"},{"issue":"9","key":"ref29","doi-asserted-by":"crossref","first-page":"1790","DOI":"10.1016\/j.bpj.2010.01.030","article-title":"Velocity Fields in a Collectively Migrating Epithelium","volume":"98","author":"L Petitjean","year":"2010","journal-title":"Biophysical Journal"},{"issue":"3","key":"ref30","doi-asserted-by":"crossref","first-page":"276","DOI":"10.1038\/ncb3115","article-title":"A molecular mechanotransduction pathway regulates collective migration of epithelial cells","volume":"17","author":"T Das","year":"2015","journal-title":"Nat Cell Biol"},{"key":"ref31","doi-asserted-by":"crossref","first-page":"248103","DOI":"10.1103\/PhysRevLett.115.248103","article-title":"Active Tensile Modulus of an Epithelial Monolayer","volume":"115","author":"R Vincent","year":"2015","journal-title":"Phys Rev Lett"},{"key":"ref32","article-title":"Biomechanics: Mechanical Properties of Living Tissues","author":"YC Fung","year":"1993"},{"issue":"4","key":"ref33","doi-asserted-by":"crossref","first-page":"863","DOI":"10.1016\/j.jmps.2010.12.011","article-title":"Perspectives on biological growth and remodeling","volume":"59","author":"D Ambrosi","year":"2011","journal-title":"Journal of the Mechanics and Physics of Solids"},{"issue":"26","key":"ref34","doi-asserted-by":"crossref","first-page":"10525","DOI":"10.1073\/pnas.1217391110","article-title":"Anisotropic growth shapes intestinal tissues during embryogenesis","volume":"110","author":"M Ben Amar","year":"2013","journal-title":"Proceedings of the National Academy of Sciences"},{"issue":"1","key":"ref35","doi-asserted-by":"crossref","first-page":"95","DOI":"10.1007\/s00424-007-0419-8","article-title":"Softening of the actin cytoskeleton by inhibition of myosin II","volume":"456","author":"JC Martens","year":"2008","journal-title":"Pfl\u00fcgers Archiv\u2014European Journal of Physiology"},{"issue":"15","key":"ref36","doi-asserted-by":"crossref","first-page":"5075","DOI":"10.1158\/0008-5472.CAN-11-0247","article-title":"Mechanical Stiffness Grades Metastatic Potential in Patient Tumor Cells and in Cancer Cell Lines","volume":"71","author":"V Swaminathan","year":"2011","journal-title":"Cancer Research"},{"key":"ref37","doi-asserted-by":"crossref","first-page":"028102","DOI":"10.1103\/PhysRevLett.113.028102","article-title":"Stress Reorganization and Response in Active Solids","volume":"113","author":"RJ Hawkins","year":"2014","journal-title":"Phys Rev Lett"},{"issue":"19","key":"ref38","doi-asserted-by":"crossref","first-page":"4091","DOI":"10.1242\/dev.094060","article-title":"The mechanical anisotropy in a tissue promotes ordering in hexagonal cell packing","volume":"140","author":"K Sugimura","year":"2013","journal-title":"Development"},{"issue":"2","key":"ref39","doi-asserted-by":"crossref","first-page":"183","DOI":"10.1038\/nmeth.2761","article-title":"Quantifying cell-generated mechanical forces within living embryonic tissues","volume":"11","author":"O Campas","year":"2014","journal-title":"Nat Meth"},{"key":"ref40","doi-asserted-by":"crossref","first-page":"e08519","DOI":"10.7554\/eLife.08519","article-title":"Unified quantitative characterization of epithelial tissue development","volume":"4","author":"B Guirao","year":"2015","journal-title":"eLife"},{"key":"ref41","doi-asserted-by":"crossref","first-page":"e14334","DOI":"10.7554\/eLife.14334","article-title":"TissueMiner: A multiscale analysis toolkit to quantify how cellular processes create tissue dynamics","volume":"5","author":"R Etournay","year":"2016","journal-title":"eLife"}],"updated-by":[{"DOI":"10.1371\/journal.pcbi.1006029","type":"new_version","label":"New version","source":"publisher","updated":{"date-parts":[[2018,3,13]],"date-time":"2018-03-13T00:00:00Z","timestamp":1520899200000}}],"container-title":["PLOS Computational Biology"],"original-title":[],"language":"en","link":[{"URL":"http:\/\/dx.plos.org\/10.1371\/journal.pcbi.1006029","content-type":"unspecified","content-version":"vor","intended-application":"similarity-checking"}],"deposited":{"date-parts":[[2024,7,1]],"date-time":"2024-07-01T17:54:44Z","timestamp":1719856484000},"score":1,"resource":{"primary":{"URL":"https:\/\/dx.plos.org\/10.1371\/journal.pcbi.1006029"}},"subtitle":[],"editor":[{"given":"Philip K","family":"Maini","sequence":"first","affiliation":[],"role":[{"vocabulary":"crossref","role":"editor"}]}],"short-title":[],"issued":{"date-parts":[[2018,3,1]]},"references-count":41,"journal-issue":{"issue":"3","published-online":{"date-parts":[[2018,3,1]]}},"URL":"https:\/\/doi.org\/10.1371\/journal.pcbi.1006029","relation":{"new_version":[{"id-type":"doi","id":"10.1371\/journal.pcbi.1006029","asserted-by":"object"}]},"ISSN":["1553-7358"],"issn-type":[{"value":"1553-7358","type":"electronic"}],"subject":[],"published":{"date-parts":[[2018,3,1]]}}}