{"status":"ok","message-type":"work","message-version":"1.0.0","message":{"indexed":{"date-parts":[[2026,4,15]],"date-time":"2026-04-15T01:24:08Z","timestamp":1776216248868,"version":"3.50.1"},"update-to":[{"DOI":"10.1371\/journal.pcbi.1007835","type":"new_version","label":"New version","source":"publisher","updated":{"date-parts":[[2020,5,20]],"date-time":"2020-05-20T00:00:00Z","timestamp":1589932800000}}],"reference-count":126,"publisher":"Public Library of Science (PLoS)","issue":"5","license":[{"start":{"date-parts":[[2020,5,8]],"date-time":"2020-05-08T00:00:00Z","timestamp":1588896000000},"content-version":"vor","delay-in-days":0,"URL":"http:\/\/creativecommons.org\/licenses\/by\/4.0\/"}],"funder":[{"DOI":"10.13039\/501100004189","name":"Max-Planck-Gesellschaft","doi-asserted-by":"publisher","award":["Independent Research Group"],"award-info":[{"award-number":["Independent Research Group"]}],"id":[{"id":"10.13039\/501100004189","id-type":"DOI","asserted-by":"publisher"}]}],"content-domain":{"domain":["www.ploscompbiol.org"],"crossmark-restriction":false},"short-container-title":["PLoS Comput Biol"],"DOI":"10.1371\/journal.pcbi.1007835","type":"journal-article","created":{"date-parts":[[2020,5,8]],"date-time":"2020-05-08T13:32:09Z","timestamp":1588944729000},"page":"e1007835","update-policy":"https:\/\/doi.org\/10.1371\/journal.pcbi.corrections_policy","source":"Crossref","is-referenced-by-count":44,"title":["Autonomous emergence of connectivity assemblies via spike triplet interactions"],"prefix":"10.1371","volume":"16","author":[{"given":"Lisandro","family":"Montangie","sequence":"first","affiliation":[]},{"ORCID":"https:\/\/orcid.org\/0000-0001-9094-2760","authenticated-orcid":true,"given":"Christoph","family":"Miehl","sequence":"additional","affiliation":[]},{"ORCID":"https:\/\/orcid.org\/0000-0001-7118-4079","authenticated-orcid":true,"given":"Julijana","family":"Gjorgjieva","sequence":"additional","affiliation":[]}],"member":"340","published-online":{"date-parts":[[2020,5,8]]},"reference":[{"key":"pcbi.1007835.ref001","volume-title":"The Organization of Behavior; a neuropsychological theory","author":"DO Hebb","year":"1949"},{"issue":"5041","key":"pcbi.1007835.ref002","doi-asserted-by":"crossref","first-page":"209","DOI":"10.1126\/science.1372754","article-title":"Selection of intrinsic horizontal connections in the visual cortex by correlated neuronal activity","volume":"255","author":"S L\u00f6wel","year":"1992","journal-title":"Science"},{"issue":"3","key":"pcbi.1007835.ref003","doi-asserted-by":"crossref","first-page":"60","DOI":"10.1038\/scientificamerican0992-60","article-title":"The Developing Brain","volume":"267","author":"CJ Shatz","year":"1992","journal-title":"Sci Am"},{"key":"pcbi.1007835.ref004","doi-asserted-by":"crossref","first-page":"461","DOI":"10.1016\/S0079-6123(08)60388-X","article-title":"Neuronal representations, assemblies and temporal coherence","volume":"95","author":"W Singer","year":"1993","journal-title":"Prog Brain Res"},{"issue":"8","key":"pcbi.1007835.ref005","doi-asserted-by":"crossref","first-page":"e1004458","DOI":"10.1371\/journal.pcbi.1004458","article-title":"Self-organization of microcircuits in networks of spiking neurons with plastic synapses","volume":"11","author":"GK Ocker","year":"2015","journal-title":"PLoS Comput Biol"},{"issue":"8","key":"pcbi.1007835.ref006","doi-asserted-by":"crossref","first-page":"e1005056","DOI":"10.1371\/journal.pcbi.1005056","article-title":"Shaping neural circuits by high order synaptic interactions","volume":"12","author":"N Ravid Tannenbaum","year":"2016","journal-title":"PLoS Comput Bio"},{"issue":"6300","key":"pcbi.1007835.ref007","doi-asserted-by":"crossref","first-page":"691","DOI":"10.1126\/science.aaf7560","article-title":"Imprinting and recalling cortical ensembles","volume":"353","author":"L Carrillo-Reid","year":"2016","journal-title":"Science"},{"key":"pcbi.1007835.ref008","doi-asserted-by":"crossref","first-page":"399","DOI":"10.1038\/nrn1669","article-title":"Neural signatures of cell assembly organization","volume":"6","author":"KD Harris","year":"2005","journal-title":"Nat Rev Neurosci"},{"key":"pcbi.1007835.ref009","doi-asserted-by":"crossref","first-page":"65","DOI":"10.1038\/nrn2303","article-title":"Synaptic plasticity, memory and the hippocampus: a neural network approach to causality","volume":"9","author":"G Neves","year":"2008","journal-title":"Nat Rev Neurosci"},{"issue":"3","key":"pcbi.1007835.ref010","doi-asserted-by":"crossref","first-page":"178","DOI":"10.1016\/j.tins.2008.12.002","article-title":"Associative memory models: from the cell-assembly theory to biophysically detailed cortex simulations","volume":"32","author":"A Lansner","year":"2009","journal-title":"Trends Neurosci"},{"issue":"3","key":"pcbi.1007835.ref011","doi-asserted-by":"crossref","first-page":"362","DOI":"10.1016\/j.neuron.2010.09.023","article-title":"Neural syntax: Cell assemblies, synapsembles, and readers","volume":"68","author":"G Buzs\u00e1ki","year":"2010","journal-title":"Neuron"},{"issue":"13","key":"pcbi.1007835.ref012","doi-asserted-by":"crossref","first-page":"5419","DOI":"10.1073\/pnas.1016051108","article-title":"A synaptic organizing principle for cortical neuronal groups","volume":"108","author":"R Perin","year":"2011","journal-title":"Proc Natl Acad Sci USA"},{"issue":"7345","key":"pcbi.1007835.ref013","doi-asserted-by":"crossref","first-page":"87","DOI":"10.1038\/nature09880","article-title":"Functional specificity of local synaptic connections in neocortical networks","volume":"473","author":"H Ko","year":"2011","journal-title":"Nature"},{"issue":"7443","key":"pcbi.1007835.ref014","doi-asserted-by":"crossref","first-page":"96","DOI":"10.1038\/nature12015","article-title":"The emergence of functional microcircuits in visual cortex","volume":"496","author":"H Ko","year":"2013","journal-title":"Nature"},{"issue":"12","key":"pcbi.1007835.ref015","doi-asserted-by":"crossref","first-page":"1553","DOI":"10.1038\/nn.4418","article-title":"Functional and structural underpinnings of neuronal assembly formation in learning","volume":"19","author":"A Holtmaat","year":"2016","journal-title":"Nat Neurosci"},{"issue":"7028","key":"pcbi.1007835.ref016","doi-asserted-by":"crossref","first-page":"868","DOI":"10.1038\/nature03252","article-title":"Excitatory cortical neurons form fine-scale functional networks","volume":"433","author":"Y Yoshimura","year":"2005","journal-title":"Nature"},{"issue":"12","key":"pcbi.1007835.ref017","doi-asserted-by":"crossref","first-page":"1472","DOI":"10.1038\/nn1798","article-title":"Cortical feed-forward networks for binding different streams of sensory information","volume":"9","author":"BM Kampa","year":"2006","journal-title":"Nat Neurosci"},{"issue":"2","key":"pcbi.1007835.ref018","doi-asserted-by":"crossref","first-page":"e1004759","DOI":"10.1371\/journal.pcbi.1004759","article-title":"Plasticity-driven self-organization under topological constraints accounts for non-random features of cortical synaptic wiring","volume":"12","author":"D Miner","year":"2016","journal-title":"PLoS Comput Bio"},{"issue":"7539","key":"pcbi.1007835.ref019","doi-asserted-by":"crossref","first-page":"399","DOI":"10.1038\/nature14182","article-title":"Functional organization of excitatory synaptic strength in primary visual cortex","volume":"518","author":"L Cossell","year":"2015","journal-title":"Nature"},{"issue":"7599","key":"pcbi.1007835.ref020","doi-asserted-by":"crossref","first-page":"370","DOI":"10.1038\/nature17192","article-title":"Anatomy and function of an excitatory network in the visual cortex","volume":"532","author":"WC Lee","year":"2016","journal-title":"Nature"},{"issue":"38","key":"pcbi.1007835.ref021","doi-asserted-by":"crossref","first-page":"E4053","DOI":"10.1073\/pnas.1406077111","article-title":"Visual stimuli recruit intrinsically generated cortical ensembles","volume":"111","author":"JK Miller","year":"2014","journal-title":"Proc Natl Acad Sci USA"},{"key":"pcbi.1007835.ref022","doi-asserted-by":"crossref","first-page":"2474","DOI":"10.1038\/ncomms3574","article-title":"Circuit reactivation dynamically regulates synaptic plasticity in neocortex","volume":"4","author":"PB Kruskal","year":"2013","journal-title":"Nat Commun"},{"key":"pcbi.1007835.ref023","doi-asserted-by":"crossref","first-page":"111","DOI":"10.3389\/fncir.2013.00111","article-title":"Direction selectivity in the visual system of the zebrafish larva","volume":"7","author":"C Gebhardt","year":"2013","journal-title":"Front Neural Circuits"},{"issue":"5","key":"pcbi.1007835.ref024","doi-asserted-by":"crossref","first-page":"1070","DOI":"10.1016\/j.neuron.2015.01.027","article-title":"Spontaneous neuronal network dynamics reveal circuit\u2019s functional adaptations for behavior","volume":"85","author":"SA Romano","year":"2015","journal-title":"Neuron"},{"issue":"16","key":"pcbi.1007835.ref025","doi-asserted-by":"crossref","first-page":"2407","DOI":"10.1016\/j.cub.2017.06.056","article-title":"Spontaneous activity in the zebrafish tectum reorganizes over development and is influenced by visual experience","volume":"27","author":"L Avitan","year":"2017","journal-title":"Curr Biol"},{"issue":"5","key":"pcbi.1007835.ref026","doi-asserted-by":"crossref","first-page":"939","DOI":"10.1016\/j.celrep.2017.04.015","article-title":"The emergence of the spatial structure of tectal spontaneous activity is independent of visual inputs","volume":"19","author":"T Pietri","year":"2017","journal-title":"Cell Rep"},{"key":"pcbi.1007835.ref027","doi-asserted-by":"crossref","first-page":"46","DOI":"10.3389\/fncir.2018.00046","article-title":"Principles of functional circuit connectivity: Insights from spontaneous activity in the zebrafish optic tectum","volume":"12","author":"E Marachlian","year":"2018","journal-title":"Front in Neural Circuits"},{"issue":"2","key":"pcbi.1007835.ref028","doi-asserted-by":"crossref","first-page":"177","DOI":"10.1007\/s10827-008-0135-1","article-title":"Correlations in spiking neuronal networks with distance dependent connections","volume":"27","author":"B Kriener","year":"2009","journal-title":"J Comput Neurosci"},{"issue":"5","key":"pcbi.1007835.ref029","doi-asserted-by":"crossref","first-page":"e1002059","DOI":"10.1371\/journal.pcbi.1002059","article-title":"How structure determines correlations in neuronal networks","volume":"7","author":"V Pernice","year":"2011","journal-title":"PLoS Comput Bio"},{"key":"pcbi.1007835.ref030","doi-asserted-by":"crossref","first-page":"28","DOI":"10.3389\/fncom.2011.00028","article-title":"Synchronization from second order network connectivity statistics","volume":"5","author":"L Zhao","year":"2011","journal-title":"Front Comput Neurosci"},{"key":"pcbi.1007835.ref031","doi-asserted-by":"crossref","first-page":"8","DOI":"10.3389\/fncom.2011.00008","article-title":"The role of degree distribution in shaping the dynamics in networks of sparsely connected spiking neurons","volume":"5","author":"A Roxin","year":"2011","journal-title":"Front Comput Neurosci"},{"key":"pcbi.1007835.ref032","doi-asserted-by":"crossref","first-page":"10","DOI":"10.3389\/fncom.2011.00010","article-title":"The Interaction of Intrinsic Dynamics and Network Topology in Determining Network Burst Synchrony","volume":"5","author":"C Gaiteri","year":"2011","journal-title":"Front Comput Neurosci"},{"issue":"11","key":"pcbi.1007835.ref033","doi-asserted-by":"crossref","first-page":"1498","DOI":"10.1038\/nn.3220","article-title":"Slow dynamics and high variability in balanced cortical networks with clustered connections","volume":"15","author":"A Litwin-Kumar","year":"2012","journal-title":"Nat Neurosci"},{"issue":"3","key":"pcbi.1007835.ref034","doi-asserted-by":"crossref","first-page":"e1002408","DOI":"10.1371\/journal.pcbi.1002408","article-title":"Impact of network structure and cellular response on spike time correlations","volume":"8","author":"J Trousdale","year":"2012","journal-title":"PLoS Comput Bio"},{"key":"pcbi.1007835.ref035","doi-asserted-by":"crossref","first-page":"72","DOI":"10.3389\/fncom.2013.00072","article-title":"The relevance of network micro-structure for neural dynamics","volume":"7","author":"V Pernice","year":"2013","journal-title":"Front Comput Neurosci"},{"issue":"1","key":"pcbi.1007835.ref036","doi-asserted-by":"crossref","first-page":"e1003428","DOI":"10.1371\/journal.pcbi.1003428","article-title":"The correlation structure of local neuronal networks intrinsically results from recurrent dynamics","volume":"10","author":"M Helias","year":"2014","journal-title":"PLoS Comput Bio"},{"issue":"6","key":"pcbi.1007835.ref037","doi-asserted-by":"crossref","first-page":"e1004963","DOI":"10.1371\/journal.pcbi.1004963","article-title":"Interplay between graph topology and correlations of third order in spiking neuronal networks","volume":"12","author":"S Jovanovi\u0107","year":"2016","journal-title":"PLoS Comput Biol"},{"key":"pcbi.1007835.ref038","doi-asserted-by":"crossref","first-page":"P03012","DOI":"10.1088\/1742-5468\/2013\/03\/P03012","article-title":"Motif statistics and spike correlations in neuronal networks","author":"Y Hu","year":"2013","journal-title":"J Stat Mech"},{"issue":"3","key":"pcbi.1007835.ref039","doi-asserted-by":"crossref","first-page":"032802","DOI":"10.1103\/PhysRevE.89.032802","article-title":"Local paths to global coherence: Cutting networks down to size","volume":"89","author":"Y Hu","year":"2014","journal-title":"Phys Rev E Stat Nonlin Soft Matter Phys"},{"key":"pcbi.1007835.ref040","doi-asserted-by":"crossref","first-page":"109","DOI":"10.1016\/j.conb.2017.07.011","article-title":"From the statistics of connectivity to the statistics of spike times in neuronal networks","volume":"46","author":"GK Ocker","year":"2017","journal-title":"Curr Opin Neurobiol"},{"issue":"5","key":"pcbi.1007835.ref041","doi-asserted-by":"crossref","first-page":"e1006153","DOI":"10.1371\/journal.pcbi.1006153","article-title":"Functional triplet motifs underlie accurate predictions of single-trial responses in populations of tuned and untuned V1 neurons","volume":"14","author":"JB Dechery","year":"2018","journal-title":"PLoS Comput Biol"},{"issue":"2","key":"pcbi.1007835.ref042","doi-asserted-by":"crossref","first-page":"81","DOI":"10.1007\/s00422-009-0319-4","article-title":"Emergence of network structure due to spike-timing-dependent plasticity in recurrent neuronal networks. I. Input selectivity\u2013strengthening correlated input pathways","volume":"101","author":"M Gilson","year":"2009","journal-title":"Biol Cybern"},{"issue":"2","key":"pcbi.1007835.ref043","doi-asserted-by":"crossref","first-page":"103","DOI":"10.1007\/s00422-009-0320-y","article-title":"Emergence of network structure due to spike-timing-dependent plasticity in recurrent neuronal networks. II. Input selectivity\u2014symmetry breaking","volume":"101","author":"M Gilson","year":"2009","journal-title":"Biol Cybern"},{"issue":"5-6","key":"pcbi.1007835.ref044","doi-asserted-by":"crossref","first-page":"427","DOI":"10.1007\/s00422-009-0346-1","article-title":"Emergence of network structure due to spike-timing-dependent plasticity in recurrent neuronal networks IV: structuring synaptic pathways among recurrent connections","volume":"101","author":"M Gilson","year":"2009","journal-title":"Biol Cybern"},{"issue":"5","key":"pcbi.1007835.ref045","doi-asserted-by":"crossref","first-page":"533","DOI":"10.1007\/s00422-007-0148-2","article-title":"Spike-timing-dependent plasticity for neurons with recurrent connections","volume":"96","author":"AN Burkitt","year":"2007","journal-title":"Biol Cybern"},{"issue":"7","key":"pcbi.1007835.ref046","doi-asserted-by":"crossref","first-page":"e101535","DOI":"10.1371\/journal.pone.0101535","article-title":"Interplay between Short- and Long-Term Plasticity in Cell-Assembly Formation","volume":"9","author":"N Hiratani","year":"2014","journal-title":"PLoS One"},{"issue":"1","key":"pcbi.1007835.ref047","doi-asserted-by":"crossref","first-page":"e84626","DOI":"10.1371\/journal.pone.0084626","article-title":"Emergence of Connectivity Motifs in Networks of Model Neurons with Short- and Long-Term Plastic Synapses","volume":"9","author":"E Vasilaki","year":"2014","journal-title":"PLoS One"},{"issue":"9","key":"pcbi.1007835.ref048","doi-asserted-by":"crossref","first-page":"e1006421","DOI":"10.1371\/journal.pcbi.1006421","article-title":"Emergence of spontaneous assembly activity in developing neural networks without afferent input","volume":"14","author":"MA Triplett","year":"2018","journal-title":"PLoS Comput Biol"},{"key":"pcbi.1007835.ref049","doi-asserted-by":"crossref","first-page":"5319","DOI":"10.1038\/ncomms6319","article-title":"Formation and maintenance of neuronal assemblies through synaptic plasticity","volume":"5","author":"A Litwin-Kumar","year":"2014","journal-title":"Nat Commun"},{"key":"pcbi.1007835.ref050","doi-asserted-by":"crossref","first-page":"6922","DOI":"10.1038\/ncomms7922","article-title":"Diverse synaptic plasticity mechanisms orchestrated to form and retrieve memories in spiking neural networks","volume":"6","author":"F Zenke","year":"2015","journal-title":"Nat Commun"},{"issue":"3","key":"pcbi.1007835.ref051","doi-asserted-by":"crossref","first-page":"937","DOI":"10.1093\/cercor\/bhy001","article-title":"Training and spontaneous reinforcement of neuronal assemblies by spike timing plasticity","volume":"29","author":"GK Ocker","year":"2019","journal-title":"Cereb Cortex"},{"issue":"4","key":"pcbi.1007835.ref052","doi-asserted-by":"crossref","first-page":"4498","DOI":"10.1103\/PhysRevE.59.4498","article-title":"Hebbian learning and spiking neurons","volume":"59","author":"R Kempter","year":"1999","journal-title":"Physical Review E"},{"key":"pcbi.1007835.ref053","doi-asserted-by":"crossref","first-page":"4","DOI":"10.3389\/fnsyn.2011.00004","article-title":"A history of spike-timing-dependent plasticity","volume":"3","author":"H Markram","year":"2011","journal-title":"Front Synaptic Neurosci"},{"issue":"38","key":"pcbi.1007835.ref054","doi-asserted-by":"crossref","first-page":"9673","DOI":"10.1523\/JNEUROSCI.1425-06.2006","article-title":"Triplets of spikes in a model of spike timing-dependent plasticity","volume":"26","author":"JP Pfister","year":"2006","journal-title":"J Neurosci"},{"issue":"6","key":"pcbi.1007835.ref055","doi-asserted-by":"crossref","first-page":"1149","DOI":"10.1016\/S0896-6273(01)00542-6","article-title":"Rate, timing, and cooperativity jointly determine cortical synaptic plasticity","volume":"32","author":"PJ Sj\u00f6str\u00f6m","year":"2001","journal-title":"Neuron"},{"issue":"2","key":"pcbi.1007835.ref056","doi-asserted-by":"crossref","first-page":"187","DOI":"10.1038\/nn1387","article-title":"Coactivation and timing-dependent integration of synaptic potentiation and depression","volume":"8","author":"HX Wang","year":"2005","journal-title":"Nat Neurosci"},{"issue":"48","key":"pcbi.1007835.ref057","doi-asserted-by":"crossref","first-page":"19383","DOI":"10.1073\/pnas.1105933108","article-title":"A triplet spike-timing-dependent plasticity model generalizes the Bienenstock-Cooper-Munro rule to higher-order spatiotemporal correlations","volume":"108","author":"J Gjorgjieva","year":"2011","journal-title":"Proc Natl Acad Sci USA"},{"issue":"7306","key":"pcbi.1007835.ref058","doi-asserted-by":"crossref","first-page":"617","DOI":"10.1038\/nature09178","article-title":"Sparse coding and high-order correlations in fine-scale cortical networks","volume":"466","author":"IE Ohiorhenuan","year":"2010","journal-title":"Nature"},{"issue":"1901","key":"pcbi.1007835.ref059","first-page":"3297","article-title":"The impact of high-order interactions on the rate of synchronous discharge and information transmission in somatosensory cortex","volume":"367","author":"F Montani","year":"2009","journal-title":"Philos Trans A Math Phys Eng Sci"},{"issue":"1","key":"pcbi.1007835.ref060","doi-asserted-by":"crossref","first-page":"125","DOI":"10.1007\/s10827-010-0257-0","article-title":"Information-geometric measure of 3-neuron firing patterns characterizes scale-dependence in cortical networks","volume":"30","author":"IE Ohiorhenuan","year":"2010","journal-title":"J Comput Neurosci"},{"issue":"48","key":"pcbi.1007835.ref061","doi-asserted-by":"crossref","first-page":"17514","DOI":"10.1523\/JNEUROSCI.3127-11.2011","article-title":"Higher-order interactions characterized in cortical activity","volume":"31","author":"S Yu","year":"2011","journal-title":"J Neuroscience"},{"issue":"1","key":"pcbi.1007835.ref062","doi-asserted-by":"crossref","first-page":"153","DOI":"10.1162\/089976600300015934","article-title":"The effects of pair-wise and higher order correlations on the firing rate of a post-synaptic neuron","volume":"12","author":"SM Bohte","year":"2000","journal-title":"Neural Comput"},{"issue":"6","key":"pcbi.1007835.ref063","doi-asserted-by":"crossref","first-page":"062707","DOI":"10.1103\/PhysRevE.92.062707","article-title":"Input nonlinearities can shape beyond-pairwise correlations and improve information transmission by neural populations","volume":"92","author":"J Zylberberg","year":"2015","journal-title":"Phys Rev E Stat Nonlin Soft Matter Phys"},{"key":"pcbi.1007835.ref064","doi-asserted-by":"crossref","first-page":"10","DOI":"10.3389\/fncom.2014.00010","article-title":"When do microcircuits produce beyond-pairwise correlations?","volume":"8","author":"AK Barreiro","year":"2014","journal-title":"Front Comput Neurosci"},{"key":"pcbi.1007835.ref065","first-page":"57","article-title":"Triplet correlations among similarly tuned cells impact population coding","volume":"9","author":"NA Cayco-Gajic","year":"2015","journal-title":"Front Comput Neurosci"},{"issue":"1","key":"pcbi.1007835.ref066","doi-asserted-by":"crossref","first-page":"127","DOI":"10.1162\/089976603321043720","article-title":"Synchronous firing and higher-order interactions in neuron pool","volume":"15","author":"S Amari","year":"2003","journal-title":"Neural Comput"},{"issue":"20","key":"pcbi.1007835.ref067","doi-asserted-by":"crossref","first-page":"208102","DOI":"10.1103\/PhysRevLett.106.208102","article-title":"Common input explains higher-order correlations and entropy in a simple model of neural population activity","volume":"106","author":"JH Macke","year":"2011","journal-title":"Phys Rev Lett"},{"key":"pcbi.1007835.ref068","doi-asserted-by":"crossref","first-page":"388","DOI":"10.1016\/j.physa.2014.11.046","article-title":"Quantifying higher-order correlations in a neuronal pool","volume":"421","author":"L Montangie","year":"2015","journal-title":"Physica A: Statistical Mechanics and its Applications"},{"issue":"6-1","key":"pcbi.1007835.ref069","doi-asserted-by":"crossref","first-page":"060302","DOI":"10.1103\/PhysRevE.97.060302","article-title":"Common inputs in subthreshold membrane potential: The role of quiescent states in neuronal activity","volume":"97","author":"L Montangie","year":"2018","journal-title":"Phys Rev E"},{"issue":"1","key":"pcbi.1007835.ref070","doi-asserted-by":"crossref","first-page":"83","DOI":"10.1093\/biomet\/58.1.83","article-title":"Spectra of some self-exciting and mutually exciting point processes","volume":"58","author":"AG Hawkes","year":"1971","journal-title":"Biometrika"},{"issue":"24","key":"pcbi.1007835.ref071","doi-asserted-by":"crossref","first-page":"10464","DOI":"10.1523\/JNEUROSCI.18-24-10464.1998","article-title":"Synaptic modifications in cultured hippocampal neurons: dependence on spike timing, synaptic strength, and postsynaptic cell type","volume":"18","author":"GQ Bi","year":"1998","journal-title":"J Neurosci"},{"key":"pcbi.1007835.ref072","doi-asserted-by":"crossref","first-page":"151","DOI":"10.3389\/fnsys.2015.00151","article-title":"Philosophy of the spike: Rate-based vs. spike-based theories of the brain","volume":"9","author":"R Brette","year":"2015","journal-title":"Front Syst Neurosci"},{"key":"pcbi.1007835.ref073","doi-asserted-by":"crossref","first-page":"44","DOI":"10.1016\/j.conb.2015.12.008","article-title":"Computational implications of biophysical diversity and multiple timescales in neurons and synapses for circuit performance","volume":"37","author":"J Gjorgjieva","year":"2016","journal-title":"Curr Opin Neurobiol"},{"issue":"1","key":"pcbi.1007835.ref074","doi-asserted-by":"crossref","first-page":"1","DOI":"10.1162\/089976603321043685","article-title":"Asynchronous states and the emergence of synchrony in large networks of interacting excitatory and inhibitory neurons","volume":"15","author":"D Hansel","year":"2003","journal-title":"Neural Comput"},{"key":"pcbi.1007835.ref075","doi-asserted-by":"crossref","first-page":"2","DOI":"10.3389\/neuro.10.002.2008","article-title":"On dynamics of integrate-and-fire neural networks with conductance based synapses","volume":"2","author":"B Cessac","year":"2008","journal-title":"Frontiers in Computational Neuroscience"},{"issue":"5965","key":"pcbi.1007835.ref076","doi-asserted-by":"crossref","first-page":"587","DOI":"10.1126\/science.1179850","article-title":"The asynchronous state in cortical circuits","volume":"327","author":"A Renart","year":"2010","journal-title":"Science"},{"issue":"6062","key":"pcbi.1007835.ref077","doi-asserted-by":"crossref","first-page":"1569","DOI":"10.1126\/science.1211095","article-title":"Inhibitory plasticity balances excitation and inhibition in sensory pathways and memory networks","volume":"334","author":"TP Vogels","year":"2011","journal-title":"Science"},{"issue":"1","key":"pcbi.1007835.ref078","doi-asserted-by":"crossref","first-page":"e1002334","DOI":"10.1371\/journal.pcbi.1002334","article-title":"Balancing feed-forward excitation and inhibition via Hebbian inhibitory synaptic plasticity","volume":"8","author":"Y Luz","year":"2012","journal-title":"PLoS Comput Biol"},{"issue":"44","key":"pcbi.1007835.ref079","doi-asserted-by":"crossref","first-page":"E9366","DOI":"10.1073\/pnas.1705841114","article-title":"Balanced excitation and inhibition are required for high-capacity, noise-robust neuronal selectivity","volume":"114","author":"R Rubin","year":"2017","journal-title":"Proc Natl Acad Sci USA"},{"issue":"4","key":"pcbi.1007835.ref080","doi-asserted-by":"crossref","first-page":"563","DOI":"10.1016\/j.neuron.2010.02.003","article-title":"Spike-time-dependent plasticity and heterosynaptic competition organize networks to produce long scale-free sequences of neural activity","volume":"65","author":"IR Fiete","year":"2010","journal-title":"Neuron"},{"key":"pcbi.1007835.ref081","doi-asserted-by":"crossref","first-page":"1","DOI":"10.1016\/j.neuron.2020.03.002","article-title":"Heterosynaptic plasticity determines the set-point for cortical excitatory-inhibitory balance","volume":"106","author":"R Field","year":"2020","journal-title":"Neuron"},{"issue":"7030","key":"pcbi.1007835.ref082","doi-asserted-by":"crossref","first-page":"221","DOI":"10.1038\/nature03366","article-title":"Spike-timing-dependent synaptic plasticity depends on dendritic location","volume":"434","author":"RC Froemke","year":"2005","journal-title":"Nature"},{"issue":"7383","key":"pcbi.1007835.ref083","doi-asserted-by":"crossref","first-page":"47","DOI":"10.1038\/nature10776","article-title":"Conditional modulation of spike-timing-dependent plasticity for olfactory learning","volume":"482","author":"S Cassenaer","year":"2012","journal-title":"Nature"},{"key":"pcbi.1007835.ref084","doi-asserted-by":"crossref","first-page":"85","DOI":"10.3389\/fncir.2015.00085","article-title":"Neuromodulated spike-timing-dependent plasticity, and theory of three-factor learning rules","volume":"9","author":"N Fr\u00e9maux","year":"2016","journal-title":"Front Neural Circuits"},{"key":"pcbi.1007835.ref085","doi-asserted-by":"crossref","first-page":"38","DOI":"10.3389\/fnsyn.2016.00038","article-title":"The role of neuromodulators in cortical plasticity. A computational perspective","volume":"8","author":"V Pedrosa","year":"2017","journal-title":"Front Synaptic Neurosci"},{"key":"pcbi.1007835.ref086","doi-asserted-by":"crossref","first-page":"49","DOI":"10.3389\/fncom.2018.00049","article-title":"Modulation of spike-timing dependent plasticity: Towards the inclusion of a third factor in computational models","volume":"12","author":"A Foncelle","year":"2018","journal-title":"Front Comput Neurosci"},{"issue":"6","key":"pcbi.1007835.ref087","doi-asserted-by":"crossref","first-page":"587","DOI":"10.1038\/nn1058","article-title":"Dopamine gates LTP induction in lateral amygdala by suppressing feedforward inhibition","volume":"6","author":"S Bissi\u00e8re","year":"2003","journal-title":"Nat Neurosci"},{"issue":"6","key":"pcbi.1007835.ref088","doi-asserted-by":"crossref","first-page":"919","DOI":"10.1016\/j.neuron.2007.08.013","article-title":"Neuromodulators control the polarity of spike-timing-dependent synaptic plasticity","volume":"55","author":"GH Seol","year":"2007","journal-title":"Neuron"},{"issue":"3","key":"pcbi.1007835.ref089","doi-asserted-by":"crossref","first-page":"344","DOI":"10.1038\/nn.2479","article-title":"Connectivity reflects coding: a model of voltage-based STDP with homeostasis","volume":"13","author":"C Clopath","year":"2010","journal-title":"Nat Neurosci"},{"issue":"13","key":"pcbi.1007835.ref090","doi-asserted-by":"crossref","first-page":"3722","DOI":"10.1523\/JNEUROSCI.2511-15.2016","article-title":"Homeostatic activity-dependent tuning of recurrent networks for robust propagation of activity","volume":"36","author":"J Gjorgjieva","year":"2016","journal-title":"J Neurosci"},{"issue":"2","key":"pcbi.1007835.ref091","doi-asserted-by":"crossref","first-page":"e1002906","DOI":"10.1371\/journal.pcbi.1002906","article-title":"Pairwise analysis can account for network structures arising from Spike-Timing Dependent Plasticity","volume":"9","author":"B Babadi","year":"2013","journal-title":"PLoS Comput Biol"},{"issue":"6684","key":"pcbi.1007835.ref092","doi-asserted-by":"crossref","first-page":"440","DOI":"10.1038\/30918","article-title":"Collective dynamics of \u2018small-world\u2019 networks","volume":"393","author":"DJ Watts","year":"1998","journal-title":"Nature"},{"issue":"23","key":"pcbi.1007835.ref093","doi-asserted-by":"crossref","first-page":"8577","DOI":"10.1073\/pnas.0601602103","article-title":"Modularity and community structure in networks","volume":"103","author":"MEJ Newman","year":"2006","journal-title":"Proc Natl Acad Sci USA"},{"issue":"12","key":"pcbi.1007835.ref094","doi-asserted-by":"crossref","first-page":"e1007360","DOI":"10.1371\/journal.pcbi.1007360","article-title":"Stability of spontaneous, correlated activity in mouse auditory cortex","volume":"15","author":"R Betzel","year":"2019","journal-title":"PLoS Comput Biol"},{"issue":"2","key":"pcbi.1007835.ref095","doi-asserted-by":"crossref","first-page":"026107","DOI":"10.1103\/PhysRevE.76.026107","article-title":"Clustering in complex directed networks","volume":"76","author":"G Fagiolo","year":"2007","journal-title":"Phys Rev E Stat Nonlin Soft Matter Phys"},{"issue":"19","key":"pcbi.1007835.ref096","doi-asserted-by":"crossref","first-page":"198701","DOI":"10.1103\/PhysRevLett.87.198701","article-title":"Efficient behavior of small-world networks","volume":"87","author":"V Latora","year":"2001","journal-title":"Phys Rev Lett"},{"issue":"1","key":"pcbi.1007835.ref097","doi-asserted-by":"crossref","first-page":"016110","DOI":"10.1103\/PhysRevE.74.016110","article-title":"Statistical mechanics of community detection","volume":"74","author":"J Reichardt","year":"2006","journal-title":"Phys Rev E Stat Nonlin Soft Matter Phys"},{"key":"pcbi.1007835.ref098","doi-asserted-by":"crossref","first-page":"P10008","DOI":"10.1088\/1742-5468\/2008\/10\/P10008","article-title":"Fast unfolding of communities in large networks","volume":"10","author":"VD Blondel","year":"2008","journal-title":"Journal of Statistical Mechanics: Theory and Experiment"},{"issue":"1","key":"pcbi.1007835.ref099","doi-asserted-by":"crossref","first-page":"45","DOI":"10.1016\/S0896-6273(00)00008-8","article-title":"Timing-Based LTP and LTD at Vertical Inputs to Layer II\/III Pyramidal Cells in Rat Barrel Cortex","volume":"27","author":"DE Feldmann","year":"2000","journal-title":"Neuron"},{"issue":"44","key":"pcbi.1007835.ref100","doi-asserted-by":"crossref","first-page":"11238","DOI":"10.1523\/JNEUROSCI.0104-16.2016","article-title":"Natural Firing Patterns Imply Low Sensitivity of Synaptic Plasticity to Spike Timing Compared with Firing Rate","volume":"36","author":"M Graupner","year":"2016","journal-title":"J Neurosci"},{"issue":"5-6","key":"pcbi.1007835.ref101","doi-asserted-by":"crossref","first-page":"411","DOI":"10.1007\/s00422-009-0343-4","article-title":"Emergence of network structure due to spike-timing-dependent plasticity in recurrent neuronal networks III: Partially connected neurons driven by spontaneous activity","volume":"101","author":"M Gilson","year":"2009","journal-title":"Biol Cybern"},{"issue":"4","key":"pcbi.1007835.ref102","doi-asserted-by":"crossref","first-page":"042802","DOI":"10.1103\/PhysRevE.91.042802","article-title":"Cumulants of Hawkes point processes","volume":"91","author":"S Jovanovi\u0107","year":"2015","journal-title":"Phys Rev E Stat Nonlin Soft Matter Phys"},{"issue":"1","key":"pcbi.1007835.ref103","doi-asserted-by":"crossref","first-page":"347","DOI":"10.1073\/pnas.0605643104","article-title":"Sequential structure of neocortical spontaneous activity in vivo","volume":"104","author":"A Luczak","year":"2007","journal-title":"Proc Natl Acad Sci USA"},{"issue":"7207","key":"pcbi.1007835.ref104","doi-asserted-by":"crossref","first-page":"995","DOI":"10.1038\/nature07140","article-title":"Spatio-temporal correlations and visual signalling in a complete neuronal population","volume":"454","author":"JW Pillow","year":"2008","journal-title":"Nature"},{"issue":"6583","key":"pcbi.1007835.ref105","doi-asserted-by":"crossref","first-page":"607","DOI":"10.1038\/381607a0","article-title":"Emergence of simple-cell receptive field properties by learning a sparse code for natural images","volume":"381","author":"BA Olshausen","year":"1996","journal-title":"Nature"},{"key":"pcbi.1007835.ref106","doi-asserted-by":"crossref","first-page":"1193","DOI":"10.1146\/annurev.neuro.24.1.1193","article-title":"Natural Image Statistics and Neural Representation","volume":"24","author":"EP Simoncelli","year":"2001","journal-title":"Annu Rev Neurosci"},{"issue":"6","key":"pcbi.1007835.ref107","doi-asserted-by":"crossref","first-page":"1129","DOI":"10.1162\/neco.1995.7.6.1129","article-title":"An information-maximization approach to blind separation and blind deconvolution","volume":"7","author":"AJ Bell","year":"1995","journal-title":"Neural Comp"},{"issue":"1","key":"pcbi.1007835.ref108","doi-asserted-by":"crossref","first-page":"3","DOI":"10.1016\/S0893-6080(05)80003-6","article-title":"Objective function formulation of the BCM theory of visual cortical plasticity: Statistical connections, stability conditions","volume":"5","author":"N Intrator","year":"1992","journal-title":"Neural Networks"},{"issue":"7","key":"pcbi.1007835.ref109","doi-asserted-by":"crossref","first-page":"1797","DOI":"10.1162\/089976698300017142","article-title":"Receptive field formation in natural scene environments. Comparison of single-cell learning rules","volume":"10","author":"BS Blais","year":"1998","journal-title":"Neural Comp"},{"issue":"10","key":"pcbi.1007835.ref110","doi-asserted-by":"crossref","first-page":"e350","DOI":"10.1371\/journal.pbio.0030350","article-title":"Highly nonrandom features of synaptic connectivity in local cortical circuits","volume":"3","author":"S Song","year":"2005","journal-title":"PLoS Biol"},{"issue":"6209","key":"pcbi.1007835.ref111","doi-asserted-by":"crossref","first-page":"651","DOI":"10.1126\/science.346.6209.651-c","article-title":"Connectomics at the cutting edge: Challenges and opportunities in high-resolution brain mapping","volume":"346","author":"J Lichtman","year":"2014","journal-title":"Science"},{"key":"pcbi.1007835.ref112","doi-asserted-by":"crossref","first-page":"197","DOI":"10.1146\/annurev-neuro-071714-033954","article-title":"From Cajal to Connectome and Beyond","volume":"39","author":"LW Swanson","year":"2016","journal-title":"Ann Rev Neurosci"},{"issue":"1","key":"pcbi.1007835.ref113","doi-asserted-by":"crossref","first-page":"18","DOI":"10.1038\/nrn2759","article-title":"Mechanisms underlying spontaneous patterned activity in developing neural circuits","volume":"11","author":"AG Blankenship","year":"2010","journal-title":"Nat Rev Neurosci"},{"issue":"7419","key":"pcbi.1007835.ref114","doi-asserted-by":"crossref","first-page":"219","DOI":"10.1038\/nature11529","article-title":"Retinal waves coordinate patterned activity throughout the developing visual system","volume":"490","author":"JB Ackman","year":"2012","journal-title":"Nature"},{"key":"pcbi.1007835.ref115","doi-asserted-by":"crossref","first-page":"39","DOI":"10.1016\/j.conb.2017.07.004","article-title":"Understanding neural circuit development through theory and models","volume":"46","author":"LM Richter","year":"2017","journal-title":"Curr Opin in Neurobiol"},{"issue":"35","key":"pcbi.1007835.ref116","doi-asserted-by":"crossref","first-page":"10890","DOI":"10.1523\/JNEUROSCI.2012-09.2009","article-title":"Internally Mediated Developmental Desynchronization of Neocortical Network Activity","volume":"29","author":"P Golshani","year":"2009","journal-title":"J Neurosci"},{"issue":"35","key":"pcbi.1007835.ref117","doi-asserted-by":"crossref","first-page":"15049","DOI":"10.1073\/pnas.0907660106","article-title":"Sparsification of neuronal activity in the visual cortex at eye-opening","volume":"106","author":"NL Rochefort","year":"2009","journal-title":"Proc Natl Acad Sci USA"},{"issue":"3","key":"pcbi.1007835.ref118","doi-asserted-by":"crossref","first-page":"e61","DOI":"10.1371\/journal.pbio.0050061","article-title":"A Burst-Based \u201cHebbian\u201d Learning Rule at Retinogeniculate Synapses Links Retinal Waves to Activity-Dependent Refinement","volume":"5","author":"DA Butts","year":"2007","journal-title":"PLoS Biol"},{"issue":"2","key":"pcbi.1007835.ref119","doi-asserted-by":"crossref","first-page":"399","DOI":"10.1016\/j.neuron.2015.06.029","article-title":"Spontaneous Activity Drives Local Synaptic Plasticity In Vivo","volume":"87","author":"J Winnubst","year":"2015","journal-title":"Neuron"},{"key":"pcbi.1007835.ref120","doi-asserted-by":"crossref","first-page":"13208","DOI":"10.1038\/ncomms13208","article-title":"Spontaneous emergence of fast attractor dynamics in a model of developing primary visual cortex","volume":"7","author":"T Miconi","year":"2016","journal-title":"Nat Commun"},{"issue":"6595","key":"pcbi.1007835.ref121","doi-asserted-by":"crossref","first-page":"76","DOI":"10.1038\/383076a0","article-title":"A neuronal learning rule for sub-millisecond temporal coding","volume":"383","author":"W Gerstner","year":"1996","journal-title":"Nature"},{"issue":"2","key":"pcbi.1007835.ref122","doi-asserted-by":"crossref","first-page":"385","DOI":"10.1162\/089976600300015844","article-title":"Modeling synaptic plasticity in conjunction with the timing of pre- and postsynaptic action potentials","volume":"12","author":"WM Kistler","year":"2000","journal-title":"Neural Comput"},{"issue":"9","key":"pcbi.1007835.ref123","doi-asserted-by":"crossref","first-page":"919","DOI":"10.1038\/78829","article-title":"Competitive Hebbian learning through spike-timing-dependent synaptic plasticity","volume":"3","author":"S Song","year":"2000","journal-title":"Nat Neurosci"},{"issue":"3","key":"pcbi.1007835.ref124","doi-asserted-by":"crossref","first-page":"1059","DOI":"10.1016\/j.neuroimage.2009.10.003","article-title":"Complex network measures of brain connectivity: Uses and interpretations","volume":"52","author":"M Rubinov","year":"2010","journal-title":"Neuroimage"},{"issue":"11","key":"pcbi.1007835.ref125","doi-asserted-by":"crossref","first-page":"118703","DOI":"10.1103\/PhysRevLett.100.118703","article-title":"Community structure in directed networks","volume":"100","author":"EA Leicht","year":"2008","journal-title":"Phys Rev Lett"},{"issue":"1","key":"pcbi.1007835.ref126","doi-asserted-by":"crossref","first-page":"012803","DOI":"10.1103\/PhysRevE.87.012803","article-title":"Spectra of random graphs with arbitrary expected degrees","volume":"87","author":"RR Nadakuditi","year":"2013","journal-title":"Phys Rev E Stat Nonlin Soft Matter Phys"}],"updated-by":[{"DOI":"10.1371\/journal.pcbi.1007835","type":"new_version","label":"New version","source":"publisher","updated":{"date-parts":[[2020,5,20]],"date-time":"2020-05-20T00:00:00Z","timestamp":1589932800000}}],"container-title":["PLOS Computational Biology"],"original-title":[],"language":"en","link":[{"URL":"https:\/\/dx.plos.org\/10.1371\/journal.pcbi.1007835","content-type":"unspecified","content-version":"vor","intended-application":"similarity-checking"}],"deposited":{"date-parts":[[2021,3,13]],"date-time":"2021-03-13T06:48:57Z","timestamp":1615618137000},"score":1,"resource":{"primary":{"URL":"https:\/\/dx.plos.org\/10.1371\/journal.pcbi.1007835"}},"subtitle":[],"editor":[{"given":"Brent","family":"Doiron","sequence":"first","affiliation":[]}],"short-title":[],"issued":{"date-parts":[[2020,5,8]]},"references-count":126,"journal-issue":{"issue":"5","published-online":{"date-parts":[[2020,5,8]]}},"URL":"https:\/\/doi.org\/10.1371\/journal.pcbi.1007835","relation":{"has-preprint":[{"id-type":"doi","id":"10.1101\/716001","asserted-by":"object"}]},"ISSN":["1553-7358"],"issn-type":[{"value":"1553-7358","type":"electronic"}],"subject":[],"published":{"date-parts":[[2020,5,8]]}}}