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(2007) Tetr\u00e1podos continentales del Cret\u00e1cico de la Argentina: una s\u00edntesis acutalizada. In: Archangelsky S, S\u00e1nchez T, Tonni EP, editors. Publicaci\u00f3n Especial 11 Ameghiniana 50\u00b0 aniversario. Buenos Aires: Asociaci\u00f3n Paleontol\u00f3gica Argentina, pp. 137\u2013153."},{"issue":"6","key":"ref5","doi-asserted-by":"crossref","first-page":"e20011","DOI":"10.1371\/journal.pone.0020011","article-title":"A new crocodylian from the Late Maastrichtian of Spain: Implications for the initial radiation of crocodyloids","volume":"6","author":"E Pu\u00e9rtolas","year":"2011","journal-title":"PLoS ONE"},{"key":"ref6","first-page":"37","article-title":"\u00dcber die <italic>Lacerta gigantea<\/italic> der Vorwet. Denksch K Akad Wiss M\u00fcnchen","volume":"6","author":"ST von S\u00f6mmerring","year":"1816","journal-title":"Kl Math Natwiss"},{"key":"ref7","unstructured":"Buffetaut E (1987) A Short History of Vertebrate Palaeontology. London: Croom Helm. 223 p."},{"key":"ref8","unstructured":"Langstom W (1973) The crocodilian skull in historical perspective. In: Gans C, Parson TS, editors. Biology of the Reptilia, Vol. 4. London: Academic Press. pp. 263\u2013284."},{"key":"ref9","unstructured":"Steel R (1973) Crocodylia. Handbuch der Pal\u00e4oherpetologie, Teil 16. Stuttgart: Gustav Fischer Verlag. 116 p."},{"key":"ref10","unstructured":"Andrews CW (1913) A Descriptive Catalogue of the Marine Reptiles of the Oxford Clay, Part Two. London: British Museum (Natural History). 285 p."},{"key":"ref11","first-page":"1","article-title":"Die Meer-Krocodilier (Thalattosuchia) des oberen Jura unter specieller ber\u00fccksichtigung von <italic>Dacosaurus<\/italic> und <italic>Geosaurus<\/italic>","volume":"49","author":"E Fraas","year":"1902","journal-title":"Paleontographica"},{"key":"ref12","doi-asserted-by":"crossref","first-page":"269","DOI":"10.1080\/002411600750053835","article-title":"Salt glands in a Tithonian metriorhynchid crocodyliform and their physiological significance","volume":"33","author":"M Fern\u00e1ndez","year":"2000","journal-title":"Lethaia"},{"key":"ref13","doi-asserted-by":"crossref","first-page":"79","DOI":"10.1007\/s00114-007-0296-1","article-title":"Salt glands in the Jurassic metriorhynchid <italic>Geosaurus<\/italic>: implications for the evolution of osmoregulation in Mesozoic crocodyliforms","volume":"95","author":"M Fern\u00e1ndez","year":"2008","journal-title":"Naturwissenschaften"},{"key":"ref14","doi-asserted-by":"crossref","first-page":"801","DOI":"10.1111\/j.1096-3642.2009.00571.x","article-title":"The evolution of Metriorhynchoidea (Mesoeucrocodylia, Thalattosuchia): an integrated approach using geometrics morphometrics, analysis of disparity and biomechanics","volume":"158","author":"MT Young","year":"2010","journal-title":"Zool J Linn Soc"},{"key":"ref15","doi-asserted-by":"crossref","first-page":"1199","DOI":"10.1111\/j.1096-3642.2011.00734.x","article-title":"Body size estimation and evolution in metriorhynchid crocodylomorphs: implications for species diversificiation and niche partitioning","volume":"163","author":"MT Young","year":"2011","journal-title":"Zool J Linn Soc"},{"key":"ref16","doi-asserted-by":"crossref","first-page":"913","DOI":"10.1098\/rsbl.2011.0357","article-title":"Craniofacial form and function in Metriorhynchidae (Crocodylomorpha: Thalattosuchia): modelling phenotypic evolution with maximum likelihood methods","volume":"7","author":"MT Young","year":"2011","journal-title":"Biol Lett"},{"key":"ref17","doi-asserted-by":"crossref","first-page":"550","DOI":"10.1016\/j.gr.2010.07.007","article-title":"The oldest known metriorhynchid crocodylian from the Middle Jurassic of North-eastern Italy: <italic>Neptunidraco ammoniticus<\/italic> gen. et sp. nov","volume":"19","author":"A Cau","year":"2011","journal-title":"Gondwana Res"},{"key":"ref18","article-title":"The oldest known metriorhynchid super-predator: a new genus and species from the Middle Jurassic of England, with implications for serration and mandibular evolution in predacious clades","author":"MT Young","journal-title":"J Syst Palaeontol"},{"key":"ref19","first-page":"277","article-title":"Ein <italic>Geosaurus<\/italic>-fragment (Reptilia, Thalattosuchia) aus dem oberen Malm von Evilard bei Biel","volume":"7","author":"O Rieppel","year":"1979","journal-title":"Burgergemeinde Bern Naturhistorisches Museum Jahrbuch"},{"key":"ref20","doi-asserted-by":"crossref","first-page":"467","DOI":"10.1016\/S0016-6995(00)80080-7","article-title":"<italic>Enaliosuchus macrospondylus<\/italic> Koken 1883 (Crocodylia, Metriorhynchidae) du Valanginien de Barret-le-Bas (Hautes Alpes, France): un cas unique de remont\u00e9e des narines externes parmi les crocodiliens","volume":"33","author":"S Hua","year":"2000","journal-title":"Geobios"},{"key":"ref21","article-title":"A new specimen of <italic>Cricosaurus saltillensis<\/italic> (Crocodylomorpha: Metriorhynchidae) from the Upper Jurassic of Mexico: evidence for craniofacial convergence within Metriorhynchidae","author":"MC Buchy","journal-title":"Oryctos"},{"key":"ref22","doi-asserted-by":"crossref","first-page":"1","DOI":"10.1127\/0077-7749\/2008\/0249-0001","article-title":"New occurrence of the genus <italic>Dakosaurus<\/italic> (Reptilia, Thalattosuchia) in the Upper Jurassic of north-eastern Mexico with comments upon skull architecture of <italic>Dakosaurus<\/italic> and <italic>Geosaurus<\/italic>","volume":"249","author":"M-C Buchy","year":"2008","journal-title":"N Jahrb Geol Pal\u00e4ontol Abh"},{"key":"ref23","doi-asserted-by":"crossref","first-page":"1467","DOI":"10.1139\/e02-060","article-title":"<italic>Geosaurus vignaudi<\/italic> n.sp. 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Unpublished PhD thesis. Poitiers: Universit\u00e9 de Poitiers. 245 pp."},{"key":"ref29","first-page":"497","article-title":"Kimmeridgian metriorhynchid crocodiles from England","volume":"39","author":"DR Grange","year":"1996","journal-title":"Palaeontology"},{"key":"ref30","doi-asserted-by":"crossref","first-page":"341","DOI":"10.1127\/njgpa\/153\/1977\/341","article-title":"<italic>Metriorhynchus casamiquelai<\/italic> n. sp. (Crocodylia, Thalattosuchia), a marine crocodile from the Jurassic (Callovian) of Chile, South America","volume":"153","author":"Z Gasparini","year":"1977","journal-title":"Neues Jahrb Geol Pal\u00e4ontol"},{"key":"ref31","first-page":"335","article-title":"Un nuevo esp\u00e9cimen de cocodrilo marino del Jur\u00e1sico Medio del norte de Chile: revalidaci\u00f3n de <italic>Metriorhynchus westermanni<\/italic> (Crocodyliformes: Metriorhynchidae)","volume":"35","author":"Z Gasparini","year":"2008","journal-title":"Rev Geol Chile"},{"key":"ref32","first-page":"5","article-title":"Catalogue descriptif, anatomique, g\u00e9ologique et historique des fossiles pr\u00e9sent\u00e9s \u00e0 l&apos;exposition \u201cLes Crocodiliens fossiles de Normandie\u201d (6 novembre\u201314 d\u00e9cembre 2008)","volume":"95","author":"Y Lepage","year":"2008","journal-title":"Bull Soc G\u00e9ol Normandies Amis Mus Hayre"},{"key":"ref33","unstructured":"Pereda Suberbiola X, Ruiz-Ome\u00f1aca JI, Canudo JI, Torcida F, Sanz JL (2012) Dinosaur faunas from the Early Cretaceous (Valanginian \u2013 Albian) of Spain. In: Godefroit P, editor. Bernissart Dinosaurs and Early Cretaceous Terrestrial Ecosystems. Bloomington: Indiana University Press. pp. 379\u2013408."},{"key":"ref34","doi-asserted-by":"crossref","first-page":"77","DOI":"10.1127\/njgpa\/239\/2006\/77","article-title":"A megatheropod tooth from the late Tithonian-lower Berriasian (Jurassic-Cretaceous transition) of Galve (Aragon, NE Spain)","volume":"239","author":"JI Canudo","year":"2006","journal-title":"Neues Jahrb Geol Palaontol Abh"},{"key":"ref35","doi-asserted-by":"crossref","first-page":"1009","DOI":"10.1111\/j.1475-4983.2009.00898.x","article-title":"High European sauropod dinosaur diversity during Jurassic-Cretaceous transition in Riodeva (Teruel, Spain)","volume":"5","author":"R Royo-Torres","year":"2009","journal-title":"Palaeontology"},{"key":"ref36","first-page":"101","article-title":"Bibliograf\u00eda del registro espa\u00f1ol de vertebrados mesozoicos","volume":"94","author":"JM Quesada","year":"1998","journal-title":"Bol R Soc Esp Hist Nat Secc Geol"},{"key":"ref37","first-page":"1245","article-title":"Mesozoic marine reptiles from the Iberian Peninsula","volume":"10","author":"N Bardet","year":"2008","journal-title":"Geo-Temas"},{"key":"ref38","first-page":"93","article-title":"El primer Crocodylia (Mesosuchia: Teleosauridae) del Jur\u00e1sico Superior de Asturias, Espa\u00f1a. Naturalia patag\u00f3nica","volume":"3","author":"RD Mart\u00ednez","year":"1995","journal-title":"Ciencias de la Tierra"},{"key":"ref39","unstructured":"Ruiz-Ome\u00f1aca JI, Pi\u00f1uela L, Garc\u00eda-Ramos JC, Bardet N, Pereda-Suberbiola X (2007) Dientes de reptiles marinos (Plesiosauroidea y Thalattosuchia) del Jur\u00e1sico Superior de Asturias. In: Braga JC, Checa A, Company M, editors. XXIII Jornadas de la Sociedad Espa\u00f1ola de Paleontolog\u00eda. Caravaca de la Cruz, 3\u20136 de octubre de 2007. Granada: Libro de Res\u00famenes IGME y Universidad de Granada. pp. 204\u2013205."},{"key":"ref40","first-page":"121","article-title":"Nota sobre a presence do g\u00e9nero <italic>Pelagosaurus<\/italic> no Lias do Tomar","volume":"41","author":"OV Ferreira","year":"1959","journal-title":"An Fac Cien do Porto"},{"key":"ref41","first-page":"1","article-title":"Un mesosuquiano do Li\u00e1sico do Tomar (Portugal). Considera\u00e7\u00f5es sobre a origem dos crocodilos","volume":"13","author":"MT Antunes","year":"1967","journal-title":"Mem\u00f3rias dos Servi\u00e7os Geol\u00f3gicos de Portugal, Nova S\u00e9rie"},{"key":"ref42","unstructured":"Ruiz-Ome\u00f1aca JI, Pi\u00f1uela L, Garc\u00eda-Ramos JC (2010) <italic>Dakosaurus<\/italic> sp. (Thalattosuchia: Metriorhynchidae) en el Kimmeridgiense de Colunga (Asturias). In: Ruiz-Ome\u00f1aca JI, Pi\u00f1uela L, Garc\u00eda-Ramos JL, editors. Comunicaciones del V Congreso del Jur\u00e1sico de Espa\u00f1a. Museo del Jur\u00e1sico de Asturias (MUJA), Colunga, 8\u201311 de septiembre de 2010. Colunga: Museo del Jur\u00e1sico de Asturias. pp. 193\u2013199."},{"issue":"9","key":"ref43","doi-asserted-by":"crossref","first-page":"e44986","DOI":"10.1371\/journal.pone.0044985","article-title":"The cranial osteology and feeding ecology of the metriorhynchid crocodylomorph genera <italic>Dakosaurus<\/italic> and <italic>Plesiosuchus<\/italic> from the Late Jurassic of Europe","volume":"7","author":"MT Young","year":"2012","journal-title":"PLoS ONE"},{"key":"ref44","first-page":"14","article-title":"El \u201cCocodrilo de Ricla\u201d el cr\u00e1neo de metriorr\u00ednquido m\u00e1s completo del Jur\u00e1sico de Espa\u00f1a","volume":"1","author":"JI Canudo","year":"1997","journal-title":"Cuadernos de Paleontolog\u00eda Aragonesa"},{"key":"ref45","first-page":"13","article-title":"Un viaje a trav\u00e9s del tiempo: La sala Lucas Mallada","volume":"1","author":"JI Canudo","year":"1998","journal-title":"Aragonia"},{"key":"ref46","unstructured":"Mel\u00e9ndez G, Soria-Llop C (2000) La protecci\u00f3n del patrimonio paleontol\u00f3gico en Espa\u00f1a:Concienciaci\u00f3n social y legislaci\u00f3n, y papel de la Sociedad Espa\u00f1ola de Paleontolog\u00eda. In: Barettino D, Wimbledon WAP, Gallego E, editors. Patrimonio Geologico, Conservacion y Gestion. Volumen 128 de Cursos y Congresos. Madrid: Instituto Tecnol\u00f3gico Geominero de Espa\u00f1a. pp. 197\u2013215."},{"key":"ref47","first-page":"160","article-title":"El patrimonio paleontol\u00f3gico en Espa\u00f1a: una aproximaci\u00f3n somera","volume":"9","author":"G Mel\u00e9ndez","year":"2001","journal-title":"Ense\u00f1anza de las Ciencias de la Tierra"},{"key":"ref48","first-page":"16","article-title":"El patrimonio geol\u00f3gico y paleontol\u00f3gico de las comarcas aragonesas","volume":"9","author":"E Li\u00f1\u00e1n","year":"2002","journal-title":"Naturaleza Aragonesa"},{"key":"ref49","first-page":"8","article-title":"Introducci\u00f3n a la paleontolog\u00eda del Jur\u00e1sico de Ricla","volume":"2","author":"I P\u00e9rez Urresti","year":"2003","journal-title":"Cuadernos de Paleontolog\u00eda Aragonesa"},{"key":"ref50","first-page":"95","article-title":"Nuestro museo y sus protagonistas. 3. <italic>Metriorhynchus<\/italic> (Von Meyer, 1830)","volume":"14","year":"2005","journal-title":"Naturaleza Aragonesa"},{"key":"ref51","unstructured":"P\u00e9rez Urresti I (2005) Gu\u00eda de los f\u00f3siles de Ricla. In: Sauras Herrera MD, G\u00e1mez Vintaned JA, editors. Paleontolog\u00eda de Ricla. Cuadernos de Paleontolog\u00eda Aragonesa 3. Zaragoza: Asociaci\u00f3n Cultural Bajo Jal\u00f3n. pp. 50\u201386"},{"key":"ref52","unstructured":"G\u00e1mez Vintaned JA, Cl\u00faa Mendez JM, Sauras Herrera MD (2006) Tesoros F\u00f3siles de Arag\u00f3n. Cat\u00e1logo de la exposici\u00f3n. 16 de febrero- 11 de marzo de 2006. Centro Joaqu\u00edn Roncal (Fundaci\u00f3n CAI-ASC). Zaragoza: Sociedad de Amigos del Museo Paleontol\u00f3gico de la Universidad de Zaragoza. 32 p."},{"key":"ref53","unstructured":"Li\u00f1\u00e1n Guijarro E, G\u00e1mez Vintaned JA (2006) El agua en Arag\u00f3n durante las eras geol\u00f3gicas. In: Centellas Salamero R, editor. Aquaria: Agua, Territorio y Paisaje en Arag\u00f3n [Cat\u00e1logo de la Exposici\u00f3n]. Zaragoza: Gobierno de Arag\u00f3n and Diputaci\u00f3n Provincial de Zaragoza. pp. 18\u201343."},{"key":"ref54","unstructured":"Genera i Monells M, Mel\u00e9ndez Hevia G (2010) Or\u00edgenes y evoluci\u00f3n de la especie humana. Reus: Pragma General d&apos;Edicions. 112 p."},{"key":"ref55","unstructured":"Li\u00f1\u00e1n E (2010) El Museo Paleontol\u00f3gico de la Universidad de Zaragoza. Veinticinco a\u00f1os despu\u00e9s. In: G\u00e1mez Vintaned JA, editor. XI Jornadas Aragonesas de Paleontolog\u00eda: La Paleontolog\u00eda en los Museos. Homenaje al Profesor Eladio Li\u00f1\u00e1n Guijarro. Zaragoza: Instituci\u00f3n Fernando el Cat\u00f3lico. pp. 17\u201328"},{"key":"ref56","first-page":"4","article-title":"Un paseo por la historia de lavida a trav\u00e9s de los f\u00f3siles de la sala \u201cLucas Mallada\u201d (Museo Paleontol\u00f3gico de Zaragoza)","volume":"25","author":"LM Sender Palomar","year":"2010","journal-title":"Naturaleza Aragonesa"},{"key":"ref57","unstructured":"Parrilla-Bel J, Canudo JI (2011) Un cr\u00e1neo de Metriorhynchidae (Thalattosuchia) del Jur\u00e1sico Medio de Ricla (Zaragoza, Espa\u00f1a). In: P\u00e9rez-Garc\u00eda A, Gasc\u00f3 F, Gasulla JM, Escaso F, editors. Viajando a Mundos Pret\u00e9ritos. Morella (Castell\u00f3n): Ayuntamiento de Morella. pp. 241\u2013247."},{"key":"ref58","unstructured":"Ramajo J (2006) Evoluci\u00f3n sedimentaria del Calloviense-Oxfordiense en el Sector Central de la Cordillera Ib\u00e9rica (Rama Aragonesa). Unpublished PhD thesis. Zaragoza: Universidad de Zaragoza. 405pp."},{"key":"ref59","unstructured":"Lend\u00ednez A, Gonzalo R, Cabra,P, P\u00e9rez A, San Rom\u00e1n J, <etal>et al<\/etal>.. (1995) Mapa Geol\u00f3gico de Espa\u00f1a. Serie Magna. \u00c9PILA. Hoja n\u00b0 382. Instituto Geol\u00f3gico y Minero de Espa\u00f1a, Madrid +1 report 164 p."},{"key":"ref60","doi-asserted-by":"crossref","first-page":"411","DOI":"10.3989\/egeol.84405-6679","article-title":"S\u00edntesis bioestratigr\u00e1fica del Calloviense de Ricla (Zaragoza, Cordillera Ib\u00e9rica)","volume":"40","author":"L Sequeiros","year":"1984","journal-title":"Estudios Geol\u00f3gicos"},{"key":"ref61","unstructured":"Cariou E, Mel\u00e9ndez G, Sequeiros L, Thierry J (1988) Biochronologie du Callovien de la province d&apos;ammonites subm\u00e9diterrane\u00e9nne: Reconnaissance dans les Cha\u00eenes Ib\u00e9riques des subdivisions fines distingu\u00e9es dans le centre-ouest de la France. In: Rocha BR, Soares AF, editors. Proceedings II International Symposium on Jurassic Stratigraphy (1987). Lisbon: Universidade Nova de Lisbon. pp. 395\u2013406."},{"key":"ref62","unstructured":"Mel\u00e9ndez G, Lardi\u00e9s MD (1988) El Calloviense y Oxfordiense de Ricla (Prov. Zaragoza). In: P\u00e9rez Lorente F., Ureta Gil MS, Comas Rengifo MJ, Fern\u00e1ndez L\u00f3pez, S, editors. III Coloquio Jur\u00e1sico de Espa\u00f1a. Gu\u00eda de Excursiones. Logro\u00f1o: Instituto de Estudios Riojanos. pp. 265\u2013281."},{"key":"ref63","unstructured":"Aurell M, Mel\u00e9ndez G, B\u00e1denas B (2002) East Iberian Basins. In Gibbons W, Moreno T, editors. The Geology of Spain. London: Geological Society. pp 223\u2013235."},{"key":"ref64","unstructured":"G\u00f3mez JJ, Fern\u00e1ndez-L\u00f3pez SR (2004) Jur\u00e1sico Medio. In: Vera JA, editor. Geolog\u00eda de Espa\u00f1a. Madrid: SGE-IGME. pp. 500\u2013503."},{"key":"ref65","doi-asserted-by":"crossref","first-page":"190","DOI":"10.1016\/j.palaeo.2005.11.008","article-title":"The Iberian Middle Jurassic carbonate-platform system: Synthesis of the palaeogeographic elements of its eastern margin (Spain)","volume":"236","author":"JJ G\u00f3mez","year":"2006","journal-title":"Paleogeogr Palaeoclimatol Palaeoecol"},{"key":"ref66","first-page":"191","article-title":"The stratigraphy of the Upper Bathonian to Middle Oxfordian succession of the Aragonese Branch of the Cordillera Ib\u00e9rica (Spain) and its European context","volume":"110","author":"KN Page","year":"2004","journal-title":"Rivista Italiana di Paleontologia e Stratigrafia"},{"key":"ref67","doi-asserted-by":"crossref","first-page":"774","DOI":"10.1111\/j.1096-0031.2008.00217.x","article-title":"TNT, a free program for phylogenetic analysis","volume":"24","author":"PA Goloboff","year":"2008","journal-title":"Cladistics"},{"key":"ref68","unstructured":"Hay OP (1930) Second Bibliography and Catalogue of the Fossil Vertebrata of North America 2. Washington DC: Carnegie Institute Washington. 1094 p."},{"key":"ref69","doi-asserted-by":"crossref","first-page":"323","DOI":"10.1098\/rstb.1970.0028","article-title":"A revision of the Jurassic reptile <italic>Hallopus victor<\/italic> (Marsh), with remarks on the classification of crocodiles","volume":"257","author":"AD Walker","year":"1970","journal-title":"Philos Transact R Soc Lond B Biol Sci"},{"key":"ref70","first-page":"409","article-title":"Die Meerkrokodile (Thalattosuchia n. g.) eine neue sauriergruppe der Juraformation","volume":"57","author":"E Fraas","year":"1901","journal-title":"Jahresh Ver vaterl Natkd W\u00fcrtt"},{"key":"ref71","doi-asserted-by":"crossref","unstructured":"Fitzinger LJFJ (1843) Systema Reptilium. Wien: Braum\u00fcller et Seidel. 136 p.","DOI":"10.5962\/bhl.title.4694"},{"key":"ref72","unstructured":"von Meyer CEH (1832) Pal\u00e4ologica zur Geschichte der Erde und irher Gesch\u00f6pfe. Frankfurt: Meyer. 560pp."},{"key":"ref73","doi-asserted-by":"crossref","first-page":"49","DOI":"10.1080\/02724634.1996.10011283","article-title":"Closure of neurocentral sutures during crocodilian ontogeny: implications for maturity assessment in fossil archosaurs","volume":"16","author":"CA Brochu","year":"1996","journal-title":"J Vert Paleontol"},{"key":"ref74","doi-asserted-by":"crossref","first-page":"163","DOI":"10.1017\/S1477201908002605","article-title":"Skull anatomy of <italic>Dakosaurus andiniensis<\/italic> (Thalattosuchia: Crocodylomorpha) and the phylogenetic position of Thalattosuchia","volume":"7","author":"D Pol","year":"2009","journal-title":"J Syst Palaeontol"},{"key":"ref75","first-page":"49","article-title":"Revision of the genus <italic>Enaliosuchus<\/italic> Koken, 1883 (Archosauromorpha: Metriorhynchidae) from the Early Cretaceous of Northwestern Germany","volume":"42","author":"HV Karl","year":"2006","journal-title":"Stud Geol Salmanticensia"},{"key":"ref76","doi-asserted-by":"crossref","first-page":"785","DOI":"10.1016\/j.crpv.2006.03.003","article-title":"A new thalattosuchian crocodyliform from the Tithonian (Upper Jurassic) of northeastern Mexico","volume":"5","author":"MC Buchy","year":"2006","journal-title":"Comptes Rendus Pal\u00e9vol"},{"key":"ref77","doi-asserted-by":"crossref","first-page":"702","DOI":"10.1671\/039.029.0323","article-title":"Paranasal sinus system of <italic>Geosaurus araucanensis<\/italic> and the homology of the antorbital fenestra of metriorhynchids (Thalattosuchia: Crocodylomorpha)","volume":"29","author":"MS Fern\u00e1ndez","year":"2009","journal-title":"J Vert Paleontol"},{"key":"ref78","doi-asserted-by":"crossref","first-page":"1009","DOI":"10.1671\/0272-4634(2006)26[1009:SGITFC]2.0.CO;2","article-title":"Salt glands in the fossil crocodile <italic>Metriorhynchus<\/italic>","volume":"26","author":"R Gandola","year":"2006","journal-title":"J Vert Paleontol"},{"key":"ref79","doi-asserted-by":"crossref","first-page":"369","DOI":"10.1080\/02724634.2011.550361","article-title":"A metriorhynchid crocodyliform braincase from the northern Chile","volume":"31","author":"MS Fern\u00e1ndez","year":"2011","journal-title":"J Vert Paleontol"},{"key":"ref80","doi-asserted-by":"crossref","first-page":"88","DOI":"10.1080\/02724634.2009.10010364","article-title":"The skull of <italic>Teleosaurus cadomensis<\/italic> (Crocodylomorpha, Thalattosuchia), and phylogenetic analysis of Thalattosuchia","volume":"29","author":"S Jouve","year":"2009","journal-title":"J Vert Paleontol"},{"key":"ref81","doi-asserted-by":"crossref","first-page":"1451","DOI":"10.1080\/02724634.2010.501442","article-title":"The evolution of extreme hypercarnivory in Metriorhynchidae (Mesoeucrocodylia: Thalattosuchia) based on evidence from microscopic denticle morphology","volume":"30","author":"MB Andrade","year":"2010","journal-title":"J Vert Paleontol"},{"key":"ref82","doi-asserted-by":"crossref","unstructured":"Massare JA (1994) Swimming capabilities of Mesozoic marine reptiles: a review. In: Maddock L, Bone Q, and Rayner JMV, editors. Mechanics and Physiology of Animal Swimming. Cambrige: Cambridge University Press. pp. 133\u2013149, pp. 205\u2013229.","DOI":"10.1017\/CBO9780511983641.011"},{"key":"ref83","doi-asserted-by":"crossref","first-page":"555","DOI":"10.1111\/j.1469-7580.2009.01137.x","article-title":"Shape and mechanics in thalattosuchian (Crocodylomorpha) skulls: Implications for feeding behavior and niche partitioning","volume":"215","author":"SE Pierce","year":"2009","journal-title":"J Anat"},{"key":"ref84","doi-asserted-by":"crossref","first-page":"1147","DOI":"10.1002\/ar.22491","article-title":"Tooth-on-tooth interlocking occlusion suggests macrophagy in the Mesozoic marine crocodylomorph <italic>Dakosaurus<\/italic>","volume":"295","author":"MT Young","year":"2012","journal-title":"Anat Rec"},{"key":"ref85","doi-asserted-by":"crossref","first-page":"1057","DOI":"10.1111\/j.1475-4983.2009.00904.x","article-title":"Morphospace occupation in thalattosuchian crocodylomorphs: Skull shape variation, species delineation, and temporal patterns","volume":"52","author":"SE Pierce","year":"2009","journal-title":"Palaeontology"}],"container-title":["PLoS ONE"],"original-title":[],"language":"en","link":[{"URL":"http:\/\/dx.plos.org\/10.1371\/journal.pone.0054275","content-type":"unspecified","content-version":"vor","intended-application":"similarity-checking"}],"deposited":{"date-parts":[[2025,4,29]],"date-time":"2025-04-29T18:21:08Z","timestamp":1745950868000},"score":1,"resource":{"primary":{"URL":"https:\/\/dx.plos.org\/10.1371\/journal.pone.0054275"}},"subtitle":[],"editor":[{"given":"Richard J.","family":"Butler","sequence":"first","affiliation":[],"role":[{"vocabulary":"crossref","role":"editor"}]}],"short-title":[],"issued":{"date-parts":[[2013,1,23]]},"references-count":85,"journal-issue":{"issue":"1","published-online":{"date-parts":[[2013,1,23]]}},"URL":"https:\/\/doi.org\/10.1371\/journal.pone.0054275","relation":{},"ISSN":["1932-6203"],"issn-type":[{"value":"1932-6203","type":"electronic"}],"subject":[],"published":{"date-parts":[[2013,1,23]]}}}