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neocortical neurons in vivo","volume":"4","author":"A Destexhe","year":"2003","journal-title":"Nature Reviews Neuroscience"},{"issue":"10","key":"ref54","doi-asserted-by":"crossref","first-page":"2345","DOI":"10.1523\/JNEUROSCI.3349-03.2004","article-title":"Neuronal integration of synaptic input in the fluctuation-driven regime","volume":"24","author":"A Kuhn","year":"2004","journal-title":"Journal of Neuroscience"},{"issue":"22","key":"ref55","doi-asserted-by":"crossref","first-page":"5216","DOI":"10.1523\/JNEUROSCI.0540-04.2004","article-title":"Neuronal avalanches are diverse and precise activity patterns that are stable for many hours in cortical slice cultures","volume":"24","author":"JM Beggs","year":"2004","journal-title":"Journal of Neuroscience"},{"issue":"3","key":"ref56","doi-asserted-by":"crossref","first-page":"1126","DOI":"10.1152\/jn.00093.2008","article-title":"Data-driven significance estimation for precise spike correlation","volume":"101","author":"S Grun","year":"2009","journal-title":"Journal of Neurophysiology"},{"key":"ref57","doi-asserted-by":"crossref","first-page":"39","DOI":"10.1016\/j.brainres.2008.04.073","article-title":"Effectiveness of systematic spike dithering depends on the precision of cortical synchronization","volume":"1225","author":"A Pazienti","year":"2008","journal-title":"Brain Research"},{"key":"ref58","doi-asserted-by":"crossref","first-page":"485","DOI":"10.1038\/srep00485","article-title":"Optimal spike-based communication in excitable networks with strong-sparse and weak-dense links","volume":"2","author":"Jn Teramae","year":"2012","journal-title":"Scientific Reports"},{"key":"ref59","doi-asserted-by":"crossref","first-page":"379","DOI":"10.1016\/j.neuroimage.2015.09.009","article-title":"The contribution of geometry to the human connectome","volume":"124","author":"JA Roberts","year":"2016","journal-title":"Neuroimage"},{"issue":"4","key":"ref60","doi-asserted-by":"crossref","first-page":"e94292","DOI":"10.1371\/journal.pone.0094292","article-title":"Dense neuron clustering explains connectivity statistics in cortical microcircuits","volume":"9","author":"VV Klinshov","year":"2014","journal-title":"PloS one"},{"issue":"2","key":"ref61","doi-asserted-by":"crossref","first-page":"021039","DOI":"10.1103\/PhysRevX.4.021039","article-title":"Balanced networks of spiking neurons with spatially dependent recurrent connections","volume":"4","author":"R Rosenbaum","year":"2014","journal-title":"Physical Review X"},{"key":"ref62","doi-asserted-by":"crossref","first-page":"8","DOI":"10.3389\/fncom.2011.00008","article-title":"The role of degree distribution in shaping the dynamics in networks of sparsely connected spiking neurons","volume":"5","author":"A Roxin","year":"2011","journal-title":"Frontiers in Computational 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Inference, validation and predictions about statistics and propagation of cortical spiking in vivo. bioRxiv. 2018.","DOI":"10.1101\/363085"},{"issue":"5","key":"ref70","doi-asserted-by":"crossref","first-page":"907","DOI":"10.1016\/j.neuron.2007.10.007","article-title":"Spatiotemporal dynamics of cortical sensorimotor integration in behaving mice","volume":"56","author":"I Ferezou","year":"2007","journal-title":"Neuron"},{"issue":"43","key":"ref71","doi-asserted-by":"crossref","first-page":"14585","DOI":"10.1523\/JNEUROSCI.4944-14.2015","article-title":"A lognormal recurrent network model for burst generation during hippocampal sharp waves","volume":"35","author":"Y Omura","year":"2015","journal-title":"The Journal of Neuroscience"},{"issue":"4","key":"ref72","doi-asserted-by":"crossref","first-page":"264","DOI":"10.1038\/nrn3687","article-title":"The log-dynamic brain: how skewed distributions affect network operations","volume":"15","author":"G Buzs\u00e1ki","year":"2014","journal-title":"Nature Reviews Neuroscience"},{"issue":"3","key":"ref73","doi-asserted-by":"crossref","first-page":"388","DOI":"10.1002\/cne.902610305","article-title":"GABA immunoreactive neurons in rat visual cortex","volume":"261","author":"DL Meinecke","year":"1987","journal-title":"Journal of Comparative Neurology"},{"issue":"3","key":"ref74","doi-asserted-by":"crossref","first-page":"263","DOI":"10.1002\/cne.902380303","article-title":"The neuronal composition of area 17 of rat visual cortex. III. Numerical considerations","volume":"238","author":"A Peters","year":"1985","journal-title":"Journal of Comparative Neurology"},{"issue":"6","key":"ref75","doi-asserted-by":"crossref","first-page":"1188","DOI":"10.1016\/j.neuron.2011.02.025","article-title":"Dense inhibitory connectivity in neocortex","volume":"69","author":"E Fino","year":"2011","journal-title":"Neuron"},{"issue":"1","key":"ref76","doi-asserted-by":"crossref","first-page":"139","DOI":"10.1113\/jphysiol.2003.044784","article-title":"Pyramidal cell communication within local networks in layer 2\/3 of rat neocortex","volume":"551","author":"C Holmgren","year":"2003","journal-title":"The Journal of physiology"},{"issue":"36","key":"ref77","doi-asserted-by":"crossref","first-page":"15927","DOI":"10.1073\/pnas.1010356107","article-title":"Neuron densities vary across and within cortical areas in primates","volume":"107","author":"CE Collins","year":"2010","journal-title":"Proceedings of the National Academy of Sciences"},{"key":"ref78","doi-asserted-by":"crossref","DOI":"10.1093\/oso\/9780195104912.001.0001","article-title":"Biophysics of Computation: Information Processing in Single Neurons","author":"C Koch","year":"1998"},{"key":"ref79","doi-asserted-by":"crossref","first-page":"319","DOI":"10.1113\/jphysiol.1984.sp015378","article-title":"Control of the repetitive discharge of rat CA1 pyramidal neurones in vitro","volume":"354","author":"D Madison","year":"1984","journal-title":"The Journal of Physiology"},{"issue":"3","key":"ref80","doi-asserted-by":"crossref","first-page":"259","DOI":"10.1088\/0954-898X_4_3_002","article-title":"Mean-field analysis of neuronal spike dynamics","volume":"4","author":"A Treves","year":"1993","journal-title":"Network: Computation in Neural Systems"},{"issue":"1","key":"ref81","doi-asserted-by":"crossref","first-page":"1","DOI":"10.1016\/0166-4328(94)00113-T","article-title":"Neuromodulation and cortical function: modeling the physiological basis of behavior","volume":"67","author":"ME Hasselmo","year":"1995","journal-title":"Behavioural Brain Research"},{"issue":"3","key":"ref82","doi-asserted-by":"crossref","first-page":"1549","DOI":"10.1152\/jn.1998.79.3.1549","article-title":"Calcium coding and adaptive temporal computation in cortical pyramidal neurons","volume":"79","author":"XJ Wang","year":"1998","journal-title":"Journal of Neurophysiology"},{"key":"ref83","first-page":"1","article-title":"Methods in Neuronal Modeling","author":"A Destexhe","year":"1998"},{"issue":"21","key":"ref84","doi-asserted-by":"crossref","first-page":"9587","DOI":"10.1523\/JNEUROSCI.19-21-09587.1999","article-title":"Synaptic basis of cortical persistent activity: the importance of NMDA receptors to working memory","volume":"19","author":"XJ Wang","year":"1999","journal-title":"Journal of Neuroscience"},{"issue":"8","key":"ref85","doi-asserted-by":"crossref","first-page":"e1000456","DOI":"10.1371\/journal.pcbi.1000456","article-title":"Towards reproducible descriptions of neuronal network models","volume":"5","author":"E Nordlie","year":"2009","journal-title":"PLoS Computational Biology"},{"issue":"430","key":"ref86","doi-asserted-by":"crossref","first-page":"773","DOI":"10.1080\/01621459.1995.10476572","article-title":"Bayes factors","volume":"90","author":"RE Kass","year":"1995","journal-title":"Journal of the American Statistical Association"}],"updated-by":[{"DOI":"10.1371\/journal.pcbi.1006902","type":"new_version","label":"New version","source":"publisher","updated":{"date-parts":[[2019,4,12]],"date-time":"2019-04-12T00:00:00Z","timestamp":1555027200000}}],"container-title":["PLOS Computational Biology"],"original-title":[],"language":"en","link":[{"URL":"http:\/\/dx.plos.org\/10.1371\/journal.pcbi.1006902","content-type":"unspecified","content-version":"vor","intended-application":"similarity-checking"}],"deposited":{"date-parts":[[2020,12,5]],"date-time":"2020-12-05T10:35:55Z","timestamp":1607164555000},"score":1,"resource":{"primary":{"URL":"https:\/\/dx.plos.org\/10.1371\/journal.pcbi.1006902"}},"subtitle":[],"editor":[{"given":"Jeff","family":"Beck","sequence":"first","affiliation":[],"role":[{"vocabulary":"crossref","role":"editor"}]}],"short-title":[],"issued":{"date-parts":[[2019,4,2]]},"references-count":86,"journal-issue":{"issue":"4","published-online":{"date-parts":[[2019,4,2]]}},"URL":"https:\/\/doi.org\/10.1371\/journal.pcbi.1006902","relation":{"new_version":[{"id-type":"doi","id":"10.1371\/journal.pcbi.1006902","asserted-by":"object"}]},"ISSN":["1553-7358"],"issn-type":[{"value":"1553-7358","type":"electronic"}],"subject":[],"published":{"date-parts":[[2019,4,2]]}}}