{"status":"ok","message-type":"work","message-version":"1.0.0","message":{"indexed":{"date-parts":[[2025,11,7]],"date-time":"2025-11-07T08:33:21Z","timestamp":1762504401751,"version":"3.40.5"},"reference-count":82,"publisher":"Public Library of Science (PLoS)","issue":"6","license":[{"start":{"date-parts":[[2021,6,4]],"date-time":"2021-06-04T00:00:00Z","timestamp":1622764800000},"content-version":"vor","delay-in-days":0,"URL":"http:\/\/creativecommons.org\/licenses\/by\/4.0\/"}],"content-domain":{"domain":["www.plosone.org"],"crossmark-restriction":false},"short-container-title":["PLoS ONE"],"abstract":"<jats:p>In modern humans, facial soft tissue thicknesses have been shown to covary with craniometric dimensions. However, to date it has not been confirmed whether these relationships are shared with non-human apes. In this study, we analyze these relationships in chimpanzees (<jats:italic>Pan troglodytes<\/jats:italic>) with the aim of producing regression models for approximating facial soft tissue thicknesses in Plio-Pleistocene hominids. Using CT scans of 19 subjects, 637 soft tissue, and 349 craniometric measurements, statistically significant multiple regression models were established for 26 points on the face and head. Examination of regression model validity resulted in minimal differences between observed and predicted soft tissue thickness values. Assessment of interspecies compatibility using a bonobo (<jats:italic>Pan paniscus<\/jats:italic>) and modern human subject resulted in minimal differences for the bonobo but large differences for the modern human. These results clearly show that (1) soft tissue thicknesses covary with craniometric dimensions in<jats:italic>P<\/jats:italic>.<jats:italic>troglodytes<\/jats:italic>, (2) confirms that such covariation is uniformly present in both extant<jats:italic>Homo<\/jats:italic>and<jats:italic>Pan<\/jats:italic>species, and (3) suggests that chimp-derived regression models have interspecies compatibility with hominids who have similar craniometric dimensions to<jats:italic>P<\/jats:italic>.<jats:italic>troglodytes<\/jats:italic>. As the craniometric dimensions of early hominids, such as South African australopithecines, are more similar to<jats:italic>P<\/jats:italic>.<jats:italic>troglodytes<\/jats:italic>than those of<jats:italic>H<\/jats:italic>.<jats:italic>sapiens<\/jats:italic>, chimpanzee-derived regression models may be used for approximating their craniofacial anatomy. It is hoped that the results of the present study and the reference dataset for facial soft tissue thicknesses of chimpanzees it provides will encourage further research into this topic.<\/jats:p>","DOI":"10.1371\/journal.pone.0245760","type":"journal-article","created":{"date-parts":[[2021,6,4]],"date-time":"2021-06-04T17:45:43Z","timestamp":1622828743000},"page":"e0245760","update-policy":"https:\/\/doi.org\/10.1371\/journal.pone.corrections_policy","source":"Crossref","is-referenced-by-count":3,"title":["Towards the restoration of ancient hominid craniofacial anatomy: Chimpanzee morphology reveals covariation between craniometrics and facial soft tissue thickness"],"prefix":"10.1371","volume":"16","author":[{"ORCID":"https:\/\/orcid.org\/0000-0003-2630-1701","authenticated-orcid":true,"given":"Ryan M.","family":"Campbell","sequence":"first","affiliation":[],"role":[{"role":"author","vocabulary":"crossref"}]},{"given":"Gabriel","family":"Vinas","sequence":"additional","affiliation":[],"role":[{"role":"author","vocabulary":"crossref"}]},{"ORCID":"https:\/\/orcid.org\/0000-0003-1941-2286","authenticated-orcid":true,"given":"Maciej","family":"Henneberg","sequence":"additional","affiliation":[],"role":[{"role":"author","vocabulary":"crossref"}]}],"member":"340","published-online":{"date-parts":[[2021,6,4]]},"reference":[{"volume-title":"Shaping Humanity: How Science, Art and Imagination Help Us Understand Our Origins","year":"2013","author":"J Gurche","key":"pone.0245760.ref001"},{"issue":"5937","key":"pone.0245760.ref002","doi-asserted-by":"crossref","first-page":"136","DOI":"10.1126\/science.325_136","article-title":"Bringing hominins back to life","volume":"325","author":"M Balter","year":"2009","journal-title":"Science"},{"key":"pone.0245760.ref003","volume-title":"Evolution: The Human Story","author":"A Roberts","year":"2018","edition":"2"},{"issue":"7012","key":"pone.0245760.ref004","doi-asserted-by":"crossref","first-page":"1055","DOI":"10.1038\/nature02999","article-title":"A new small-bodied hominin from the Late Pleistocene of Flores, Indonesia","volume":"431","author":"P Brown","year":"2004","journal-title":"Nature"},{"issue":"7061","key":"pone.0245760.ref005","doi-asserted-by":"crossref","first-page":"1012","DOI":"10.1038\/nature04022","article-title":"Further evidence for small-bodied hominins from the Late Pleistocene of Flores, Indonesia","volume":"437","author":"MJ Morwood","year":"2005","journal-title":"Nature"},{"issue":"12","key":"pone.0245760.ref006","doi-asserted-by":"crossref","first-page":"4400","DOI":"10.1016\/j.jas.2013.06.028","article-title":"Faces of Homo floresiensis (LB1)","volume":"40","author":"S Hayes","year":"2013","journal-title":"Journal of Archaeological Science"},{"key":"pone.0245760.ref007","unstructured":"Anderson K. Hominin Representations in Museum Displays: Their role in forming public understanding through the non-verbal communication of science: Ph.D. Dissertation, The University of Adelaide, Adelaide; 2011."},{"key":"pone.0245760.ref008","doi-asserted-by":"crossref","DOI":"10.3389\/fevo.2021.639048","article-title":"Visual depictions of our evolutionary past: A broad case study concerning the need for quantitative methods of soft tissue reconstruction and art-science collaborations","volume":"9","author":"RM Campbell","year":"2021","journal-title":"Frontiers in Ecology and Evolution"},{"issue":"1","key":"pone.0245760.ref009","doi-asserted-by":"crossref","first-page":"48","DOI":"10.1002\/ajpa.10166","article-title":"Facial approximation: An evaluation of mouth-width determination","volume":"121","author":"CN Stephan","year":"2003","journal-title":"American Journal of Physical Anthropology"},{"issue":"5","key":"pone.0245760.ref010","doi-asserted-by":"crossref","first-page":"1305","DOI":"10.1520\/JFS11181J","article-title":"The lateral craniographic method of facial reconstruction","volume":"32","author":"RM George","year":"1987","journal-title":"Journal of Forensic Sciences"},{"volume-title":"Schiller\u2019s Sch\u00e4del und Todenmaske nebst Mittheilungen \u00fcber Sch\u00e4del und Todenmaske Kants","year":"1883","author":"H Welcker","key":"pone.0245760.ref011"},{"volume-title":"The Face Finder","year":"1971","author":"M Gerasimov","key":"pone.0245760.ref012"},{"volume-title":"The Reconstruction of the Face on the Skull","year":"1955","author":"M Gerasimov","key":"pone.0245760.ref013"},{"volume-title":"Making Faces: Using Forensic and Archaeological Evidence","year":"1997","author":"J Prag","key":"pone.0245760.ref014"},{"key":"pone.0245760.ref015","doi-asserted-by":"crossref","DOI":"10.1017\/CBO9781107340961","volume-title":"Forensic Facial Reconstruction","author":"C Wilkinson","year":"2004"},{"key":"pone.0245760.ref016","doi-asserted-by":"crossref","DOI":"10.1016\/j.forsciint.2015.04.015","article-title":"Soft tissue thickness values for black and coloured South African children aged 6\u201313 years","volume":"252","author":"N Briers","year":"2015","journal-title":"Forensic Science International"},{"key":"pone.0245760.ref017","doi-asserted-by":"crossref","first-page":"113","DOI":"10.1016\/j.forsciint.2017.09.016","article-title":"2018 tallied facial soft tissue thicknesses for adults and sub-adults","volume":"280","author":"CN Stephan","year":"2017","journal-title":"Forensic Science International"},{"key":"pone.0245760.ref018","unstructured":"Hanebrink JR. Datum is only skin deep: in vivo measurements of facial tissue thickness in chimpanzees [Ph.D. Dissertation]: Ph.D. Dissertation, Louisiana State University; 2006."},{"issue":"Suppl. 1","key":"pone.0245760.ref019","doi-asserted-by":"crossref","first-page":"S126","DOI":"10.1016\/j.forsciint.2006.02.034","article-title":"Large-scale in-vivo Caucasian soft tissue thickness database for craniofacial reconstruction","volume":"159","author":"S Greef","year":"2006","journal-title":"Forensic science international"},{"issue":"2","key":"pone.0245760.ref020","doi-asserted-by":"crossref","first-page":"303","DOI":"10.1002\/ajpa.1330300215","article-title":"Bizygomatic diameter: The thickness of the soft tissues over the zygions","volume":"30","author":"PRN Sutton","year":"1969","journal-title":"American Journal of Physical Anthropology"},{"issue":"2","key":"pone.0245760.ref021","doi-asserted-by":"crossref","first-page":"121","DOI":"10.1002\/ajpa.10073","article-title":"Variation in soft-tissue thicknesses on the human face and their relation to craniometric dimensions","volume":"118","author":"E Simpson","year":"2002","journal-title":"American Journal of Physical Anthropology"},{"key":"pone.0245760.ref022","doi-asserted-by":"crossref","first-page":"128","DOI":"10.1016\/j.forsciint.2018.03.011","article-title":"Facial soft tissue thickness (FSTT) estimation models\u2014And the strength of correlations between craniometric dimensions and FSTTs","volume":"286","author":"CN Stephan","year":"2018","journal-title":"Forensic Science International"},{"key":"pone.0245760.ref023","doi-asserted-by":"crossref","first-page":"117","DOI":"10.1007\/978-3-642-19953-0_12","volume-title":"New Challenges for Intelligent Information and Database Systems","author":"QH Dinh","year":"2011"},{"issue":"1","key":"pone.0245760.ref024","doi-asserted-by":"crossref","first-page":"608","DOI":"10.1038\/s41598-017-00548-3","article-title":"Bonobo anatomy reveals stasis and mosaicism in chimpanzee evolution, and supports bonobos as the most appropriate extant model for the common ancestor of chimpanzees and humans","volume":"7","author":"R Diogo","year":"2017","journal-title":"Scientific Reports"},{"volume-title":"Photographic and descriptive musculoskeletal atlas of bonobos\u2014with notes on the weight, attachments, variations, and innervation of the muscles and comparisons with common chimpanzees and humans","year":"2017","author":"R Diogo","key":"pone.0245760.ref025"},{"journal-title":"Photographic and Descriptive Musculoskeletal Atlas of Chimpanzees (Pan)","year":"2013","author":"R Diogo","key":"pone.0245760.ref026"},{"issue":"3","key":"pone.0245760.ref027","doi-asserted-by":"crossref","first-page":"205","DOI":"10.1007\/s10278-004-1014-6","article-title":"OsiriX: An open-source software for navigating in multidimensional DICOM images","volume":"17","author":"A Rosset","year":"2004","journal-title":"Journal of Digital Imaging"},{"issue":"2","key":"pone.0245760.ref028","doi-asserted-by":"crossref","first-page":"285","DOI":"10.1002\/jcsm.12158","article-title":"A comparative study of software programmes for cross-sectional skeletal muscle and adipose tissue measurements on abdominal computed tomography scans of rectal cancer patients","volume":"8","author":"JLA van Vugt","year":"2017","journal-title":"Journal of Cachexia, Sarcopenia and Muscle"},{"issue":"6","key":"pone.0245760.ref029","doi-asserted-by":"crossref","first-page":"1257","DOI":"10.1111\/j.1556-4029.2008.00852.x","article-title":"Facial soft tissue depths in craniofacial identification (part I): An analytical review of the published adult data","volume":"53","author":"CN Stephan","year":"2008","journal-title":"Journal of Forensic Sciences"},{"issue":"3","key":"pone.0245760.ref030","doi-asserted-by":"crossref","first-page":"863","DOI":"10.1007\/s00414-015-1292-1","article-title":"A standardized nomenclature for craniofacial and facial anthropometry","volume":"130","author":"J Caple","year":"2016","journal-title":"International Journal of Legal Medicine"},{"volume-title":"The Human Skeleton in Forensic Medicine","year":"2013","author":"MY Iscan","key":"pone.0245760.ref031"},{"key":"pone.0245760.ref032","doi-asserted-by":"crossref","first-page":"109965","DOI":"10.1016\/j.forsciint.2019.109965","article-title":"Facial soft tissue thicknesses in craniofacial identification: Data collection protocols and associated measurement errors","volume":"304","author":"CN Stephan","year":"2019","journal-title":"Forensic Science International"},{"key":"pone.0245760.ref033","doi-asserted-by":"crossref","DOI":"10.1525\/9780520351288","volume-title":"Bonobo: The forgotten ape","author":"F de Waal","year":"1997"},{"issue":"7","key":"pone.0245760.ref034","doi-asserted-by":"crossref","first-page":"2124","DOI":"10.1073\/pnas.0808730106","article-title":"The feeding biomechanics and dietary ecology of Australopithecus africanus","volume":"106","author":"DS Strait","year":"2009","journal-title":"Proceedings of the National Academy of Sciences USA"},{"issue":"2","key":"pone.0245760.ref035","doi-asserted-by":"crossref","first-page":"324","DOI":"10.1002\/ajpa.22225","article-title":"Individual tooth macrowear pattern guides the reconstruction of Sts 52 (Australopithecus africanus) dental arches","volume":"150","author":"S Benazzi","year":"2013","journal-title":"American Journal of Physical Anthropology"},{"issue":"1","key":"pone.0245760.ref036","doi-asserted-by":"crossref","first-page":"75","DOI":"10.1016\/j.jhevol.2011.02.005","article-title":"A new OH5 reconstruction with an assessment of its uncertainty","volume":"61","author":"S Benazzi","year":"2011","journal-title":"Journal of Human Evolution"},{"issue":"2","key":"pone.0245760.ref037","doi-asserted-by":"crossref","first-page":"107","DOI":"10.1080\/00359190509520486","article-title":"Dating of the Sterkfontein hominids: progress and possibilities","volume":"60","author":"TC Partridge","year":"2005","journal-title":"Transactions of the Royal Society of South Africa"},{"issue":"4046","key":"pone.0245760.ref038","doi-asserted-by":"crossref","first-page":"672","DOI":"10.1038\/159672a0","article-title":"Discovery of a new skull of the South African ape-man","volume":"159","author":"R Broom","year":"1947","journal-title":"Plesianthropus. Nature"},{"issue":"6031","key":"pone.0245760.ref039","doi-asserted-by":"crossref","first-page":"788","DOI":"10.1038\/316788a0","article-title":"Early Homo erectus skeleton from west Lake Turkana, Kenya","volume":"316","author":"F Brown","year":"1985","journal-title":"Nature"},{"key":"pone.0245760.ref040","doi-asserted-by":"crossref","first-page":"102634","DOI":"10.1016\/j.jhevol.2019.06.005","article-title":"The skull of StW 573, a 3.67 Ma Australopithecus prometheus skeleton from Sterkfontein Caves, South Africa","volume":"134","author":"RJ Clarke","year":"2019","journal-title":"Journal of Human Evolution"},{"issue":"1556","key":"pone.0245760.ref041","doi-asserted-by":"crossref","first-page":"3365","DOI":"10.1098\/rstb.2010.0070","article-title":"The cranial base of Australopithecus afarensis: new insights from the female skull","volume":"365","author":"WH Kimbel","year":"2010","journal-title":"Philosophical Transactions of the Royal Society of London: Bioligical Sciences"},{"key":"pone.0245760.ref042","doi-asserted-by":"crossref","DOI":"10.1016\/B978-0-12-576280-9.50006-7","volume-title":"The Australopithecine Face","author":"Y Rak","year":"1983"},{"issue":"1","key":"pone.0245760.ref043","doi-asserted-by":"crossref","first-page":"97","DOI":"10.1002\/(SICI)1096-8644(199901)108:1<97::AID-AJPA6>3.0.CO;2-O","article-title":"Sexual dimorphism in the face of Australopithecus africanus","volume":"108","author":"CA Lockwood","year":"1999","journal-title":"American Journal of Physical Anthropology"},{"key":"pone.0245760.ref044","doi-asserted-by":"crossref","DOI":"10.17159\/sajs.2018\/a0249","article-title":"The cranium of Sts 5 (\u2018Mrs Ples\u2019) in relation to sexual dimorphism of Australopithecus africanus","volume":"114","author":"G Tawane","year":"2018","journal-title":"South African Journal of Science"},{"key":"pone.0245760.ref045","doi-asserted-by":"crossref","first-page":"533","DOI":"10.1016\/j.jhevol.2009.11.005","article-title":"Temporal lines and open sutures revealed on cranial bone adhering to matrix associated with Sts 5 (\u201cMrs Ples\u201d), Sterkfontein, South Africa","volume":"58","author":"S Potze","year":"2010","journal-title":"Journal of human evolution"},{"key":"pone.0245760.ref046","doi-asserted-by":"crossref","first-page":"1707","DOI":"10.1002\/ar.20795","article-title":"Revisiting the developmental stage and age-at-death of the \u201cMrs. Ples\u201d (Sts 5) and Sts 14 specimens from Sterkfontein (South Africa): Do they belong to the same individual?","volume":"291","author":"A Bonmat\u00ed","year":"2008","journal-title":"Anatomical Record"},{"issue":"7","key":"pone.0245760.ref047","doi-asserted-by":"crossref","first-page":"4289","DOI":"10.1002\/ece3.5050","article-title":"3D printing: Applications in evolution and ecology","volume":"9","author":"M Walker","year":"2019","journal-title":"Ecol Evol"},{"key":"pone.0245760.ref048","doi-asserted-by":"crossref","first-page":"513","DOI":"10.1520\/JFS2004251","article-title":"Does sexual dimorphism in facial soft tissue depths justify sex distinction in craniofacial identification?","volume":"50","author":"C Stephan","year":"2005","journal-title":"Journal of forensic sciences"},{"issue":"5","key":"pone.0245760.ref049","doi-asserted-by":"crossref","first-page":"471","DOI":"10.1016\/S0940-9602(04)80086-4","article-title":"Craniofacial sexual dimorphism patterns and allometry among extant hominids","volume":"186","author":"K Schaefer","year":"2004","journal-title":"Annals of Anatomy\u2014Anatomischer Anzeiger"},{"issue":"3","key":"pone.0245760.ref050","doi-asserted-by":"crossref","first-page":"183","DOI":"10.1006\/jhev.1993.1014","article-title":"Sexual dimorphism in hominoids: Further studies of craniofacial shape differences in Pan, Gorilla, and Pongo","volume":"24","author":"P O\u2019Higgins","year":"1993","journal-title":"Journal of Human Evolution"},{"key":"pone.0245760.ref051","doi-asserted-by":"crossref","first-page":"72","DOI":"10.1016\/j.jhevol.2019.02.006","article-title":"Brain size growth in Australopithecus","volume":"130","author":"Z Cofran","year":"2019","journal-title":"Journal of Human Evolution"},{"issue":"14","key":"pone.0245760.ref052","doi-asserted-by":"crossref","DOI":"10.1126\/sciadv.aaz4729","article-title":"Australopithecus afarensis endocasts suggest ape-like brain organization and prolonged brain growth","volume":"6","author":"P Gunz","year":"2020","journal-title":"Science Advances"},{"key":"pone.0245760.ref053","doi-asserted-by":"crossref","first-page":"94","DOI":"10.1016\/j.jhevol.2017.03.011","article-title":"Australopithecus sediba and the emergence of Homo: Questionable evidence from the cranium of the juvenile holotype MH 1","volume":"107","author":"WH Kimbel","year":"2017","journal-title":"Journal of Human Evolution"},{"key":"pone.0245760.ref054","first-page":"345","article-title":"A study of man embracing error","volume":"49","author":"MFA Montagu","year":"1947","journal-title":"Technology Review"},{"key":"pone.0245760.ref055","doi-asserted-by":"crossref","unstructured":"Diogo, Potau JM, Pastor, Paz D, Ferrero EM, Bello, et al. Photographic and Descriptive Musculoskeletal Atlas of Orangutans2013.","DOI":"10.1201\/b14592"},{"key":"pone.0245760.ref056","doi-asserted-by":"crossref","DOI":"10.1201\/b10299","author":"R Diogo","year":"2010","journal-title":"Photographic and Descriptive Musculoskeletal Atlas of Gorilla"},{"volume-title":"An Introduction to Human Evolutionary Anatomy","year":"1990","author":"L Aiello","key":"pone.0245760.ref057"},{"issue":"2","key":"pone.0245760.ref058","first-page":"97","article-title":"Mikhail Mikhaylovich Gerasimov\u2019s authentic approach to plastic facial reconstruction","volume":"52","author":"H Ullrich","year":"2016","journal-title":"Anthropologie"},{"key":"pone.0245760.ref059","first-page":"329","article-title":"Die Persistenz der Rassen und die Reconstruction der Physiognomie pr\u00e4historischer Sch\u00e4del","volume":"25","author":"J Kollman","year":"1898","journal-title":"Archiv f\u00fcr Anthropologie"},{"key":"pone.0245760.ref060","unstructured":"Kern MW. Regional Phylogenetic Analysis of the Myology of Primates and Correlates in the Human Skeleton [Masters Thesis]. Madison: Masters Thesis, University of Wisconsin; 2012."},{"key":"pone.0245760.ref061","doi-asserted-by":"crossref","first-page":"47","DOI":"10.1016\/j.jhevol.2017.07.014","article-title":"A volumetric technique for fossil body mass estimation applied to Australopithecus afarensis","volume":"115","author":"CA Brassey","year":"2018","journal-title":"Journal of Human Evolution"},{"issue":"1","key":"pone.0245760.ref062","doi-asserted-by":"crossref","first-page":"438","DOI":"10.1038\/s41598-019-57247-4","article-title":"Corruption of the Pearson correlation coefficient by measurement error and its estimation, bias, and correction under different error models","volume":"10","author":"E Saccenti","year":"2020","journal-title":"Scientific Reports"},{"issue":"5","key":"pone.0245760.ref063","doi-asserted-by":"crossref","first-page":"1271","DOI":"10.1111\/j.1556-4029.2012.02075.x","article-title":"Anatomical placement of the human eyeball in the orbit\u2014Validation using CT scans of living adults and prediction for facial approximation","volume":"57","author":"P Guyomarc\u2019h","year":"2012","journal-title":"Journal of Forensic Sciences"},{"issue":"1","key":"pone.0245760.ref064","doi-asserted-by":"crossref","first-page":"29","DOI":"10.1016\/S0379-0738(02)00279-7","article-title":"Position of superciliare in relation to the lateral iris: testing a suggested facial approximation guideline","volume":"130","author":"CN Stephan","year":"2002","journal-title":"Forensic Science International"},{"key":"pone.0245760.ref065","first-page":"59","article-title":"Studies in eye brows among the Bengalees","volume":"6","author":"RN Basu","year":"1941","journal-title":"University of Calcutta Anthropological Papers"},{"key":"pone.0245760.ref066","first-page":"245","article-title":"The position of the eyebrow","volume":"21","author":"JH Oestreicher","year":"1990","journal-title":"Ophthalmic Surgery"},{"volume-title":"Forensic Analysis of the Skull: Craniofacial Analysis, Reconstruction, and Identification","year":"1993","author":"BA Fedosyutkin","key":"pone.0245760.ref067"},{"key":"pone.0245760.ref068","doi-asserted-by":"crossref","first-page":"364","DOI":"10.1002\/ajpa.20337","article-title":"Appraisal of traditional and recently proposed relationships between the hard and soft nose in profile","volume":"130","author":"C Rynn","year":"2006","journal-title":"American journal of physical anthropology"},{"key":"pone.0245760.ref069","doi-asserted-by":"crossref","first-page":"1305","DOI":"10.1520\/JFS11181J","article-title":"The lateral craniographic method of facial reconstruction","volume":"32","author":"R George","year":"1987","journal-title":"Journal of Forensic Sciences"},{"key":"pone.0245760.ref070","doi-asserted-by":"crossref","first-page":"1391","DOI":"10.1520\/JFS11917J","article-title":"An appraisal of plastic reconstruction of the external Nose","volume":"31","author":"G Macho","year":"1986","journal-title":"Journal of forensic sciences"},{"key":"pone.0245760.ref071","first-page":"1","article-title":"Reconstructing the shape of the nose according to the skull","volume":"4","author":"M Prokopec","year":"2002","journal-title":"Forensic Science Communications"},{"issue":"3","key":"pone.0245760.ref072","doi-asserted-by":"crossref","first-page":"240","DOI":"10.1002\/ajpa.10300","article-title":"Predicting nose projection and pronasale position in facial approximation: A test of published methods and proposal of new guidelines","volume":"122","author":"CN Stephan","year":"2003","journal-title":"American Journal of Physical Anthropology"},{"key":"pone.0245760.ref073","doi-asserted-by":"crossref","first-page":"725","DOI":"10.1520\/JFS2002418","article-title":"Predicting mouth width from inter-canine width\u2014A 75% rule","volume":"48","author":"C Stephan","year":"2003","journal-title":"Journal of forensic sciences"},{"key":"pone.0245760.ref074","doi-asserted-by":"crossref","first-page":"1343","DOI":"10.1097\/01.prs.0000279558.86727.5a","article-title":"Location of the infraorbital and mental foramen with reference to the soft-tissue landmarks","volume":"120","author":"W-C Song","year":"2007","journal-title":"Plast Reconstr Surg"},{"key":"pone.0245760.ref075","first-page":"2","article-title":"Mouth width prediction in craniofacial identification: Cadaver tests of four recent methods, including two techniques for edentulous skulls","volume":"26","author":"C Stephan","year":"2008","journal-title":"The Journal of Forensic Odonto-Stomatology"},{"issue":"1","key":"pone.0245760.ref076","doi-asserted-by":"crossref","first-page":"1","DOI":"10.1097\/00006534-195707000-00001","article-title":"Artistic relationships in surface anatomy of the face: application to reconstructive surgery","volume":"20","author":"TR Broadbend","year":"1957","journal-title":"Plast Reconstr Surg"},{"key":"pone.0245760.ref077","doi-asserted-by":"crossref","first-page":"1427","DOI":"10.1111\/j.1556-4029.2012.02181.x","article-title":"The validity of ear prediction guidelines used in facial approximation","volume":"57","author":"P Guyomarc\u2019h","year":"2012","journal-title":"Journal of forensic sciences"},{"issue":"4685","key":"pone.0245760.ref078","doi-asserted-by":"crossref","first-page":"491","DOI":"10.1038\/184491a0","article-title":"A new fossil skull from olduvai","volume":"184","author":"LSB Leakey","year":"1959","journal-title":"Nature"},{"key":"pone.0245760.ref079","first-page":"189","article-title":"The Drimolen skull: The most complete australopithecine cranium and mandible to date","volume":"96","author":"AW Keyser","year":"2000","journal-title":"South African Journal of Science"},{"key":"pone.0245760.ref080","doi-asserted-by":"crossref","first-page":"195","DOI":"10.1038\/115195a0","article-title":"Australopithecus africanus: The man-ape of South Africa","volume":"115","author":"RA Dart","year":"1925","journal-title":"Nature"},{"issue":"7109","key":"pone.0245760.ref081","doi-asserted-by":"crossref","first-page":"296","DOI":"10.1038\/nature05047","article-title":"A juvenile early hominin skeleton from Dikika, Ethiopia","volume":"443","author":"Z Alemseged","year":"2006","journal-title":"Nature"},{"issue":"6","key":"pone.0245760.ref082","doi-asserted-by":"crossref","first-page":"1641","DOI":"10.1111\/1556-4029.13768","article-title":"Growth patterns and individual variation in mid-sagittal facial soft tissue depth from childhood to adulthood","volume":"63","author":"MR McKinnon","year":"2018","journal-title":"Journal of Forensic Sciences"}],"container-title":["PLOS 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