{"status":"ok","message-type":"work","message-version":"1.0.0","message":{"indexed":{"date-parts":[[2026,5,28]],"date-time":"2026-05-28T01:24:15Z","timestamp":1779931455317,"version":"3.53.1"},"update-to":[{"DOI":"10.1371\/journal.pcbi.1013599","type":"new_version","label":"New version","source":"publisher","updated":{"date-parts":[[2025,11,4]],"date-time":"2025-11-04T00:00:00Z","timestamp":1762214400000}}],"reference-count":82,"publisher":"Public Library of Science (PLoS)","issue":"10","license":[{"start":{"date-parts":[[2025,10,27]],"date-time":"2025-10-27T00:00:00Z","timestamp":1761523200000},"content-version":"vor","delay-in-days":0,"URL":"http:\/\/creativecommons.org\/licenses\/by\/4.0\/"}],"funder":[{"DOI":"10.13039\/100000025","name":"National Institute of Mental Health","doi-asserted-by":"publisher","award":["R01-EY027401"],"award-info":[{"award-number":["R01-EY027401"]}],"id":[{"id":"10.13039\/100000025","id-type":"DOI","asserted-by":"publisher"}]},{"DOI":"10.13039\/100000025","name":"National Institute of Mental Health","doi-asserted-by":"publisher","award":["P30EY013079"],"award-info":[{"award-number":["P30EY013079"]}],"id":[{"id":"10.13039\/100000025","id-type":"DOI","asserted-by":"publisher"}]},{"name":"NYU Center for Brain Imaging"},{"name":"NYUAD Center for Brain and Health"}],"content-domain":{"domain":["www.ploscompbiol.org"],"crossmark-restriction":false},"short-container-title":["PLoS Comput Biol"],"abstract":"<jats:p>Primary visual cortex (V1) has long served as a model system for understanding cortical organization. Although its structural and functional properties vary markedly across its surface, patterns of covariation suggest possible underlying constancies. Such constancies would imply that V1 is composed of multiple identical units whose visual properties differ only due to differences in their inputs. To test this, we used fMRI to investigate how V1 cortical magnification and preferred spatial frequency covary with eccentricity and polar angle, measured in 40 observers. V1 cortical magnification and preferred spatial frequency were strongly correlated across eccentricity and around polar angle, however their relation differed between these dimensions: they were proportional across eccentricity but not polar angle. The constant ratio of cortical magnification to preferred spatial frequency when measured as a function of eccentricity suggests a shared underlying cause of variation in the two properties, e.g., the gradient of retinal ganglion cell density across eccentricity. In contrast, the deviation from proportionality around polar angle implies that cortical variation differs from that in retina along this dimension. Thus, a constancy hypothesis is supported for one of the two spatial dimensions of V1, highlighting the importance of examining the full 2D-map to understand how V1 is organized.<\/jats:p>","DOI":"10.1371\/journal.pcbi.1013599","type":"journal-article","created":{"date-parts":[[2025,10,27]],"date-time":"2025-10-27T17:46:37Z","timestamp":1761587197000},"page":"e1013599","update-policy":"https:\/\/doi.org\/10.1371\/journal.pcbi.corrections_policy","source":"Crossref","is-referenced-by-count":6,"title":["Unpacking the V1 map: Differential covariation of preferred spatial frequency and cortical magnification across spatial dimensions"],"prefix":"10.1371","volume":"21","author":[{"ORCID":"https:\/\/orcid.org\/0000-0001-9133-7984","authenticated-orcid":true,"given":"Marc M.","family":"Himmelberg","sequence":"first","affiliation":[],"role":[{"vocabulary":"crossref","role":"author"}]},{"given":"Yuna","family":"Kwak","sequence":"additional","affiliation":[],"role":[{"vocabulary":"crossref","role":"author"}]},{"given":"Marisa","family":"Carrasco","sequence":"additional","affiliation":[],"role":[{"vocabulary":"crossref","role":"author"}]},{"given":"Jonathan","family":"Winawer","sequence":"additional","affiliation":[],"role":[{"vocabulary":"crossref","role":"author"}]}],"member":"340","published-online":{"date-parts":[[2025,10,27]]},"reference":[{"issue":"4","key":"pcbi.1013599.ref001","doi-asserted-by":"crossref","first-page":"640","DOI":"10.1016\/j.neuron.2018.07.002","article-title":"Parcellating cerebral cortex: How invasive animal studies inform noninvasive mapmaking in humans","volume":"99","author":"DC Van Essen","year":"2018","journal-title":"Neuron"},{"key":"pcbi.1013599.ref002","volume-title":"Die sehstorungen bei schussverletzungen der kortikalen sehsphare: Nach beobachtungen an verwundeten der letszten japanischen kriege","author":"T Inouye","year":"1909"},{"issue":"11","key":"pcbi.1013599.ref003","doi-asserted-by":"crossref","first-page":"1255","DOI":"10.1016\/S0002-9394(41)91363-6","article-title":"Physiological studies on neural mechanisms of visual localization and discrimination","volume":"24","author":"SA Talbot","year":"1941","journal-title":"Am J Ophthalmol"},{"issue":"2","key":"pcbi.1013599.ref004","doi-asserted-by":"crossref","first-page":"203","DOI":"10.1113\/jphysiol.1961.sp006803","article-title":"The representation of the visual field on the cerebral cortex in monkeys","volume":"159","author":"PM Daniel","year":"1961","journal-title":"J Physiol"},{"issue":"3","key":"pcbi.1013599.ref005","doi-asserted-by":"crossref","first-page":"295","DOI":"10.1002\/cne.901580305","article-title":"Uniformity of monkey striate cortex: a parallel relationship between field size, scatter, and magnification factor","volume":"158","author":"DH Hubel","year":"1974","journal-title":"J Comp Neurol"},{"issue":"5","key":"pcbi.1013599.ref006","doi-asserted-by":"crossref","first-page":"545","DOI":"10.1016\/0042-6989(82)90113-4","article-title":"Spatial frequency selectivity of cells in macaque visual cortex","volume":"22","author":"RL De Valois","year":"1982","journal-title":"Vision Res"},{"issue":"6","key":"pcbi.1013599.ref007","doi-asserted-by":"crossref","first-page":"816","DOI":"10.1001\/archopht.1991.01080060080030","article-title":"The representation of the visual field in human striate cortex. 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