{"status":"ok","message-type":"work","message-version":"1.0.0","message":{"indexed":{"date-parts":[[2026,9,29]],"date-time":"2026-09-29T18:15:04Z","timestamp":1790705704681,"version":"4.1.0"},"update-to":[{"DOI":"10.1371\/journal.pbio.2006926","type":"new_version","label":"New version","source":"publisher","updated":{"date-parts":[[2019,1,16]],"date-time":"2019-01-16T00:00:00Z","timestamp":1547596800000}}],"reference-count":108,"publisher":"Public Library of Science (PLoS)","issue":"1","license":[{"start":{"date-parts":[[2019,1,4]],"date-time":"2019-01-04T00:00:00Z","timestamp":1546560000000},"content-version":"vor","delay-in-days":0,"URL":"http:\/\/creativecommons.org\/licenses\/by\/4.0\/"}],"content-domain":{"domain":["www.plosbiology.org"],"crossmark-restriction":false},"short-container-title":["PLoS Biol"],"DOI":"10.1371\/journal.pbio.2006926","type":"journal-article","created":{"date-parts":[[2019,1,4]],"date-time":"2019-01-04T18:39:01Z","timestamp":1546627141000},"page":"e2006926","update-policy":"https:\/\/doi.org\/10.1371\/journal.pbio.corrections_policy","source":"Crossref","is-referenced-by-count":86,"title":["Differential and convergent utilization of autophagy components by positive-strand RNA viruses"],"prefix":"10.1371","volume":"17","author":[{"given":"Emma","family":"Abernathy","sequence":"first","affiliation":[],"role":[{"vocabulary":"crossref","role":"author"}]},{"given":"Roberto","family":"Mateo","sequence":"additional","affiliation":[],"role":[{"vocabulary":"crossref","role":"author"}]},{"given":"Karim","family":"Majzoub","sequence":"additional","affiliation":[],"role":[{"vocabulary":"crossref","role":"author"}]},{"given":"Nick","family":"van Buuren","sequence":"additional","affiliation":[],"role":[{"vocabulary":"crossref","role":"author"}]},{"given":"Sara W.","family":"Bird","sequence":"additional","affiliation":[],"role":[{"vocabulary":"crossref","role":"author"}]},{"given":"Jan E.","family":"Carette","sequence":"additional","affiliation":[],"role":[{"vocabulary":"crossref","role":"author"}]},{"ORCID":"https:\/\/orcid.org\/0000-0001-7628-3770","authenticated-orcid":true,"given":"Karla","family":"Kirkegaard","sequence":"additional","affiliation":[],"role":[{"vocabulary":"crossref","role":"author"}]}],"member":"340","published-online":{"date-parts":[[2019,1,4]]},"reference":[{"key":"ref1","doi-asserted-by":"crossref","first-page":"107","DOI":"10.1146\/annurev-cellbio-092910-154005","article-title":"The role of Atg proteins in autophagosome formation","volume":"27","author":"N Mizushima","year":"2011","journal-title":"Annu Rev Cell Dev Biol"},{"issue":"9","key":"ref2","doi-asserted-by":"crossref","first-page":"831","DOI":"10.1038\/ncb0910-831","article-title":"The origin of the autophagosomal membrane","volume":"12","author":"SA Tooze","year":"2010","journal-title":"Nat Cell Biol"},{"issue":"4","key":"ref3","doi-asserted-by":"crossref","first-page":"656","DOI":"10.1016\/j.cell.2010.04.009","article-title":"Mitochondria supply membranes for autophagosome biogenesis during starvation","volume":"141","author":"DW Hailey","year":"2010","journal-title":"Cell"},{"issue":"8","key":"ref4","doi-asserted-by":"crossref","first-page":"747","DOI":"10.1038\/ncb2078","article-title":"Plasma membrane contributes to the formation of pre-autophagosomal structures","volume":"12","author":"B Ravikumar","year":"2010","journal-title":"Nat Cell Biol"},{"key":"ref5","doi-asserted-by":"crossref","first-page":"e04135","DOI":"10.7554\/eLife.04135","article-title":"Phosphatidylinositol 3-kinase and COPII generate LC3 lipidation vesicles from the ER-Golgi intermediate compartment","volume":"3","author":"L Ge","year":"2014","journal-title":"Elife"},{"issue":"6","key":"ref6","doi-asserted-by":"crossref","first-page":"1285","DOI":"10.1016\/j.cell.2013.08.044","article-title":"Diverse autophagosome membrane sources coalesce in recycling endosomes","volume":"154","author":"C Puri","year":"2013","journal-title":"Cell"},{"issue":"18","key":"ref7","doi-asserted-by":"crossref","first-page":"12297","DOI":"10.1074\/jbc.M900573200","article-title":"ULK1.ATG13.FIP200 complex mediates mTOR signaling and is essential for autophagy","volume":"284","author":"IG Ganley","year":"2009","journal-title":"J Biol Chem"},{"issue":"2","key":"ref8","doi-asserted-by":"crossref","first-page":"132","DOI":"10.1038\/ncb2152","article-title":"AMPK and mTOR regulate autophagy through direct phosphorylation of Ulk1","volume":"13","author":"J Kim","year":"2011","journal-title":"Nat Cell Biol"},{"issue":"6","key":"ref9","doi-asserted-by":"crossref","first-page":"585","DOI":"10.1042\/EBC20170021","article-title":"The mammalian ULK1 complex and autophagy initiation","volume":"61","author":"M Zachari","year":"2017","journal-title":"Essays Biochem"},{"issue":"7","key":"ref10","doi-asserted-by":"crossref","first-page":"741","DOI":"10.1038\/ncb2757","article-title":"ULK1 induces autophagy by phosphorylating Beclin-1 and activating VPS34 lipid kinase","volume":"15","author":"RC Russell","year":"2013","journal-title":"Nat Cell Biol"},{"issue":"6","key":"ref11","doi-asserted-by":"crossref","first-page":"773","DOI":"10.1083\/jcb.200907014","article-title":"Coordination of membrane events during autophagy by multiple class III PI3-kinase complexes","volume":"186","author":"A Simonsen","year":"2009","journal-title":"J Cell Biol"},{"issue":"7","key":"ref12","doi-asserted-by":"crossref","first-page":"671","DOI":"10.1016\/j.semcdb.2010.04.002","article-title":"Regulation of membrane biogenesis in autophagy via PI3P dynamics","volume":"21","author":"T Noda","year":"2010","journal-title":"Semin Cell Dev Biol"},{"issue":"10","key":"ref13","doi-asserted-by":"crossref","first-page":"1860","DOI":"10.1091\/mbc.e11-09-0746","article-title":"Dynamic and transient interactions of Atg9 with autophagosomes, but not membrane integration, are required for autophagy","volume":"23","author":"A Orsi","year":"2012","journal-title":"Mol Biol Cell"},{"issue":"6","key":"ref14","doi-asserted-by":"crossref","first-page":"1005","DOI":"10.1083\/jcb.200912089","article-title":"An Atg9-containing compartment that functions in the early steps of autophagosome biogenesis","volume":"190","author":"M Mari","year":"2010","journal-title":"J Cell Biol"},{"issue":"2","key":"ref15","doi-asserted-by":"crossref","first-page":"219","DOI":"10.1083\/jcb.201202061","article-title":"Atg9 vesicles are an important membrane source during early steps of autophagosome formation","volume":"198","author":"H Yamamoto","year":"2012","journal-title":"J Cell Biol"},{"issue":"Pt 18","key":"ref16","doi-asserted-by":"crossref","first-page":"3888","DOI":"10.1242\/jcs.03172","article-title":"Starvation and ULK1-dependent cycling of mammalian Atg9 between the TGN and endosomes","volume":"119","author":"AR Young","year":"2006","journal-title":"J Cell Sci"},{"issue":"4","key":"ref17","doi-asserted-by":"crossref","DOI":"10.15252\/embr.201744837","article-title":"SNX18 regulates ATG9A trafficking from recycling endosomes by recruiting Dynamin-2","volume":"19","author":"K Soreng","year":"2018","journal-title":"EMBO Rep"},{"issue":"2","key":"ref18","doi-asserted-by":"crossref","first-page":"231","DOI":"10.1016\/j.semcdb.2003.12.004","article-title":"Two ubiquitin-like conjugation systems essential for autophagy","volume":"15","author":"Y Ohsumi","year":"2004","journal-title":"Semin Cell Dev Biol"},{"issue":"8","key":"ref19","doi-asserted-by":"crossref","first-page":"3290","DOI":"10.1091\/mbc.e07-12-1292","article-title":"Atg8 controls phagophore expansion during autophagosome formation","volume":"19","author":"Z Xie","year":"2008","journal-title":"Mol Biol Cell"},{"issue":"21","key":"ref20","doi-asserted-by":"crossref","first-page":"5720","DOI":"10.1093\/emboj\/19.21.5720","article-title":"LC3, a mammalian homologue of yeast Apg8p, is localized in autophagosome membranes after processing","volume":"19","author":"Y Kabeya","year":"2000","journal-title":"EMBO J"},{"issue":"2","key":"ref21","doi-asserted-by":"crossref","first-page":"167","DOI":"10.1016\/j.molcel.2013.12.014","article-title":"Interactions between autophagy receptors and ubiquitin-like proteins form the molecular basis for selective autophagy","volume":"53","author":"V Rogov","year":"2014","journal-title":"Mol Cell"},{"issue":"4","key":"ref22","doi-asserted-by":"crossref","first-page":"457","DOI":"10.1016\/j.jmb.2017.01.006","article-title":"Sensing Membrane Curvature in Macroautophagy","volume":"429","author":"N Nguyen","year":"2017","journal-title":"J Mol Biol"},{"issue":"2","key":"ref23","doi-asserted-by":"crossref","first-page":"144","DOI":"10.1038\/nsmb.2475","article-title":"Interaction between FIP200 and ATG16L1 distinguishes ULK1 complex-dependent and -independent autophagy","volume":"20","author":"N Gammoh","year":"2013","journal-title":"Nat Struct Mol Biol"},{"issue":"10","key":"ref24","doi-asserted-by":"crossref","first-page":"1115","DOI":"10.4161\/auto.7.10.16608","article-title":"Beclin 1-independent autophagy contributes to apoptosis in cortical neurons","volume":"7","author":"Y Grishchuk","year":"2011","journal-title":"Autophagy"},{"issue":"7264","key":"ref25","doi-asserted-by":"crossref","first-page":"654","DOI":"10.1038\/nature08455","article-title":"Discovery of Atg5\/Atg7-independent alternative macroautophagy","volume":"461","author":"Y Nishida","year":"2009","journal-title":"Nature"},{"key":"ref26","doi-asserted-by":"crossref","first-page":"106","DOI":"10.1016\/j.ceb.2015.04.016","article-title":"Secretory autophagy","volume":"35","author":"M Ponpuak","year":"2015","journal-title":"Curr Opin Cell Biol"},{"issue":"6","key":"ref27","doi-asserted-by":"crossref","first-page":"359","DOI":"10.1002\/rmv.630","article-title":"Viruses and autophagy","volume":"19","author":"SB Kudchodkar","year":"2009","journal-title":"Rev Med Virol"},{"issue":"3","key":"ref28","doi-asserted-by":"crossref","first-page":"268","DOI":"10.4161\/auto.5210","article-title":"Downregulation of autophagy by herpesvirus Bcl-2 homologs","volume":"4","author":"C Liang","year":"2008","journal-title":"Autophagy"},{"issue":"2","key":"ref29","doi-asserted-by":"crossref","first-page":"115","DOI":"10.1016\/j.chom.2010.01.007","article-title":"Autophagy protects against Sindbis virus infection of the central nervous system","volume":"7","author":"A Orvedahl","year":"2010","journal-title":"Cell Host Microbe"},{"issue":"5817","key":"ref30","doi-asserted-by":"crossref","first-page":"1398","DOI":"10.1126\/science.1136880","article-title":"Autophagy-dependent viral recognition by plasmacytoid dendritic cells","volume":"315","author":"HK Lee","year":"2007","journal-title":"Science"},{"issue":"5","key":"ref31","doi-asserted-by":"crossref","first-page":"363","DOI":"10.1038\/nrmicro1890","article-title":"Modification of intracellular membrane structures for virus replication","volume":"6","author":"S Miller","year":"2008","journal-title":"Nat Rev Microbiol"},{"key":"ref32","first-page":"139","article-title":"Viral RNA replication in association with cellular membranes","volume":"285","author":"A Salonen","year":"2005","journal-title":"Curr Top Microbiol Immunol"},{"issue":"31","key":"ref33","doi-asserted-by":"crossref","first-page":"11263","DOI":"10.1073\/pnas.0404157101","article-title":"Alternate, virus-induced membrane rearrangements support positive-strand RNA virus genome replication","volume":"101","author":"M Schwartz","year":"2004","journal-title":"Proc Natl Acad Sci U S A"},{"issue":"10","key":"ref34","doi-asserted-by":"crossref","first-page":"6576","DOI":"10.1128\/JVI.70.10.6576-6588.1996","article-title":"Cellular origin and ultrastructure of membranes induced during poliovirus infection","volume":"70","author":"A Schlegel","year":"1996","journal-title":"J Virol"},{"issue":"19","key":"ref35","doi-asserted-by":"crossref","first-page":"8953","DOI":"10.1128\/JVI.74.19.8953-8965.2000","article-title":"Remodeling the endoplasmic reticulum by poliovirus infection and by individual viral proteins: an autophagy-like origin for virus-induced vesicles","volume":"74","author":"DA Suhy","year":"2000","journal-title":"J Virol"},{"issue":"5","key":"ref36","doi-asserted-by":"crossref","first-page":"e156","DOI":"10.1371\/journal.pbio.0030156","article-title":"Subversion of cellular autophagosomal machinery by RNA viruses","volume":"3","author":"WT Jackson","year":"2005","journal-title":"PLoS Biol"},{"issue":"18","key":"ref37","doi-asserted-by":"crossref","first-page":"9143","DOI":"10.1128\/JVI.00641-08","article-title":"Autophagosome supports coxsackievirus B3 replication in host cells","volume":"82","author":"J Wong","year":"2008","journal-title":"J Virol"},{"issue":"11","key":"ref38","doi-asserted-by":"crossref","first-page":"e1003046","DOI":"10.1371\/journal.ppat.1003046","article-title":"Intracellular vesicle acidification promotes maturation of infectious poliovirus particles","volume":"8","author":"AL Richards","year":"2012","journal-title":"PLoS Pathog"},{"issue":"36","key":"ref39","doi-asserted-by":"crossref","first-page":"13081","DOI":"10.1073\/pnas.1401437111","article-title":"Nonlytic viral spread enhanced by autophagy components","volume":"111","author":"SW Bird","year":"2014","journal-title":"Proc Natl Acad Sci U S A"},{"issue":"4","key":"ref40","doi-asserted-by":"crossref","first-page":"619","DOI":"10.1016\/j.cell.2015.01.032","article-title":"Phosphatidylserine vesicles enable efficient en bloc transmission of enteroviruses","volume":"160","author":"YH Chen","year":"2015","journal-title":"Cell"},{"issue":"4","key":"ref41","doi-asserted-by":"crossref","first-page":"e1004045","DOI":"10.1371\/journal.ppat.1004045","article-title":"Coxsackievirus B exits the host cell in shed microvesicles displaying autophagosomal markers","volume":"10","author":"SM Robinson","year":"2014","journal-title":"PLoS Pathog"},{"issue":"2","key":"ref42","doi-asserted-by":"crossref","first-page":"240","DOI":"10.1016\/j.virol.2008.02.016","article-title":"Autophagic machinery activated by dengue virus enhances virus replication","volume":"374","author":"YR Lee","year":"2008","journal-title":"Virology"},{"issue":"25","key":"ref43","doi-asserted-by":"crossref","first-page":"22147","DOI":"10.1074\/jbc.M110.192500","article-title":"Flavivirus NS4A-induced autophagy protects cells against death and enhances virus replication","volume":"286","author":"JE McLean","year":"2011","journal-title":"J Biol Chem"},{"issue":"15","key":"ref44","doi-asserted-by":"crossref","first-page":"8026","DOI":"10.1128\/JVI.00787-15","article-title":"Dengue Virus Inhibition of Autophagic Flux and Dependency of Viral Replication on Proteasomal Degradation of the Autophagy Receptor p62","volume":"89","author":"P Metz","year":"2015","journal-title":"J Virol"},{"issue":"5","key":"ref45","doi-asserted-by":"crossref","first-page":"422","DOI":"10.1016\/j.chom.2010.10.006","article-title":"Dengue virus-induced autophagy regulates lipid metabolism","volume":"8","author":"NS Heaton","year":"2010","journal-title":"Cell Host Microbe"},{"issue":"3","key":"ref46","doi-asserted-by":"crossref","first-page":"1312","DOI":"10.1128\/JVI.02177-12","article-title":"Inhibition of cellular autophagy deranges dengue virion maturation","volume":"87","author":"R Mateo","year":"2013","journal-title":"J Virol"},{"issue":"17","key":"ref47","doi-asserted-by":"crossref","first-page":"8880","DOI":"10.1128\/JVI.00354-15","article-title":"Biology of Zika Virus Infection in Human Skin Cells","volume":"89","author":"R Hamel","year":"2015","journal-title":"J Virol"},{"issue":"5","key":"ref48","doi-asserted-by":"crossref","first-page":"663","DOI":"10.1016\/j.stem.2016.07.019","article-title":"Zika Virus NS4A and NS4B Proteins Deregulate Akt-mTOR Signaling in Human Fetal Neural Stem Cells to Inhibit Neurogenesis and Induce Autophagy","volume":"19","author":"Q Liang","year":"2016","journal-title":"Cell Stem Cell"},{"key":"ref49","first-page":"206","article-title":"The Role of Secretory Autophagy in Zika Virus Transfer through the Placental Barrier","volume":"6","author":"ZW Zhang","year":"2016","journal-title":"Front Cell Infect Microbiol"},{"issue":"2","key":"ref50","doi-asserted-by":"crossref","first-page":"117","DOI":"10.1002\/path.2694","article-title":"Autophagy: assays and artifacts","volume":"221","author":"S Barth","year":"2010","journal-title":"J Pathol"},{"issue":"1","key":"ref51","doi-asserted-by":"crossref","first-page":"157","DOI":"10.1128\/MCB.01082-08","article-title":"Kinase-inactivated ULK proteins inhibit autophagy via their conserved C-terminal domains using an Atg13-independent mechanism","volume":"29","author":"EY Chan","year":"2009","journal-title":"Mol Cell Biol"},{"issue":"7","key":"ref52","doi-asserted-by":"crossref","first-page":"689","DOI":"10.4161\/auto.7.7.15450","article-title":"The requirement of uncoordinated 51-like kinase 1 (ULK1) and ULK2 in the regulation of autophagy","volume":"7","author":"EJ Lee","year":"2011","journal-title":"Autophagy"},{"key":"ref53","first-page":"1","article-title":"The Exoribonuclease Xrn1 Is a Post-Transcriptional Negative Regulator of Autophagy","author":"E Delorme-Axford","year":"2018","journal-title":"Autophagy"},{"key":"ref54","doi-asserted-by":"crossref","first-page":"129","DOI":"10.1007\/978-1-4939-3292-4_7","article-title":"A Transgenic Mouse Model of Poliomyelitis","volume":"1387","author":"S Koike","year":"2016","journal-title":"Methods Mol Biol"},{"issue":"7095","key":"ref55","doi-asserted-by":"crossref","first-page":"885","DOI":"10.1038\/nature04724","article-title":"Suppression of basal autophagy in neural cells causes neurodegenerative disease in mice","volume":"441","author":"T Hara","year":"2006","journal-title":"Nature"},{"issue":"5","key":"ref56","doi-asserted-by":"crossref","first-page":"295","DOI":"10.1038\/nchembio.79","article-title":"Novel targets for Huntington&apos;s disease in an mTOR-independent autophagy pathway","volume":"4","author":"A Williams","year":"2008","journal-title":"Nat Chem Biol"},{"issue":"48","key":"ref57","doi-asserted-by":"crossref","first-page":"19023","DOI":"10.1073\/pnas.0709695104","article-title":"Small molecule regulators of autophagy identified by an image-based high-throughput screen","volume":"104","author":"L Zhang","year":"2007","journal-title":"Proc Natl Acad Sci U S A"},{"issue":"20","key":"ref58","doi-asserted-by":"crossref","first-page":"9018","DOI":"10.1128\/JVI.01448-16","article-title":"Kinetics of HIV-1 Latency Reversal Quantified on the Single-Cell Level Using a Novel Flow-Based Technique","volume":"90","author":"G Martrus","year":"2016","journal-title":"J Virol"},{"key":"ref59","doi-asserted-by":"crossref","DOI":"10.7554\/eLife.32579","article-title":"Transmission genetics of drug-resistant hepatitis C virus","volume":"7","author":"N van Buuren","year":"2018","journal-title":"Elife"},{"issue":"8","key":"ref60","doi-asserted-by":"crossref","first-page":"1197","DOI":"10.4161\/auto.20586","article-title":"AMPK-dependent phosphorylation of ULK1 regulates ATG9 localization","volume":"8","author":"HI Mack","year":"2012","journal-title":"Autophagy"},{"issue":"8","key":"ref61","doi-asserted-by":"crossref","first-page":"924","DOI":"10.4161\/auto.7.8.15860","article-title":"AMPK and mTOR coordinate the regulation of Ulk1 and mammalian autophagy initiation","volume":"7","author":"L Shang","year":"2011","journal-title":"Autophagy"},{"issue":"50","key":"ref62","doi-asserted-by":"crossref","first-page":"E3405","DOI":"10.1073\/pnas.1216539109","article-title":"Autophagy hijacked through viroporin-activated calcium\/calmodulin-dependent kinase kinase-beta signaling is required for rotavirus replication","volume":"109","author":"SE Crawford","year":"2012","journal-title":"Proc Natl Acad Sci U S A"},{"issue":"3","key":"ref63","doi-asserted-by":"crossref","first-page":"471","DOI":"10.1016\/j.cell.2006.01.016","article-title":"TOR signaling in growth and metabolism","volume":"124","author":"S Wullschleger","year":"2006","journal-title":"Cell"},{"issue":"2","key":"ref64","doi-asserted-by":"crossref","first-page":"285","DOI":"10.1016\/j.molcel.2015.05.031","article-title":"Small Molecule Inhibition of the Autophagy Kinase ULK1 and Identification of ULK1 Substrates","volume":"59","author":"DF Egan","year":"2015","journal-title":"Mol Cell"},{"issue":"12","key":"ref65","doi-asserted-by":"crossref","first-page":"1013","DOI":"10.1038\/nchembio.1681","article-title":"A highly potent and selective Vps34 inhibitor alters vesicle trafficking and autophagy","volume":"10","author":"B Ronan","year":"2014","journal-title":"Nat Chem Biol"},{"issue":"1","key":"ref66","doi-asserted-by":"crossref","first-page":"302","DOI":"10.1128\/JVI.05937-11","article-title":"Complex dynamic development of poliovirus membranous replication complexes","volume":"86","author":"GA Belov","year":"2012","journal-title":"J Virol"},{"issue":"4","key":"ref67","doi-asserted-by":"crossref","first-page":"365","DOI":"10.1016\/j.chom.2009.03.007","article-title":"Composition and three-dimensional architecture of the dengue virus replication and assembly sites","volume":"5","author":"S Welsch","year":"2009","journal-title":"Cell Host Microbe"},{"issue":"6","key":"ref68","doi-asserted-by":"crossref","first-page":"764","DOI":"10.4161\/auto.6.6.12709","article-title":"Characterization of autophagosome formation site by a hierarchical analysis of mammalian Atg proteins","volume":"6","author":"E Itakura","year":"2010","journal-title":"Autophagy"},{"issue":"1","key":"ref69","doi-asserted-by":"crossref","first-page":"131","DOI":"10.4161\/auto.5233","article-title":"Consideration about negative controls for LC3 and expression vectors for four colored fluorescent protein-LC3 negative controls","volume":"4","author":"I Tanida","year":"2008","journal-title":"Autophagy"},{"issue":"2","key":"ref70","doi-asserted-by":"crossref","first-page":"129","DOI":"10.1111\/j.1600-0854.2005.00368.x","article-title":"Microtubules facilitate autophagosome formation and fusion of autophagosomes with endosomes","volume":"7","author":"R Kochl","year":"2006","journal-title":"Traffic"},{"issue":"6811","key":"ref71","doi-asserted-by":"crossref","first-page":"488","DOI":"10.1038\/35044114","article-title":"A ubiquitin-like system mediates protein lipidation","volume":"408","author":"Y Ichimura","year":"2000","journal-title":"Nature"},{"issue":"46","key":"ref72","doi-asserted-by":"crossref","first-page":"47704","DOI":"10.1074\/jbc.M407016200","article-title":"Human light chain 3\/MAP1LC3B is cleaved at its carboxyl-terminal Met121 to expose Gly120 for lipidation and targeting to autophagosomal membranes","volume":"279","author":"I Tanida","year":"2004","journal-title":"J Biol Chem"},{"issue":"9","key":"ref73","doi-asserted-by":"crossref","first-page":"1341","DOI":"10.1038\/emboj.2009.80","article-title":"The structure of Atg4B-LC3 complex reveals the mechanism of LC3 processing and delipidation during autophagy","volume":"28","author":"K Satoo","year":"2009","journal-title":"EMBO J"},{"issue":"52","key":"ref74","doi-asserted-by":"crossref","first-page":"37298","DOI":"10.1074\/jbc.C700195200","article-title":"The Atg12-Atg5 conjugate has a novel E3-like activity for protein lipidation in autophagy","volume":"282","author":"T Hanada","year":"2007","journal-title":"J Biol Chem"},{"issue":"Pt 16","key":"ref75","doi-asserted-by":"crossref","first-page":"2685","DOI":"10.1242\/jcs.026005","article-title":"The N-terminus and Phe52 residue of LC3 recruit p62\/SQSTM1 into autophagosomes","volume":"121","author":"E Shvets","year":"2008","journal-title":"J Cell Sci"},{"issue":"7","key":"ref76","doi-asserted-by":"crossref","first-page":"683","DOI":"10.4161\/auto.7.7.15279","article-title":"Dissecting the involvement of LC3B and GATE-16 in p62 recruitment into autophagosomes","volume":"7","author":"E Shvets","year":"2011","journal-title":"Autophagy"},{"issue":"2","key":"ref77","doi-asserted-by":"crossref","first-page":"253","DOI":"10.1083\/jcb.200907015","article-title":"FYCO1 is a Rab7 effector that binds to LC3 and PI3P to mediate microtubule plus end-directed vesicle transport","volume":"188","author":"S Pankiv","year":"2010","journal-title":"J Cell Biol"},{"issue":"2","key":"ref78","doi-asserted-by":"crossref","first-page":"263","DOI":"10.1083\/jcb.151.2.263","article-title":"The reversible modification regulates the membrane-binding state of Apg8\/Aut7 essential for autophagy and the cytoplasm to vacuole targeting pathway","volume":"151","author":"T Kirisako","year":"2000","journal-title":"J Cell Biol"},{"issue":"39","key":"ref79","doi-asserted-by":"crossref","first-page":"40584","DOI":"10.1074\/jbc.M405860200","article-title":"In vivo and in vitro reconstitution of Atg8 conjugation essential for autophagy","volume":"279","author":"Y Ichimura","year":"2004","journal-title":"J Biol Chem"},{"issue":"22","key":"ref80","doi-asserted-by":"crossref","first-page":"12543","DOI":"10.1128\/JVI.00755-07","article-title":"Modification of cellular autophagy protein LC3 by poliovirus","volume":"81","author":"MP Taylor","year":"2007","journal-title":"J Virol"},{"issue":"33","key":"ref81","doi-asserted-by":"crossref","first-page":"22847","DOI":"10.1074\/jbc.M802182200","article-title":"Structural basis for sorting mechanism of p62 in selective autophagy","volume":"283","author":"Y Ichimura","year":"2008","journal-title":"J Biol Chem"},{"issue":"47","key":"ref82","doi-asserted-by":"crossref","first-page":"39275","DOI":"10.1074\/jbc.M112.378109","article-title":"ATG8 family proteins act as scaffolds for assembly of the ULK complex: sequence requirements for LC3-interacting region (LIR) motifs","volume":"287","author":"EA Alemu","year":"2012","journal-title":"J Biol Chem"},{"issue":"6","key":"ref83","doi-asserted-by":"crossref","first-page":"4611","DOI":"10.1128\/JVI.73.6.4611-4621.1999","article-title":"Genetic interaction of flavivirus nonstructural proteins NS1 and NS4A as a determinant of replicase function","volume":"73","author":"BD Lindenbach","year":"1999","journal-title":"J Virol"},{"issue":"7","key":"ref84","doi-asserted-by":"crossref","first-page":"3471","DOI":"10.1128\/JVI.03454-14","article-title":"Mapping the Interactions between the NS4B and NS3 proteins of dengue virus","volume":"89","author":"J Zou","year":"2015","journal-title":"J Virol"},{"issue":"1","key":"ref85","doi-asserted-by":"crossref","first-page":"7","DOI":"10.1038\/nrm3249","article-title":"Canonical and non-canonical autophagy: variations on a common theme of self-eating?","volume":"13","author":"P Codogno","year":"2011","journal-title":"Nat Rev Mol Cell Biol"},{"issue":"27","key":"ref86","doi-asserted-by":"crossref","first-page":"11121","DOI":"10.1073\/pnas.1107969108","article-title":"Ammonia-induced autophagy is independent of ULK1\/ULK2 kinases","volume":"108","author":"H Cheong","year":"2011","journal-title":"Proc Natl Acad Sci U S A"},{"issue":"8","key":"ref87","doi-asserted-by":"crossref","first-page":"1032","DOI":"10.4161\/auto.6.8.13336","article-title":"Beclin 1-independent autophagy induced by a Bcl-XL\/Bcl-2 targeting compound, Z18","volume":"6","author":"S Tian","year":"2010","journal-title":"Autophagy"},{"issue":"12","key":"ref88","doi-asserted-by":"crossref","first-page":"1867","DOI":"10.1038\/cdd.2010.53","article-title":"Arsenic trioxide induces a beclin-1-independent autophagic pathway via modulation of SnoN\/SkiL expression in ovarian carcinoma cells","volume":"17","author":"DM Smith","year":"2010","journal-title":"Cell Death Differ"},{"issue":"9","key":"ref89","doi-asserted-by":"crossref","first-page":"1429","DOI":"10.1080\/15548627.2016.1203489","article-title":"Alternative autophagy, brefeldin A and viral trafficking pathways","volume":"12","author":"C Grose","year":"2016","journal-title":"Autophagy"},{"issue":"6","key":"ref90","doi-asserted-by":"crossref","first-page":"500","DOI":"10.1016\/j.chom.2010.05.013","article-title":"Coronaviruses Hijack the LC3-I-positive EDEMosomes, ER-derived vesicles exporting short-lived ERAD regulators, for replication","volume":"7","author":"F Reggiori","year":"2010","journal-title":"Cell Host Microbe"},{"issue":"2","key":"ref91","doi-asserted-by":"crossref","first-page":"164","DOI":"10.4161\/auto.22743","article-title":"An autophagy-independent role for LC3 in equine arteritis virus replication","volume":"9","author":"I Monastyrska","year":"2013","journal-title":"Autophagy"},{"issue":"9","key":"ref92","doi-asserted-by":"crossref","first-page":"1637","DOI":"10.4161\/auto.29455","article-title":"Japanese encephalitis virus replication is negatively regulated by autophagy and occurs on LC3-I- and EDEM1-containing membranes","volume":"10","author":"M Sharma","year":"2014","journal-title":"Autophagy"},{"issue":"3","key":"ref93","doi-asserted-by":"crossref","first-page":"405","DOI":"10.1016\/j.bbrc.2008.04.098","article-title":"Segregation and rapid turnover of EDEM1 by an autophagy-like mechanism modulates standard ERAD and folding activities","volume":"371","author":"T Cali","year":"2008","journal-title":"Biochem Biophys Res Commun"},{"issue":"9","key":"ref94","doi-asserted-by":"crossref","first-page":"1440","DOI":"10.1080\/15548627.2016.1191724","article-title":"Autophagy proteins are not universally required for phagosome maturation","volume":"12","author":"M Cemma","year":"2016","journal-title":"Autophagy"},{"issue":"2","key":"ref95","doi-asserted-by":"crossref","first-page":"239","DOI":"10.1016\/j.chom.2014.01.006","article-title":"A LC3-interacting motif in the influenza A virus M2 protein is required to subvert autophagy and maintain virion stability","volume":"15","author":"R Beale","year":"2014","journal-title":"Cell Host Microbe"},{"issue":"14","key":"ref96","doi-asserted-by":"crossref","first-page":"6570","DOI":"10.1128\/JVI.74.14.6570-6580.2000","article-title":"Formation of the poliovirus replication complex requires coupled viral translation, vesicle production, and viral RNA synthesis","volume":"74","author":"D Egger","year":"2000","journal-title":"J Virol"},{"issue":"2","key":"ref97","doi-asserted-by":"crossref","first-page":"676","DOI":"10.1016\/0042-6822(92)90522-Q","article-title":"Immunocytochemical localization of capsid-related particles in subcellular fractions of poliovirus-infected cells","volume":"188","author":"T Pfister","year":"1992","journal-title":"Virology"},{"key":"ref98","doi-asserted-by":"crossref","first-page":"269","DOI":"10.3389\/fmicb.2017.00269","article-title":"The Autophagic Machinery in Viral Exocytosis","volume":"8","author":"C Munz","year":"2017","journal-title":"Front Microbiol"},{"issue":"4","key":"ref99","doi-asserted-by":"crossref","first-page":"537","DOI":"10.1083\/jcb.200911149","article-title":"Unconventional secretion of Pichia pastoris Acb1 is dependent on GRASP protein, peroxisomal functions, and autophagosome formation","volume":"188","author":"R Manjithaya","year":"2010","journal-title":"J Cell Biol"},{"issue":"4","key":"ref100","doi-asserted-by":"crossref","first-page":"527","DOI":"10.1083\/jcb.200911154","article-title":"Unconventional secretion of Acb1 is mediated by autophagosomes","volume":"188","author":"JM Duran","year":"2010","journal-title":"J Cell Biol"},{"issue":"6","key":"ref101","doi-asserted-by":"crossref","first-page":"979","DOI":"10.1083\/jcb.201106098","article-title":"Biogenesis of a novel compartment for autophagosome-mediated unconventional protein secretion","volume":"195","author":"C Bruns","year":"2011","journal-title":"J Cell Biol"},{"key":"ref102","doi-asserted-by":"crossref","DOI":"10.7554\/eLife.11205","article-title":"Translocation of interleukin-1beta into a vesicle intermediate in autophagy-mediated secretion","volume":"4","author":"M Zhang","year":"2015","journal-title":"Elife"},{"issue":"9","key":"ref103","doi-asserted-by":"crossref","first-page":"1161","DOI":"10.1091\/mbc.e16-11-0762","article-title":"The link between autophagy and secretion: a story of multitasking proteins","volume":"28","author":"H Farhan","year":"2017","journal-title":"Mol Biol Cell"},{"issue":"7445","key":"ref104","doi-asserted-by":"crossref","first-page":"367","DOI":"10.1038\/nature12029","article-title":"A pathogenic picornavirus acquires an envelope by hijacking cellular membranes","volume":"496","author":"Z Feng","year":"2013","journal-title":"Nature"},{"issue":"10","key":"ref105","first-page":"1207","article-title":"Malnutrition as an underlying cause of childhood deaths associated with infectious diseases in developing countries","volume":"78","author":"AL Rice","year":"2000","journal-title":"Bull World Health Organ"},{"issue":"4523","key":"ref106","doi-asserted-by":"crossref","first-page":"916","DOI":"10.1126\/science.6272391","article-title":"Cloned poliovirus complementary DNA is infectious in mammalian cells","volume":"214","author":"VR Racaniello","year":"1981","journal-title":"Science"},{"issue":"11","key":"ref107","doi-asserted-by":"crossref","first-page":"2281","DOI":"10.1038\/nprot.2013.143","article-title":"Genome engineering using the CRISPR-Cas9 system","volume":"8","author":"FA Ran","year":"2013","journal-title":"Nat Protoc"},{"issue":"7610","key":"ref108","doi-asserted-by":"crossref","first-page":"159","DOI":"10.1038\/nature18631","article-title":"Genetic dissection of Flaviviridae host factors through genome-scale CRISPR screens","volume":"535","author":"CD Marceau","year":"2016","journal-title":"Nature"}],"updated-by":[{"DOI":"10.1371\/journal.pbio.2006926","type":"new_version","label":"New version","source":"publisher","updated":{"date-parts":[[2019,1,16]],"date-time":"2019-01-16T00:00:00Z","timestamp":1547596800000}}],"container-title":["PLOS Biology"],"original-title":[],"language":"en","link":[{"URL":"http:\/\/dx.plos.org\/10.1371\/journal.pbio.2006926","content-type":"unspecified","content-version":"vor","intended-application":"similarity-checking"}],"deposited":{"date-parts":[[2020,11,21]],"date-time":"2020-11-21T18:10:46Z","timestamp":1605982246000},"score":1,"resource":{"primary":{"URL":"https:\/\/dx.plos.org\/10.1371\/journal.pbio.2006926"}},"subtitle":[],"editor":[{"given":"Bill","family":"Sugden","sequence":"first","affiliation":[],"role":[{"vocabulary":"crossref","role":"editor"}]}],"short-title":[],"issued":{"date-parts":[[2019,1,4]]},"references-count":108,"journal-issue":{"issue":"1","published-online":{"date-parts":[[2019,1,4]]}},"URL":"https:\/\/doi.org\/10.1371\/journal.pbio.2006926","relation":{"new_version":[{"id-type":"doi","id":"10.1371\/journal.pbio.2006926","asserted-by":"object"}]},"ISSN":["1545-7885"],"issn-type":[{"value":"1545-7885","type":"electronic"}],"subject":[],"published":{"date-parts":[[2019,1,4]]}}}