{"status":"ok","message-type":"work","message-version":"1.0.0","message":{"indexed":{"date-parts":[[2026,2,27]],"date-time":"2026-02-27T09:34:25Z","timestamp":1772184865393,"version":"3.50.1"},"update-to":[{"DOI":"10.1371\/journal.pcbi.1008102","type":"new_version","label":"New version","source":"publisher","updated":{"date-parts":[[2020,8,24]],"date-time":"2020-08-24T00:00:00Z","timestamp":1598227200000}}],"reference-count":56,"publisher":"Public Library of Science (PLoS)","issue":"7","license":[{"start":{"date-parts":[[2020,7,30]],"date-time":"2020-07-30T00:00:00Z","timestamp":1596067200000},"content-version":"vor","delay-in-days":0,"URL":"http:\/\/creativecommons.org\/licenses\/by\/4.0\/"}],"funder":[{"DOI":"10.13039\/100000893","name":"Simons Foundation","doi-asserted-by":"publisher","award":["376199"],"award-info":[{"award-number":["376199"]}],"id":[{"id":"10.13039\/100000893","id-type":"DOI","asserted-by":"publisher"}]},{"DOI":"10.13039\/100000913","name":"James S. McDonnell Foundation","doi-asserted-by":"publisher","award":["220020439"],"award-info":[{"award-number":["220020439"]}],"id":[{"id":"10.13039\/100000913","id-type":"DOI","asserted-by":"publisher"}]}],"content-domain":{"domain":["www.ploscompbiol.org"],"crossmark-restriction":false},"short-container-title":["PLoS Comput Biol"],"DOI":"10.1371\/journal.pcbi.1008102","type":"journal-article","created":{"date-parts":[[2020,7,30]],"date-time":"2020-07-30T19:00:53Z","timestamp":1596135653000},"page":"e1008102","update-policy":"https:\/\/doi.org\/10.1371\/journal.pcbi.corrections_policy","source":"Crossref","is-referenced-by-count":32,"title":["Emergent neutrality in consumer-resource dynamics"],"prefix":"10.1371","volume":"16","author":[{"ORCID":"https:\/\/orcid.org\/0000-0001-9687-6178","authenticated-orcid":true,"given":"Rafael","family":"D\u2019Andrea","sequence":"first","affiliation":[]},{"ORCID":"https:\/\/orcid.org\/0000-0002-1515-8420","authenticated-orcid":true,"given":"Theo","family":"Gibbs","sequence":"additional","affiliation":[]},{"ORCID":"https:\/\/orcid.org\/0000-0003-1180-8622","authenticated-orcid":true,"given":"James P.","family":"O\u2019Dwyer","sequence":"additional","affiliation":[]}],"member":"340","published-online":{"date-parts":[[2020,7,30]]},"reference":[{"issue":"3","key":"pcbi.1008102.ref001","doi-asserted-by":"crossref","first-page":"679","DOI":"10.1034\/j.1600-0706.2003.12617.x","article-title":"Strong and weak tests of macroecological theory","volume":"102","author":"BJ McGill","year":"2003","journal-title":"Oikos"},{"issue":"10","key":"pcbi.1008102.ref002","doi-asserted-by":"crossref","first-page":"995","DOI":"10.1111\/j.1461-0248.2007.01094.x","article-title":"Species abundance distributions: moving beyond single prediction theories to integration within an ecological framework","volume":"10","author":"BJ McGill","year":"2007","journal-title":"Ecology letters"},{"key":"pcbi.1008102.ref003","doi-asserted-by":"crossref","DOI":"10.1093\/acprof:oso\/9780199593415.001.0001","volume-title":"Maximum entropy and ecology: a theory of abundance, distribution, and energetics","author":"J Harte","year":"2011"},{"issue":"6952","key":"pcbi.1008102.ref004","doi-asserted-by":"crossref","first-page":"1006","DOI":"10.1038\/4241006a","article-title":"Tail of death and resurrection","volume":"424","author":"J Harte","year":"2003","journal-title":"Nature"},{"issue":"5","key":"pcbi.1008102.ref005","doi-asserted-by":"crossref","first-page":"793","DOI":"10.1111\/j.2006.0906-7590.04714.x","article-title":"Null versus neutral models: what\u2019s the difference?","volume":"29","author":"NJ Gotelli","year":"2006","journal-title":"Ecography"},{"issue":"8","key":"pcbi.1008102.ref006","doi-asserted-by":"crossref","first-page":"451","DOI":"10.1016\/j.tree.2006.03.019","article-title":"The merits of neutral theory","volume":"21","author":"D Alonso","year":"2006","journal-title":"Trends in Ecology & Evolution"},{"issue":"1","key":"pcbi.1008102.ref007","doi-asserted-by":"crossref","first-page":"8","DOI":"10.1016\/j.tree.2008.09.004","article-title":"Beyond neutral science","volume":"24","author":"JS Clark","year":"2009","journal-title":"Trends in ecology & evolution"},{"issue":"7","key":"pcbi.1008102.ref008","doi-asserted-by":"crossref","first-page":"340","DOI":"10.1016\/j.tree.2011.03.024","article-title":"The unified neutral theory of biodiversity and biogeography at age ten","volume":"26","author":"J Rosindell","year":"2011","journal-title":"Trends in ecology & evolution"},{"issue":"4","key":"pcbi.1008102.ref009","doi-asserted-by":"crossref","DOI":"10.1016\/j.tree.2012.02.001","article-title":"The coherence problem with the Unified Neutral Theory of Biodiversity","volume":"27","author":"JS Clark","year":"2012","journal-title":"Trends in Ecology & Evolution"},{"issue":"4","key":"pcbi.1008102.ref010","doi-asserted-by":"crossref","first-page":"204","DOI":"10.1016\/j.tree.2012.01.004","article-title":"The case for ecological neutral theory","volume":"27","author":"J Rosindell","year":"2012","journal-title":"Trends in Ecology & Evolution"},{"issue":"3","key":"pcbi.1008102.ref011","doi-asserted-by":"crossref","first-page":"327","DOI":"10.2307\/1942257","article-title":"Community Structure: A Neutral Model Analysis","volume":"46","author":"HAL Caswell","year":"1976","journal-title":"Ecological Monographs"},{"key":"pcbi.1008102.ref012","volume-title":"The Unified Neutral Theory of Biodiversity and Biogeography","author":"SP Hubbell","year":"2001"},{"issue":"3","key":"pcbi.1008102.ref013","doi-asserted-by":"crossref","first-page":"241","DOI":"10.1111\/j.1461-0248.2003.00566.x","article-title":"Neutral theory and community ecology","volume":"7","author":"J Chave","year":"2004","journal-title":"Ecology Letters"},{"key":"pcbi.1008102.ref014","doi-asserted-by":"crossref","first-page":"309","DOI":"10.1038\/nature03211","article-title":"Field parameterization and experimental test of the neutral theory of biodiversity","volume":"433","author":"JT Wootton","year":"2005","journal-title":"Nature"},{"issue":"6","key":"pcbi.1008102.ref015","doi-asserted-by":"crossref","first-page":"1411","DOI":"10.1890\/0012-9658(2006)87[1411:EEONT]2.0.CO;2","article-title":"Empirical evaluation of neutral theory","volume":"87","author":"BJ McGill","year":"2006","journal-title":"Ecology"},{"issue":"6197","key":"pcbi.1008102.ref016","doi-asserted-by":"crossref","first-page":"45","DOI":"10.1038\/nature06197","article-title":"Patterns of relative species abundance in rainforests and coral reefs","volume":"450","author":"I Volkov","year":"2007","journal-title":"Nature"},{"issue":"5539","key":"pcbi.1008102.ref017","doi-asserted-by":"crossref","first-page":"2413","DOI":"10.1126\/science.293.5539.2413","article-title":"Neutral macroecology","volume":"293","author":"G Bell","year":"2001","journal-title":"Science (New York, NY)"},{"issue":"6952","key":"pcbi.1008102.ref018","doi-asserted-by":"crossref","first-page":"1035","DOI":"10.1038\/nature01883","article-title":"Neutral theory and relative species abundance in ecology","volume":"424","author":"I Volkov","year":"2003","journal-title":"Nature"},{"issue":"6940","key":"pcbi.1008102.ref019","doi-asserted-by":"crossref","first-page":"635","DOI":"10.1038\/nature01632","article-title":"Stability of forest biodiversity","volume":"423","author":"JS Clark","year":"2003","journal-title":"Nature"},{"issue":"6976","key":"pcbi.1008102.ref020","doi-asserted-by":"crossref","first-page":"696","DOI":"10.1038\/427696a","article-title":"The Stability of Forest Biodiversity\u2014Reply","volume":"427","author":"I Volkov","year":"2004","journal-title":"Nature"},{"issue":"44","key":"pcbi.1008102.ref021","doi-asserted-by":"crossref","first-page":"15747","DOI":"10.1073\/pnas.0502648102","article-title":"Species lifetime distribution for simple models of ecologies","volume":"102","author":"S Pigolotti","year":"2005","journal-title":"Proceedings of the National Academy of Sciences of the United States of America"},{"issue":"1","key":"pcbi.1008102.ref022","doi-asserted-by":"crossref","first-page":"52","DOI":"10.1093\/jpe\/rtr035","article-title":"Large-scale spatial synchrony and the stability of forest biodiversity revisited","volume":"5","author":"AM Ostling","year":"2012","journal-title":"Journal of Plant Ecology"},{"issue":"40","key":"pcbi.1008102.ref023","doi-asserted-by":"crossref","first-page":"16039","DOI":"10.1073\/pnas.1314992110","article-title":"Why abundant tropical tree species are phylogenetically old","volume":"110","author":"S Wang","year":"2013","journal-title":"Proceedings of the National Academy of Sciences"},{"issue":"4","key":"pcbi.1008102.ref024","doi-asserted-by":"crossref","first-page":"439","DOI":"10.1086\/677930","article-title":"Niche versus Neutrality: A Dynamical Analysis","volume":"184","author":"M Kalyuzhny","year":"2014","journal-title":"The American Naturalist"},{"issue":"27","key":"pcbi.1008102.ref025","doi-asserted-by":"crossref","first-page":"201419341","DOI":"10.1073\/pnas.1419341112","article-title":"Backbones of evolutionary history test biodiversity theory for microbes","volume":"112","author":"JP O\u2019Dwyer","year":"2015","journal-title":"Proceedings of the National Academy of Sciences"},{"issue":"5","key":"pcbi.1008102.ref026","doi-asserted-by":"crossref","first-page":"1207","DOI":"10.1890\/15-0984.1","article-title":"Reproducing static and dynamic biodiversity patterns in tropical forests: the critical role of environmental variance","volume":"97","author":"T Fung","year":"2016","journal-title":"Ecology"},{"issue":"12","key":"pcbi.1008102.ref027","doi-asserted-by":"crossref","first-page":"3211","DOI":"10.1002\/ecy.2020","article-title":"Biodiversity maintenance may be lower under partial niche differentiation than under neutrality","volume":"98","author":"R D\u2019Andrea","year":"2017","journal-title":"Ecology"},{"issue":"1","key":"pcbi.1008102.ref028","doi-asserted-by":"crossref","first-page":"1","DOI":"10.1086\/324112","article-title":"Comparing classical community models: theoretical consequences for patterns of diversity","volume":"159","author":"J Chave","year":"2002","journal-title":"The American Naturalist"},{"issue":"36","key":"pcbi.1008102.ref029","doi-asserted-by":"crossref","first-page":"15821","DOI":"10.1073\/pnas.1009387107","article-title":"Niche and neutral models predict asymptotically equivalent species abundance distributions in high-diversity ecological communities","volume":"107","author":"RA Chisholm","year":"2010","journal-title":"Proceedings of the National Academy of Sciences of the United States of America"},{"issue":"3","key":"pcbi.1008102.ref030","doi-asserted-by":"crossref","first-page":"e1004134","DOI":"10.1371\/journal.pcbi.1004134","article-title":"When Can Species Abundance Data Reveal Non-neutrality?","volume":"11","author":"O Al Hammal","year":"2015","journal-title":"PLOS Computational Biology"},{"key":"pcbi.1008102.ref031","article-title":"Emergent niche structuring leads to increased differences from neutrality in species abundance distributions","author":"RC Rael","year":"2018","journal-title":"Ecology"},{"issue":"1","key":"pcbi.1008102.ref032","doi-asserted-by":"crossref","first-page":"150","DOI":"10.1016\/j.jtbi.2010.10.006","article-title":"A mathematical synthesis of niche and neutral theories in community ecology","volume":"269","author":"B Haegeman","year":"2011","journal-title":"Journal of theoretical biology"},{"issue":"36","key":"pcbi.1008102.ref033","doi-asserted-by":"crossref","first-page":"13111","DOI":"10.1073\/pnas.1405637111","article-title":"The transition between the niche and neutral regimes in ecology","volume":"111","author":"CK Fisher","year":"2014","journal-title":"Proceedings of the National Academy of Sciences"},{"issue":"January","key":"pcbi.1008102.ref034","first-page":"1","article-title":"Metabolic Trade-Offs Promote Diversity in a Model Ecosystem","volume":"118","author":"A Posfai","year":"2017","journal-title":"Physical Review Letters"},{"issue":"3","key":"pcbi.1008102.ref035","doi-asserted-by":"crossref","first-page":"522","DOI":"10.1016\/j.jtbi.2007.06.010","article-title":"The zero-sum assumption in neutral biodiversity theory","volume":"248","author":"RS Etienne","year":"2007","journal-title":"Journal of theoretical biology"},{"key":"pcbi.1008102.ref036","doi-asserted-by":"crossref","first-page":"56","DOI":"10.1007\/s00285-003-0195-9","article-title":"The evolution of resource use","volume":"47","author":"SJ Schreiber","year":"2003","journal-title":"Journal of Mathematical Biology"},{"issue":"6","key":"pcbi.1008102.ref037","doi-asserted-by":"crossref","first-page":"716","DOI":"10.1111\/j.1461-0248.2010.01463.x","article-title":"Protracted speciation revitalizes the neutral theory of biodiversity","volume":"13","author":"J Rosindell","year":"2010","journal-title":"Ecology Letters"},{"issue":"3","key":"pcbi.1008102.ref038","doi-asserted-by":"crossref","first-page":"470","DOI":"10.1002\/ecm.1263","article-title":"A structural approach for understanding multispecies coexistence","volume":"87","author":"S Saavedra","year":"2017","journal-title":"Ecological Monographs"},{"issue":"16","key":"pcbi.1008102.ref039","doi-asserted-by":"crossref","first-page":"6230","DOI":"10.1073\/pnas.0508024103","article-title":"Self-organized similarity, the evolutionary emergence of groups of similar species","volume":"103","author":"M Scheffer","year":"2006","journal-title":"Proceedings of the National Academy of Sciences of the United States of America"},{"issue":"3","key":"pcbi.1008102.ref040","doi-asserted-by":"crossref","first-page":"195","DOI":"10.1006\/tpbi.1998.1384","article-title":"Shaken not stirred: On permanence in ecological communities","volume":"54","author":"VAA Jansen","year":"1998","journal-title":"Theoretical Population Biology"},{"key":"pcbi.1008102.ref041","doi-asserted-by":"crossref","first-page":"317","DOI":"10.1016\/0040-5809(76)90051-4","article-title":"Coexistence of Species Competing for Shared Resources","volume":"9","author":"RA Armstrong","year":"1976","journal-title":"Theoretical Population Biology"},{"issue":"2","key":"pcbi.1008102.ref042","doi-asserted-by":"crossref","first-page":"151","DOI":"10.1086\/283553","article-title":"Competitive Exclusion","volume":"115","author":"RA Armstrong","year":"1980","journal-title":"The American Naturalist"},{"key":"pcbi.1008102.ref043","doi-asserted-by":"crossref","first-page":"1479","DOI":"10.1111\/ele.12350","article-title":"Sensitivity analysis of coexistence in ecological communities: theory and application","volume":"17","author":"G Barab\u00e1s","year":"2014","journal-title":"Ecology Letters"},{"issue":"3","key":"pcbi.1008102.ref044","doi-asserted-by":"crossref","first-page":"171","DOI":"10.1007\/s12080-009-0040-x","article-title":"The paradox of the clumps mathematically explained","volume":"2","author":"H Fort","year":"2009","journal-title":"Theoretical Ecology"},{"issue":"7068","key":"pcbi.1008102.ref045","doi-asserted-by":"crossref","first-page":"658","DOI":"10.1038\/nature04030","article-title":"Density dependence explains tree species abundance and diversity in tropical forests","volume":"438","author":"I Volkov","year":"2005","journal-title":"Nature"},{"issue":"6 Pt 1","key":"pcbi.1008102.ref046","doi-asserted-by":"crossref","first-page":"061902","DOI":"10.1103\/PhysRevE.68.061902","article-title":"Analytical solution of a neutral model of biodiversity","volume":"68","author":"M Vallade","year":"2003","journal-title":"Physical review E, Statistical, nonlinear, and soft matter physics"},{"issue":"3","key":"pcbi.1008102.ref047","doi-asserted-by":"crossref","first-page":"1","DOI":"10.1103\/RevModPhys.88.035003","article-title":"Statistical mechanics of ecological systems: Neutral theory and beyond","volume":"88","author":"S Azaele","year":"2016","journal-title":"Reviews of Modern Physics"},{"issue":"1","key":"pcbi.1008102.ref048","doi-asserted-by":"crossref","first-page":"42","DOI":"10.2307\/1411","article-title":"The Relation Between the Number of Species and the Number of Individuals in a Random Sample of an Animal Population","volume":"12","author":"RA Fisher","year":"1943","journal-title":"Journal of Animal Ecology"},{"issue":"10","key":"pcbi.1008102.ref049","doi-asserted-by":"crossref","first-page":"901","DOI":"10.1111\/j.1461-0248.2004.00640.x","article-title":"Sampling Hubbell\u2019s neutral theory of biodiversity","volume":"7","author":"D Alonso","year":"2004","journal-title":"Ecology Letters"},{"key":"pcbi.1008102.ref050","unstructured":"Arnold TB, Emerson JW. cvm.test: Discrete Cramer von Mises Goodness-of-Fit Tests; 2010. Available from: https:\/\/r-forge.r-project.org\/projects\/ks-test\/."},{"key":"pcbi.1008102.ref051","doi-asserted-by":"crossref","first-page":"85","DOI":"10.1016\/j.tpb.2014.02.002","article-title":"Species ages in neutral biodiversity models","volume":"93","author":"RA Chisholm","year":"2014","journal-title":"Theoretical Population Biology"},{"key":"pcbi.1008102.ref052","volume-title":"Handbook of mathematical functions: with formulas, graphs, and mathematical tables","author":"M Abramowitz","year":"1965"},{"issue":"3","key":"pcbi.1008102.ref053","doi-asserted-by":"crossref","first-page":"663","DOI":"10.1103\/RevModPhys.76.663","article-title":"Universality classes in nonequilibrium lattice systems","volume":"76","author":"G \u00d3dor","year":"2004","journal-title":"Reviews of modern physics"},{"key":"pcbi.1008102.ref054","unstructured":"Odor G. Phase transition universality classes of classical, nonequilibrium systems. arXiv preprint cond-mat\/0205644. 2002."},{"issue":"1-2","key":"pcbi.1008102.ref055","doi-asserted-by":"crossref","first-page":"1","DOI":"10.1023\/A:1018632005018","article-title":"Nonequilibrium phase transition in a model of diffusion, aggregation, and fragmentation","volume":"99","author":"SN Majumdar","year":"2000","journal-title":"Journal of Statistical Physics"},{"issue":"August","key":"pcbi.1008102.ref056","doi-asserted-by":"crossref","first-page":"469","DOI":"10.1126\/science.aat1168","article-title":"Emergent simplicity in microbial community assembly","volume":"361","author":"JE Goldford","year":"2018","journal-title":"Science"}],"updated-by":[{"DOI":"10.1371\/journal.pcbi.1008102","type":"new_version","label":"New version","source":"publisher","updated":{"date-parts":[[2020,8,24]],"date-time":"2020-08-24T00:00:00Z","timestamp":1598227200000}}],"container-title":["PLOS Computational Biology"],"original-title":[],"language":"en","link":[{"URL":"https:\/\/dx.plos.org\/10.1371\/journal.pcbi.1008102","content-type":"unspecified","content-version":"vor","intended-application":"similarity-checking"}],"deposited":{"date-parts":[[2020,8,24]],"date-time":"2020-08-24T13:37:34Z","timestamp":1598276254000},"score":1,"resource":{"primary":{"URL":"https:\/\/dx.plos.org\/10.1371\/journal.pcbi.1008102"}},"subtitle":[],"editor":[{"given":"Jacopo","family":"Grilli","sequence":"first","affiliation":[]}],"short-title":[],"issued":{"date-parts":[[2020,7,30]]},"references-count":56,"journal-issue":{"issue":"7","published-online":{"date-parts":[[2020,7,30]]}},"URL":"https:\/\/doi.org\/10.1371\/journal.pcbi.1008102","relation":{"has-preprint":[{"id-type":"doi","id":"10.1101\/710541","asserted-by":"object"}]},"ISSN":["1553-7358"],"issn-type":[{"value":"1553-7358","type":"electronic"}],"subject":[],"published":{"date-parts":[[2020,7,30]]}}}