{"status":"ok","message-type":"work","message-version":"1.0.0","message":{"indexed":{"date-parts":[[2026,4,13]],"date-time":"2026-04-13T23:26:13Z","timestamp":1776122773401,"version":"3.50.1"},"update-to":[{"DOI":"10.1371\/journal.pcbi.1007787","type":"new_version","label":"New version","source":"publisher","updated":{"date-parts":[[2020,5,5]],"date-time":"2020-05-05T00:00:00Z","timestamp":1588636800000}}],"reference-count":76,"publisher":"Public Library of Science (PLoS)","issue":"4","license":[{"start":{"date-parts":[[2020,4,23]],"date-time":"2020-04-23T00:00:00Z","timestamp":1587600000000},"content-version":"vor","delay-in-days":0,"URL":"http:\/\/creativecommons.org\/licenses\/by\/4.0\/"}],"content-domain":{"domain":["www.ploscompbiol.org"],"crossmark-restriction":false},"short-container-title":["PLoS Comput Biol"],"DOI":"10.1371\/journal.pcbi.1007787","type":"journal-article","created":{"date-parts":[[2020,4,23]],"date-time":"2020-04-23T17:45:43Z","timestamp":1587663943000},"page":"e1007787","update-policy":"https:\/\/doi.org\/10.1371\/journal.pcbi.corrections_policy","source":"Crossref","is-referenced-by-count":24,"title":["Telling ecological networks apart by their structure: An environment-dependent approach"],"prefix":"10.1371","volume":"16","author":[{"ORCID":"https:\/\/orcid.org\/0000-0001-7490-8626","authenticated-orcid":true,"given":"Chuliang","family":"Song","sequence":"first","affiliation":[]},{"ORCID":"https:\/\/orcid.org\/0000-0003-1768-363X","authenticated-orcid":true,"given":"Serguei","family":"Saavedra","sequence":"additional","affiliation":[]}],"member":"340","published-online":{"date-parts":[[2020,4,23]]},"reference":[{"key":"pcbi.1007787.ref001","doi-asserted-by":"crossref","DOI":"10.1515\/9781400883790","volume-title":"The theory of ecological communities (MPB-57)","author":"M Vellend","year":"2016"},{"issue":"5993","key":"pcbi.1007787.ref002","doi-asserted-by":"crossref","first-page":"765","DOI":"10.1126\/science.1194255","article-title":"Structure and dynamics of ecological networks","volume":"329","author":"J Bascompte","year":"2010","journal-title":"Science"},{"issue":"5993","key":"pcbi.1007787.ref003","doi-asserted-by":"crossref","first-page":"853","DOI":"10.1126\/science.1188321","article-title":"Stability of ecological communities and the architecture of mutualistic and trophic networks","volume":"329","author":"E Th\u00e9bault","year":"2010","journal-title":"Science"},{"key":"pcbi.1007787.ref004","doi-asserted-by":"crossref","DOI":"10.23943\/princeton\/9780691131269.001.0001","volume-title":"Mutualistic Networks","author":"J Bascompte","year":"2013"},{"key":"pcbi.1007787.ref005","doi-asserted-by":"crossref","first-page":"9383","DOI":"10.1073\/pnas.1633576100","article-title":"The nested assembly of plant-animal mutualistic networks","volume":"100","author":"J Bascompte","year":"2003","journal-title":"Proceedings of the National Academy of Sciences"},{"key":"pcbi.1007787.ref006","volume-title":"Food webs (MPB-50)","author":"KS McCann","year":"2011"},{"issue":"9","key":"pcbi.1007787.ref007","doi-asserted-by":"crossref","first-page":"3648","DOI":"10.1073\/pnas.1014353108","article-title":"Compartmentalization increases food-web persistence","volume":"108","author":"DB Stouffer","year":"2011","journal-title":"Proceedings of the National Academy of Sciences"},{"issue":"1802","key":"pcbi.1007787.ref008","doi-asserted-by":"crossref","first-page":"20142925","DOI":"10.1098\/rspb.2014.2925","article-title":"Geographical variation in mutualistic networks: similarity, turnover and partner fidelity","volume":"282","author":"K Tr\u00f8jelsgaard","year":"2015","journal-title":"Proceedings of the Royal Society B: Biological Sciences"},{"issue":"1","key":"pcbi.1007787.ref009","doi-asserted-by":"crossref","first-page":"21","DOI":"10.1002\/ecy.2063","article-title":"Interaction frequency, network position, and the temporal persistence of interactions in a plant\u2013pollinator network","volume":"99","author":"NP Chacoff","year":"2018","journal-title":"Ecology"},{"issue":"3","key":"pcbi.1007787.ref010","doi-asserted-by":"crossref","first-page":"031024","DOI":"10.1103\/PhysRevX.9.031024","article-title":"Breaking the Spell of Nestedness: The Entropic Origin of Nestedness in Mutualistic Systems","volume":"9","author":"C Payrat\u00f3-Borr\u00e0s","year":"2019","journal-title":"Physical Review X"},{"key":"pcbi.1007787.ref011","doi-asserted-by":"crossref","first-page":"1243","DOI":"10.1111\/ele.13289","article-title":"Superorganisms or loose collections of species? A unifying theory of community patterns along environmental gradients","volume":"22","author":"K Liautaud","year":"2019","journal-title":"Ecology Letters"},{"issue":"6","key":"pcbi.1007787.ref012","doi-asserted-by":"crossref","first-page":"1","DOI":"10.1371\/journal.pcbi.1007076","article-title":"Telling ecological networks apart by their structure: A computational challenge","volume":"15","author":"MJ Michalska-Smith","year":"2019","journal-title":"PLOS Computational Biology"},{"key":"pcbi.1007787.ref013","first-page":"78","volume-title":"Theoretical Ecology. Principles and Applications","author":"R May","year":"1981"},{"issue":"7388","key":"pcbi.1007787.ref014","doi-asserted-by":"crossref","first-page":"205","DOI":"10.1038\/nature10832","article-title":"Stability criteria for complex ecosystems","volume":"483","author":"S Allesina","year":"2012","journal-title":"Nature"},{"issue":"6092","key":"pcbi.1007787.ref015","doi-asserted-by":"crossref","first-page":"349","DOI":"10.1126\/science.1220529","article-title":"Diversity of interaction types and ecological community stability","volume":"337","author":"A Mougi","year":"2012","journal-title":"Science"},{"key":"pcbi.1007787.ref016","doi-asserted-by":"crossref","first-page":"2156","DOI":"10.1073\/pnas.1710352115","article-title":"Generic assembly patterns in complex ecological communities","volume":"115","author":"M Barbier","year":"2018","journal-title":"Proceedings of the National Academy of Sciences"},{"key":"pcbi.1007787.ref017","doi-asserted-by":"crossref","first-page":"e02708","DOI":"10.1002\/ecy.2708","article-title":"Coexistence and competitive exclusion in mutualism","volume":"100","author":"CA Johnson","year":"2019","journal-title":"Ecology"},{"key":"pcbi.1007787.ref018","article-title":"The balance of interaction types determines the assembly and stability of ecological communities","author":"JJ Qian","year":"2019","journal-title":"BioRxiv"},{"issue":"5663","key":"pcbi.1007787.ref019","doi-asserted-by":"crossref","first-page":"1538","DOI":"10.1126\/science.1089167","article-title":"Superfamilies of evolved and designed networks","volume":"303","author":"R Milo","year":"2004","journal-title":"Science"},{"issue":"5","key":"pcbi.1007787.ref020","doi-asserted-by":"crossref","first-page":"808","DOI":"10.1111\/1365-2656.12964","article-title":"Beware z-scores","volume":"88","author":"C Song","year":"2019","journal-title":"Journal of Animal Ecology"},{"issue":"2","key":"pcbi.1007787.ref021","doi-asserted-by":"crossref","first-page":"785","DOI":"10.1111\/brv.12366","article-title":"Comparing species interaction networks along environmental gradients","volume":"93","author":"L Pellissier","year":"2018","journal-title":"Biological Reviews"},{"key":"pcbi.1007787.ref022","doi-asserted-by":"crossref","DOI":"10.1515\/9780691209418","volume-title":"Evolution in Changing Environments: Some Theoretical Explorations","author":"R Levins","year":"1968"},{"key":"pcbi.1007787.ref023","doi-asserted-by":"crossref","first-page":"21","DOI":"10.1016\/S0016-6995(89)80006-3","article-title":"The logic of monsters: Evidence for internal constraint in development and evolution","volume":"22","author":"P Alberch","year":"1989","journal-title":"Geobios"},{"issue":"6","key":"pcbi.1007787.ref024","doi-asserted-by":"crossref","first-page":"429","DOI":"10.1016\/j.tree.2017.03.004","article-title":"Should environmental filtering be abandoned?","volume":"32","author":"MW Cadotte","year":"2017","journal-title":"Trends in Ecology & Evolution"},{"issue":"6","key":"pcbi.1007787.ref025","doi-asserted-by":"crossref","first-page":"912","DOI":"10.1038\/s41559-019-0879-1","article-title":"Non-parametric estimation of the structural stability of non-equilibrium community dynamics","volume":"3","author":"S Cenci","year":"2019","journal-title":"Nature ecology & evolution"},{"key":"pcbi.1007787.ref026","doi-asserted-by":"crossref","first-page":"844","DOI":"10.1038\/nature00812","article-title":"Positive interactions among alpine plants increase with stress","volume":"417","author":"RM Callaway","year":"2002","journal-title":"Nature"},{"key":"pcbi.1007787.ref027","doi-asserted-by":"crossref","first-page":"881","DOI":"10.1111\/ele.12279","article-title":"How context dependent are species interactions?","volume":"17","author":"SA Chamberlain","year":"2014","journal-title":"Ecology Letters"},{"issue":"2","key":"pcbi.1007787.ref028","doi-asserted-by":"crossref","first-page":"326","DOI":"10.1002\/ece3.1371","article-title":"Structure of trophic and mutualistic networks across broad environmental gradients","volume":"5","author":"EA Welti","year":"2015","journal-title":"Ecology and evolution"},{"key":"pcbi.1007787.ref029","doi-asserted-by":"crossref","first-page":"25","DOI":"10.1146\/annurev-ecolsys-110316-022821","article-title":"Ecological networks across environmental gradients","volume":"48","author":"JM Tylianakis","year":"2017","journal-title":"Annual Review of Ecology, Evolution, and Systematics"},{"issue":"14","key":"pcbi.1007787.ref030","doi-asserted-by":"crossref","first-page":"6852","DOI":"10.1002\/ece3.4252","article-title":"Rethinking the importance of the structure of ecological networks under an environment-dependent framework","volume":"8","author":"S Cenci","year":"2018","journal-title":"Ecology & Evolution"},{"key":"pcbi.1007787.ref031","doi-asserted-by":"crossref","DOI":"10.2307\/j.ctvcm4j72","volume-title":"Mostly harmless econometrics: An empiricist\u2019s companion","author":"JD Angrist","year":"2008"},{"issue":"6","key":"pcbi.1007787.ref032","doi-asserted-by":"crossref","first-page":"1417","DOI":"10.1111\/1365-2656.12749","article-title":"Why are some plant\u2013pollinator networks more nested than others?","volume":"86","author":"C Song","year":"2017","journal-title":"Journal of Animal Ecology"},{"issue":"12","key":"pcbi.1007787.ref033","doi-asserted-by":"crossref","first-page":"4302","DOI":"10.1002\/joc.5086","article-title":"WorldClim 2: new 1-km spatial resolution climate surfaces for global land areas","volume":"37","author":"SE Fick","year":"2017","journal-title":"International Journal of Climatology"},{"key":"pcbi.1007787.ref034","doi-asserted-by":"crossref","first-page":"1391","DOI":"10.1038\/ncomms2422","article-title":"The ghost of nestedness in ecological networks","volume":"4","author":"PP Staniczenko","year":"2013","journal-title":"Nature Communications"},{"issue":"4","key":"pcbi.1007787.ref035","doi-asserted-by":"crossref","first-page":"042822","DOI":"10.1103\/PhysRevE.88.042822","article-title":"Spectral methods for community detection and graph partitioning","volume":"88","author":"ME Newman","year":"2013","journal-title":"Physical Review E"},{"key":"pcbi.1007787.ref036","volume-title":"Numerical Ecology","author":"P Legendre","year":"2012"},{"issue":"47","key":"pcbi.1007787.ref037","doi-asserted-by":"crossref","first-page":"12017","DOI":"10.1073\/pnas.1809088115","article-title":"The geographic mosaic of coevolution in mutualistic networks","volume":"115","author":"LP Medeiros","year":"2018","journal-title":"Proceedings of the National Academy of Sciences"},{"issue":"1","key":"pcbi.1007787.ref038","doi-asserted-by":"crossref","first-page":"27","DOI":"10.1111\/j.1600-0587.2012.07348.x","article-title":"Collinearity: a review of methods to deal with it and a simulation study evaluating their performance","volume":"36","author":"CF Dormann","year":"2013","journal-title":"Ecography"},{"issue":"3","key":"pcbi.1007787.ref039","first-page":"300","article-title":"A note on the use of principal components in regression","volume":"31","author":"IT Jolliffe","year":"1982","journal-title":"Journal of the Royal Statistical Society: Series C (Applied Statistics)"},{"issue":"919","key":"pcbi.1007787.ref040","doi-asserted-by":"crossref","first-page":"233","DOI":"10.1086\/282487","article-title":"Why mountain passes are higher in the tropics","volume":"101","author":"DH Janzen","year":"1967","journal-title":"The American Naturalist"},{"issue":"7","key":"pcbi.1007787.ref041","doi-asserted-by":"crossref","first-page":"1771","DOI":"10.1890\/03-9000","article-title":"Toward a metabolic theory of ecology","volume":"85","author":"JH Brown","year":"2004","journal-title":"Ecology"},{"issue":"1880","key":"pcbi.1007787.ref042","doi-asserted-by":"crossref","first-page":"20180767","DOI":"10.1098\/rspb.2018.0767","article-title":"Structural stability as a consistent predictor of phenological events","volume":"285","author":"C Song","year":"2018","journal-title":"Proceedings of the Royal Society B"},{"issue":"5","key":"pcbi.1007787.ref043","doi-asserted-by":"crossref","first-page":"553","DOI":"10.1086\/697046","article-title":"Fifty years of mountain passes: A perspective on Dan Janzen\u2019s classic article","volume":"191","author":"KS Sheldon","year":"2018","journal-title":"The American Naturalist"},{"issue":"10","key":"pcbi.1007787.ref044","doi-asserted-by":"crossref","first-page":"727","DOI":"10.1016\/j.tree.2017.07.001","article-title":"Mutualisms are not on the verge of breakdown","volume":"32","author":"ME Frederickson","year":"2017","journal-title":"Trends in Ecology & Evolution"},{"key":"pcbi.1007787.ref045","article-title":"Towards a probabilistic understanding about the context-dependency of species interactions","author":"C Song","year":"2020","journal-title":"Trends in Ecology & Evolution"},{"issue":"1-3","key":"pcbi.1007787.ref046","doi-asserted-by":"crossref","first-page":"159","DOI":"10.1016\/S0167-8809(00)00224-3","article-title":"Temperature variability and the yield of annual crops","volume":"82","author":"TR Wheeler","year":"2000","journal-title":"Agriculture, Ecosystems & Environment"},{"issue":"6972","key":"pcbi.1007787.ref047","doi-asserted-by":"crossref","first-page":"332","DOI":"10.1038\/nature02300","article-title":"The role of increasing temperature variability in European summer heatwaves","volume":"427","author":"C Sch\u00e4r","year":"2004","journal-title":"Nature"},{"issue":"5972","key":"pcbi.1007787.ref048","doi-asserted-by":"crossref","first-page":"1461","DOI":"10.1126\/science.1186473","article-title":"Phenology under global warming","volume":"327","author":"C K\u00f6rner","year":"2010","journal-title":"Science"},{"key":"pcbi.1007787.ref049","doi-asserted-by":"crossref","DOI":"10.1017\/CBO9780511803161","volume-title":"Causality","author":"J Pearl","year":"2009"},{"issue":"8","key":"pcbi.1007787.ref050","doi-asserted-by":"crossref","first-page":"e1002540","DOI":"10.1371\/journal.pbio.1002540","article-title":"Resource availability modulates the cooperative and competitive nature of a microbial cross-feeding mutualism","volume":"14","author":"TA Hoek","year":"2016","journal-title":"PLOS Biology"},{"key":"pcbi.1007787.ref051","first-page":"265","volume-title":"Advances In Ecological Research","author":"K Layer","year":"2010"},{"issue":"2","key":"pcbi.1007787.ref052","doi-asserted-by":"crossref","first-page":"295","DOI":"10.1111\/ecog.03443","article-title":"Food-web structure varies along environmental gradients in a high-latitude marine ecosystem","volume":"42","author":"S Kortsch","year":"2019","journal-title":"Ecography"},{"issue":"6","key":"pcbi.1007787.ref053","doi-asserted-by":"crossref","first-page":"1943","DOI":"10.2307\/1941447","article-title":"The problem of pattern and scale in ecology: the Robert H. MacArthur award lecture","volume":"73","author":"SA Levin","year":"1992","journal-title":"Ecology"},{"issue":"1","key":"pcbi.1007787.ref054","doi-asserted-by":"crossref","first-page":"63","DOI":"10.1007\/s10144-014-0471-0","article-title":"The stability\u2013complexity relationship at age 40: a random matrix perspective","volume":"57","author":"S Allesina","year":"2015","journal-title":"Population Ecology"},{"key":"pcbi.1007787.ref055","doi-asserted-by":"crossref","first-page":"115","DOI":"10.1016\/j.jtbi.2017.10.016","article-title":"Estimating the effect of the reorganization of interactions on the adaptability of species to changing environments","volume":"437","author":"S Cenci","year":"2018","journal-title":"Journal of theoretical biology"},{"key":"pcbi.1007787.ref056","doi-asserted-by":"crossref","first-page":"30","DOI":"10.1016\/j.jtbi.2018.04.030","article-title":"A guideline to study the feasibility domain of multi-trophic and changing ecological communities","volume":"450","author":"C Song","year":"2018","journal-title":"Journal of Theoretical Biology"},{"issue":"8","key":"pcbi.1007787.ref057","doi-asserted-by":"crossref","first-page":"1221","DOI":"10.1111\/ele.13091","article-title":"Structural changes within trophic levels are constrained by within-family assembly rules at lower trophic levels","volume":"21","author":"C Song","year":"2018","journal-title":"Ecology Letters"},{"key":"pcbi.1007787.ref058","doi-asserted-by":"crossref","first-page":"20140693","DOI":"10.1098\/rsif.2014.0693","article-title":"How structurally stable are global socioeconomic systems?","volume":"11","author":"S Saavedra","year":"2014","journal-title":"J R Soc Interface"},{"issue":"4","key":"pcbi.1007787.ref059","doi-asserted-by":"crossref","first-page":"411","DOI":"10.1086\/688046","article-title":"Persist or produce: a community trade-off tuned by species evenness","volume":"188","author":"RP Rohr","year":"2016","journal-title":"The American Naturalist"},{"issue":"4P2","key":"pcbi.1007787.ref060","doi-asserted-by":"crossref","first-page":"1","DOI":"10.1080\/00029890.1966.11970915","article-title":"Can one hear the shape of a drum?","volume":"73","author":"M Kac","year":"1966","journal-title":"The American Mathematical Monthly"},{"issue":"7463","key":"pcbi.1007787.ref061","doi-asserted-by":"crossref","first-page":"449","DOI":"10.1038\/nature12438","article-title":"Emergence of structural and dynamical properties of ecological mutualistic networks","volume":"500","author":"S Suweis","year":"2013","journal-title":"Nature"},{"issue":"6195","key":"pcbi.1007787.ref062","doi-asserted-by":"crossref","first-page":"1253497","DOI":"10.1126\/science.1253497","article-title":"On the structural stability of mutualistic systems","volume":"345","author":"RP Rohr","year":"2014","journal-title":"Science"},{"key":"pcbi.1007787.ref063","doi-asserted-by":"crossref","first-page":"997","DOI":"10.1002\/ece3.1930","article-title":"Nested species interactions promote feasibility over stability during the assembly of a pollinator community","volume":"6","author":"S Saavedra","year":"2016","journal-title":"Ecology & Evolution"},{"issue":"2","key":"pcbi.1007787.ref064","doi-asserted-by":"crossref","first-page":"238","DOI":"10.1111\/j.2517-6161.1951.tb00088.x","article-title":"The interpretation of interaction in contingency tables","volume":"13","author":"EH Simpson","year":"1951","journal-title":"Journal of the Royal Statistical Society: Series B (Methodological)"},{"issue":"2","key":"pcbi.1007787.ref065","doi-asserted-by":"crossref","first-page":"192","DOI":"10.1086\/381004","article-title":"On the generality of the latitudinal diversity gradient","volume":"163","author":"H Hillebrand","year":"2004","journal-title":"The American Naturalist"},{"issue":"1","key":"pcbi.1007787.ref066","doi-asserted-by":"crossref","first-page":"1","DOI":"10.1086\/689739","article-title":"Shaping the latitudinal diversity gradient: new perspectives from a synthesis of paleobiology and biogeography","volume":"189","author":"D Jablonski","year":"2017","journal-title":"The American Naturalist"},{"issue":"40","key":"pcbi.1007787.ref067","doi-asserted-by":"crossref","first-page":"14472","DOI":"10.1073\/pnas.1408471111","article-title":"Collapse of an ecological network in Ancient Egypt","volume":"111","author":"JD Yeakel","year":"2014","journal-title":"Proceedings of the National Academy of Sciences"},{"key":"pcbi.1007787.ref068","doi-asserted-by":"crossref","first-page":"80","DOI":"10.1016\/j.soilbio.2018.05.007","article-title":"Deforestation decreases spatial turnover and alters the network interactions in soil bacterial communities","volume":"123","author":"J Tian","year":"2018","journal-title":"Soil Biology and Biochemistry"},{"issue":"7241","key":"pcbi.1007787.ref069","doi-asserted-by":"crossref","first-page":"1018","DOI":"10.1038\/nature07950","article-title":"The architecture of mutualistic networks minimizes competition and increases biodiversity","volume":"458","author":"U Bastolla","year":"2009","journal-title":"Nature"},{"issue":"5941","key":"pcbi.1007787.ref070","doi-asserted-by":"crossref","first-page":"747","DOI":"10.1126\/science.1173536","article-title":"Generalized models reveal stabilizing factors in food webs","volume":"325","author":"T Gross","year":"2009","journal-title":"Science"},{"issue":"8","key":"pcbi.1007787.ref071","doi-asserted-by":"crossref","first-page":"797","DOI":"10.1111\/j.1461-0248.2011.01647.x","article-title":"An interaction switch predicts the nested architecture of mutualistic networks","volume":"14","author":"F Zhang","year":"2011","journal-title":"Ecology Letters"},{"issue":"1","key":"pcbi.1007787.ref072","doi-asserted-by":"crossref","first-page":"94","DOI":"10.1038\/s41559-017-0383-4","article-title":"The architecture of mutualistic networks as an evolutionary spandrel","volume":"2","author":"S Valverde","year":"2018","journal-title":"Nature Ecology & Evolution"},{"key":"pcbi.1007787.ref073","doi-asserted-by":"crossref","first-page":"1517","DOI":"10.1111\/ele.13279","article-title":"Mutualistic networks: moving closer to a predictive theory","volume":"22","author":"FS Valdovinos","year":"2019","journal-title":"Ecology Letters"},{"issue":"3","key":"pcbi.1007787.ref074","doi-asserted-by":"crossref","first-page":"324","DOI":"10.1111\/ele.12912","article-title":"Network spandrels reflect ecological assembly","volume":"21","author":"DS Maynard","year":"2018","journal-title":"Ecology Letters"},{"issue":"1","key":"pcbi.1007787.ref075","doi-asserted-by":"crossref","first-page":"18","DOI":"10.1111\/geb.12842","article-title":"Where Newton might have taken ecology","volume":"28","author":"DJ Currie","year":"2019","journal-title":"Global Ecology and Biogeography"},{"key":"pcbi.1007787.ref076","unstructured":"Oscar Perpi\u00f1\u00e1n and Robert Hijmans. rasterVis. 2019."}],"updated-by":[{"DOI":"10.1371\/journal.pcbi.1007787","type":"new_version","label":"New version","source":"publisher","updated":{"date-parts":[[2020,5,5]],"date-time":"2020-05-05T00:00:00Z","timestamp":1588636800000}}],"container-title":["PLOS Computational Biology"],"original-title":[],"language":"en","link":[{"URL":"https:\/\/dx.plos.org\/10.1371\/journal.pcbi.1007787","content-type":"unspecified","content-version":"vor","intended-application":"similarity-checking"}],"deposited":{"date-parts":[[2024,8,4]],"date-time":"2024-08-04T15:59:53Z","timestamp":1722787193000},"score":1,"resource":{"primary":{"URL":"https:\/\/dx.plos.org\/10.1371\/journal.pcbi.1007787"}},"subtitle":[],"editor":[{"given":"James","family":"O\u2019Dwyer","sequence":"first","affiliation":[]}],"short-title":[],"issued":{"date-parts":[[2020,4,23]]},"references-count":76,"journal-issue":{"issue":"4","published-online":{"date-parts":[[2020,4,23]]}},"URL":"https:\/\/doi.org\/10.1371\/journal.pcbi.1007787","relation":{"new_version":[{"id-type":"doi","id":"10.1371\/journal.pcbi.1007787","asserted-by":"object"}]},"ISSN":["1553-7358"],"issn-type":[{"value":"1553-7358","type":"electronic"}],"subject":[],"published":{"date-parts":[[2020,4,23]]}}}