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Harmful Algae, Xunta de Galicia and Intergovernmental Oceanographic Commission of UNESCO. 245\u2013248."},{"key":"ref5","doi-asserted-by":"crossref","first-page":"91","DOI":"10.1016\/S0168-1605(00)00382-2","article-title":"Ciguatera: recent advances but the risk remains","volume":"61","author":"L Lehane","year":"2000","journal-title":"Int J Food Microbiol"},{"key":"ref6","doi-asserted-by":"crossref","first-page":"674","DOI":"10.1016\/j.toxicon.2009.05.032","article-title":"Ciguatera risk management in French Polynesia: The case study of Raivavae Island (Australes Archipelago)","volume":"56","author":"M Chinain","year":"2010","journal-title":"Toxicon"},{"key":"ref7","doi-asserted-by":"crossref","first-page":"698","DOI":"10.1016\/j.toxicon.2010.02.026","article-title":"Ciguatera fish poisoning and sea surface temperatures in the Caribbean Sea and the West Indies","volume":"56","author":"PA Tester","year":"2010","journal-title":"Toxicon"},{"key":"ref8","doi-asserted-by":"crossref","first-page":"97","DOI":"10.1016\/S0041-0101(00)00161-6","article-title":"The changing face of ciguatera","volume":"39","author":"RJ Lewis","year":"2001","journal-title":"Toxicon"},{"key":"ref9","doi-asserted-by":"crossref","first-page":"656","DOI":"10.1016\/j.toxicon.2009.05.036","article-title":"Ciguatera incidence and fish toxicity in Okinawa, Japan","volume":"56","author":"N Oshiro","year":"2010","journal-title":"Toxicon"},{"key":"ref10","doi-asserted-by":"crossref","first-page":"917","DOI":"10.2331\/suisan.74.917","article-title":"The occurrence of palytoxin-like poisoning and ciguatera in parts of the main land of Japan","volume":"74","author":"S Taniyama","year":"2008","journal-title":"Nippon Suisan Gakkaishi"},{"key":"ref11","first-page":"13","article-title":"Occurrence of the toxic benthic dinoflagellate <italic>Gambierdiscus toxicus<\/italic> in Ago Bay, central part of Japan","volume":"74","author":"A Ishikawa","year":"2010","journal-title":"Bull Jpn Soc Fish Oceanogr"},{"key":"ref12","doi-asserted-by":"crossref","first-page":"53","DOI":"10.2149\/tmh.2011-03","article-title":"Ciguatera fish poisoning on Kakeroma Island, Kagoshima Prefecture, Japan","volume":"39","author":"N Oshiro","year":"2011","journal-title":"Trop Med Health"},{"key":"ref13","doi-asserted-by":"crossref","first-page":"1021","DOI":"10.2331\/suisan.43.1021","article-title":"Finding of a dinoflagellate as a likely culprit of ciguatera","volume":"43","author":"T Yasumoto","year":"1977","journal-title":"Bull Japan Soc Sci Fish"},{"key":"ref14","doi-asserted-by":"crossref","first-page":"67","DOI":"10.2331\/suisan.45.67","article-title":"The thecal structure of a marine toxic dinoflagellate <italic>Gambierdiscus toxicus<\/italic> gen. et sp. nov. collected in a ciguatera-endemic area","volume":"45","author":"R Adachi","year":"1979","journal-title":"Bull Japan Soc Sci Fish"},{"key":"ref15","doi-asserted-by":"crossref","first-page":"10","DOI":"10.1016\/j.hal.2011.06.013","article-title":"<italic>Gambierdiscus excentricus<\/italic> sp. nov. 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In: Proceedings of the Regional Symposium on Environment and Natural Resources 10\u201311<sup>th<\/sup> April 2002 Malaysia. 144\u2013153."},{"key":"ref30","doi-asserted-by":"crossref","first-page":"629","DOI":"10.1111\/j.1756-1051.2004.tb01938.x","article-title":"Marine epibenthic dinoflagellates from Malaysia - a study of live cultures and preserved samples based on light and scanning electron microscopy","volume":"24","author":"N Mohammad-Noor","year":"2004","journal-title":"Nord J Bot"},{"key":"ref31","doi-asserted-by":"crossref","first-page":"143","DOI":"10.1111\/j.1440-1835.2011.00612.x","article-title":"First report of the benthic dinoflagellate, <italic>Gambierdiscus belizeanus<\/italic> (Gonyaulacales: Dinophyceae) for the east coast of Sabah, Malaysian Borneo","volume":"59","author":"CP Leaw","year":"2011","journal-title":"Phycol Res"},{"key":"ref32","doi-asserted-by":"crossref","first-page":"205","DOI":"10.1007\/s12601-011-0016-9","article-title":"Abundance of epiphytic dinoflagellates from coastal waters off Jeju Island, Korea During Autumn 2009","volume":"46","author":"HS Kim","year":"2011","journal-title":"Ocean Sci J"},{"key":"ref33","first-page":"277","article-title":"Algal biotoxin research, monitoring, and management in shellfish aquaculture of Taiwan. Aquaculture and fisheries resources management","volume":"4","author":"HN Chou","year":"2001","journal-title":"In: Proceedings of the Joint Taiwan-Australia Aquaculture and Fisheries Resources and Management Forum"},{"key":"ref34","first-page":"99","article-title":"A floristic study of the marine microalgae along the coast of the Izu Peninsula","volume":"15","author":"Y Hara","year":"1982","journal-title":"Mem Natl Sci Mus"},{"key":"ref35","doi-asserted-by":"crossref","first-page":"2261","DOI":"10.2331\/suisan.57.2261","article-title":"Distributions of benthic dinoflagellates in Akajima Island, Okinawa, Japan","volume":"57","author":"K Koike","year":"1991","journal-title":"Bull Japan Soc Sci Fish"},{"key":"ref36","unstructured":"Fukuyo Y, Imai I, Kodama M, Tamai K (2002) Red tides and other harmful algal blooms in Japan. In: Taylor FJR, Trainer VL, eds. Harmful algal blooms in the PICES region of the North Pacific. PICES Scientific Report No. 23. North Pacific Marine Sci Org (PICES), Canada. 7\u201320."},{"key":"ref37","unstructured":"Omura T, Nagahama Y, Fukuyo Y (2010) Ciguatera causative species found in main land of Japan. In: GEOHAB Meeting Honolulu, Hawaii, USA, 21\u201323 June 2010. Available: <ext-link xmlns:xlink=\"http:\/\/www.w3.org\/1999\/xlink\" ext-link-type=\"uri\" xlink:href=\"http:\/\/www.scor-int.org\/GEOHAB_BHAB_Program_Book.pdf\" xlink:type=\"simple\">http:\/\/www.scor-int.org\/GEOHAB_BHAB_Program_Book.pdf<\/ext-link>. Accessed 13 November 2012."},{"key":"ref38","doi-asserted-by":"crossref","first-page":"685","DOI":"10.2331\/suisan.77.685","article-title":"First report of the benthic dinoflagellate of the genus <italic>Gambierdiscus<\/italic> from western Wakasa Bay in the Sea of Japan","volume":"77","author":"Y Hatayama","year":"2011","journal-title":"Nippon Suisan Gakkaishi"},{"key":"ref39","doi-asserted-by":"crossref","first-page":"344","DOI":"10.1111\/j.1529-8817.2007.00320.x","article-title":"Recognizing dinoflagellate species using ITS rDNA sequences","volume":"43","author":"RW Litaker","year":"2007","journal-title":"J Phycol"},{"key":"ref40","doi-asserted-by":"crossref","first-page":"426","DOI":"10.1111\/j.1529-8817.2010.00950.x","article-title":"Discriminative power of nuclear rDNA sequences for the DNA taxonomy of the dinoflagellate genus <italic>Peridinium<\/italic> (Dinophyceae)","volume":"47","author":"JS Ki","year":"2011","journal-title":"J Phycol"},{"key":"ref41","doi-asserted-by":"crossref","first-page":"319","DOI":"10.1007\/s10811-009-9462-5","article-title":"Molecular genetic divergence of the centric diatom <italic>Cyclotella<\/italic> and <italic>Discostella<\/italic> (Bacillariophyceae) revealed by nuclear ribosomal DNA comparisons","volume":"22","author":"SW Jung","year":"2010","journal-title":"J Appl Phycol"},{"key":"ref42","doi-asserted-by":"crossref","first-page":"65","DOI":"10.1111\/j.1755-0998.2009.02633.x","article-title":"Barcoding diatoms: Is there a good marker?","volume":"9","author":"MBJ Moniz","year":"2009","journal-title":"Mol Ecol Resour"},{"key":"ref43","doi-asserted-by":"crossref","first-page":"739","DOI":"10.1016\/j.toxicon.2009.06.013","article-title":"Growth and toxin production in the ciguatera-causing dinoflagellate <italic>Gambierdiscus polynesiensis<\/italic> (Dinophyceae) in culture","volume":"56","author":"M Chinain","year":"2010","journal-title":"Toxicon"},{"key":"ref44","doi-asserted-by":"crossref","first-page":"754","DOI":"10.1046\/j.1529-8817.2003.02112.x","article-title":"Identification of <italic>Pfiesteria piscicida<\/italic> (Dinophyceae) and <italic>Pfiesteria<\/italic>-like organisms using internal transcribed spacer-specific PCR assays","volume":"39","author":"RW Litaker","year":"2003","journal-title":"J Phycol"},{"key":"ref45","doi-asserted-by":"crossref","first-page":"411","DOI":"10.1111\/j.1529-8817.2005.04168.x","article-title":"Phylogeny of dinoflagellates based on mitochondrial cytochrome b and nuclear small subunit rDNA sequence comparisons","volume":"41","author":"H Zhang","year":"2005","journal-title":"J Phycol"},{"key":"ref46","doi-asserted-by":"crossref","first-page":"251","DOI":"10.1111\/j.1529-8817.2006.00177.x","article-title":"Acquiring scanning electron microscopical, light microscopical and multiple gene sequence data from a single dinoflagellate cell","volume":"42","author":"Y Takano","year":"2005","journal-title":"J Phycol"},{"key":"ref47","doi-asserted-by":"crossref","first-page":"e27983","DOI":"10.1371\/journal.pone.0027983","article-title":"Phylogeography of <italic>Ostreopsis<\/italic> along west Pacific coast, with special reference to a novel clade from Japan","volume":"6","author":"S Sato","year":"2011","journal-title":"PLoS ONE"},{"key":"ref48","doi-asserted-by":"crossref","first-page":"999","DOI":"10.1111\/j.0022-3646.1994.00999.x","article-title":"Identification of group- and strain-specific genetic markers for globally distributed <italic>Alexandrium<\/italic> (Dinophyceae). 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