{"status":"ok","message-type":"work","message-version":"1.0.0","message":{"indexed":{"date-parts":[[2026,8,2]],"date-time":"2026-08-02T15:19:30Z","timestamp":1785683970335,"version":"3.56.0"},"reference-count":100,"publisher":"Public Library of Science (PLoS)","issue":"4","license":[{"start":{"date-parts":[[2021,4,15]],"date-time":"2021-04-15T00:00:00Z","timestamp":1618444800000},"content-version":"vor","delay-in-days":0,"URL":"http:\/\/creativecommons.org\/licenses\/by\/4.0\/"}],"funder":[{"name":"Ministry of Science and ICT","award":["IITP-2019-2017-0-01630"],"award-info":[{"award-number":["IITP-2019-2017-0-01630"]}]},{"DOI":"10.13039\/501100000837","name":"University of Nottingham","doi-asserted-by":"crossref","award":["Beacon Anne McLaren Research Fellowship"],"award-info":[{"award-number":["Beacon Anne McLaren Research Fellowship"]}],"id":[{"id":"10.13039\/501100000837","id-type":"DOI","asserted-by":"crossref"}]}],"content-domain":{"domain":["www.plosone.org"],"crossmark-restriction":false},"short-container-title":["PLoS ONE"],"abstract":"<jats:p>When people confuse and reject a non-word that is created by switching two adjacent letters from an actual word, is called the transposition confusability effect (TCE). The TCE is known to occur at the very early stages of visual word recognition with such unit exchange as letters or syllables, but little is known about the brain mechanisms of TCE. In this study, we examined the neural correlates of TCE and the effect of a morpheme boundary placement on TCE. We manipulated the placement of a morpheme boundary by exchanging places of two syllables embedded in Korean morphologically complex words made up of lexical morpheme and grammatical morpheme. In the two experimental conditions, the transposition syllable within-boundary condition (TSW) involved exchanging two syllables within the same morpheme, whereas the across-boundary condition (TSA) involved the exchange of syllables across the stem and grammatical morpheme boundary. During fMRI, participants performed the lexical decision task. Behavioral results revealed that the TCE was found in TSW condition, and the morpheme boundary, which is manipulated in TSA, modulated the TCE. In the fMRI results, TCE induced activation in the left inferior parietal lobe (IPL) and intraparietal sulcus (IPS). The IPS activation was specific to a TCE and its strength of activation was associated with task performance. Furthermore, two functional networks were involved in the TCE: the central executive network and the dorsal attention network. Morpheme boundary modulation suppressed the TCE by recruiting the prefrontal and temporal regions, which are the key regions involved in semantic processing. Our findings propose the role of the dorsal visual pathway in syllable position processing and that its interaction with other higher cognitive systems is modulated by the morphological boundary in the early phases of visual word recognition.<\/jats:p>","DOI":"10.1371\/journal.pone.0249111","type":"journal-article","created":{"date-parts":[[2021,4,15]],"date-time":"2021-04-15T18:06:03Z","timestamp":1618509963000},"page":"e0249111","update-policy":"https:\/\/doi.org\/10.1371\/journal.pone.corrections_policy","source":"Crossref","is-referenced-by-count":8,"title":["Neural correlates of confusability in recognition of morphologically complex Korean words"],"prefix":"10.1371","volume":"16","author":[{"ORCID":"https:\/\/orcid.org\/0000-0003-1289-2424","authenticated-orcid":true,"given":"Jeahong","family":"Kim","sequence":"first","affiliation":[],"role":[{"vocabulary":"crossref","role":"author"}]},{"given":"JeYoung","family":"Jung","sequence":"additional","affiliation":[],"role":[{"vocabulary":"crossref","role":"author"}]},{"given":"Kichun","family":"Nam","sequence":"additional","affiliation":[],"role":[{"vocabulary":"crossref","role":"author"}]}],"member":"340","published-online":{"date-parts":[[2021,4,15]]},"reference":[{"issue":"2","key":"pone.0249111.ref001","doi-asserted-by":"crossref","first-page":"225","DOI":"10.1016\/S0022-5371(79)90136-1","article-title":"Letter and order information in lexical access","volume":"18","author":"SM Chambers","year":"1979","journal-title":"Journal of Verbal Learning and Verbal Behavior"},{"issue":"2","key":"pone.0249111.ref002","doi-asserted-by":"crossref","first-page":"189","DOI":"10.1016\/j.jml.2006.11.002","article-title":"SIHGT and SUNOD: The role of orthography and phonology in the perception of transposed letter anagrams","volume":"56","author":"C Frankish","year":"2007","journal-title":"Journal of Memory and Language"},{"issue":"1","key":"pone.0249111.ref003","doi-asserted-by":"crossref","first-page":"78","DOI":"10.3758\/BF03196953","article-title":"Criterion bias and search sequence bias in word recognition","volume":"9","author":"RE O\u2019Connor","year":"1981","journal-title":"Memory & Cognition"},{"issue":"4","key":"pone.0249111.ref004","doi-asserted-by":"crossref","first-page":"308","DOI":"10.1027\/1618-3169.53.4.308","article-title":"Do transposed-letter similarity effects occur at a syllable level?","volume":"53","author":"M Perea","year":"2006","journal-title":"Experimental Psychology"},{"issue":"7","key":"pone.0249111.ref005","doi-asserted-by":"crossref","first-page":"917","DOI":"10.1080\/01690965.2012.679662","article-title":"Morphological decomposition and the transposed-letter (TL) position effect","volume":"28","author":"M Taft","year":"2013","journal-title":"Language and Cognitive Processes"},{"issue":"6","key":"pone.0249111.ref006","doi-asserted-by":"crossref","first-page":"775","DOI":"10.1006\/jmla.1996.0040","article-title":"Lexical retrieval and selection processes: Effects of transposed-letter confusability","volume":"35","author":"S Andrews","year":"1996","journal-title":"Journal of Memory and Language"},{"issue":"2","key":"pone.0249111.ref007","doi-asserted-by":"crossref","first-page":"203","DOI":"10.1023\/A:1023202116609","article-title":"The pros and cons of masked priming","volume":"27","author":"KI Forster","year":"1998","journal-title":"Journal of Psycholinguistic Research"},{"issue":"6","key":"pone.0249111.ref008","doi-asserted-by":"crossref","first-page":"829","DOI":"10.3758\/BF03196438","article-title":"Does jugde activate COURT? Transposed-letter similarity effects in masked associative priming","volume":"31","author":"M Perea","year":"2003","journal-title":"Memory & Cognition"},{"issue":"2","key":"pone.0249111.ref009","doi-asserted-by":"crossref","first-page":"231","DOI":"10.1016\/j.jml.2004.05.005","article-title":"Can CANISO activate CASINO? Transposed-letter similarity effects with nonadjacent letter positions","volume":"51","author":"M Perea","year":"2004","journal-title":"Journal of Memory and Language"},{"issue":"3","key":"pone.0249111.ref010","doi-asserted-by":"crossref","first-page":"333","DOI":"10.1080\/01690960344000198","article-title":"Letter position coding in printed word perception: Effects of repeated and transposed letters","volume":"19","author":"S Schoonbaert","year":"2004","journal-title":"Language and Cognitive Processes"},{"key":"pone.0249111.ref011","doi-asserted-by":"crossref","first-page":"69","DOI":"10.1080\/01690960701578146","article-title":"Do orthotactics and phonology constrain the transposed-letter effect?","volume":"23","author":"M Perea","year":"2008","journal-title":"Language and Cognitive Processes"},{"key":"pone.0249111.ref012","doi-asserted-by":"crossref","first-page":"52","DOI":"10.1111\/bjop.12469","article-title":"Does orthographic processing emerge rapidly after learning a new script?","volume":"112","author":"M Fern\u00e1ndez\u2010L\u00f3pez","year":"2021","journal-title":"British Journal of Psychology"},{"issue":"7","key":"pone.0249111.ref013","doi-asserted-by":"crossref","first-page":"1610","DOI":"10.1162\/jocn_a_00180","article-title":"Differential sensitivity of letters, numbers, and symbols to character transpositions","volume":"24","author":"JA Du\u00f1abeitia","year":"2009","journal-title":"Journal of Cognitive Neuroscience"},{"issue":"6","key":"pone.0249111.ref014","doi-asserted-by":"crossref","first-page":"1344","DOI":"10.1177\/1747021818789876","article-title":"Can letter position encoding be modified by visual perceptual elements?","volume":"72","author":"A Marcet","year":"2019","journal-title":"Quarterly Journal of Experimental Psychology"},{"issue":"7","key":"pone.0249111.ref015","doi-asserted-by":"crossref","first-page":"e68460","DOI":"10.1371\/journal.pone.0068460","article-title":"Evidence for letter-specific position coding mechanisms","volume":"8","author":"S Massol","year":"2013","journal-title":"PLoS One"},{"key":"pone.0249111.ref016","first-page":"1","volume-title":"Mental lexicon: \u201cSome words to talk about words\u201d","author":"J Grainger","year":"2003"},{"issue":"3","key":"pone.0249111.ref017","doi-asserted-by":"crossref","first-page":"713","DOI":"10.1037\/a0019738","article-title":"The spatial coding model of visual word identification","volume":"117","author":"CJ Davis","year":"2010","journal-title":"Psychological Review"},{"issue":"5","key":"pone.0249111.ref018","first-page":"923","article-title":"What do letter migration errors reveal about letter position coding in visual word recognition?","volume":"30","author":"CJ Davis","year":"2004","journal-title":"Journal of Experimental Psychology: Human Perception and Performance"},{"issue":"3","key":"pone.0249111.ref019","first-page":"535","article-title":"Contrasting five different theories of letter position coding: Evidence from orthographic similarity effects","volume":"32","author":"CJ Davis","year":"2006","journal-title":"Journal of Experimental Psychology: Human Perception and Performance"},{"issue":"3","key":"pone.0249111.ref020","doi-asserted-by":"crossref","first-page":"577","DOI":"10.1037\/a0012667","article-title":"The overlap model: A model of letter position coding","volume":"115","author":"P Gomez","year":"2008","journal-title":"Psychological Review"},{"issue":"4","key":"pone.0249111.ref021","doi-asserted-by":"crossref","first-page":"530","DOI":"10.1016\/j.jml.2009.01.002","article-title":"Are onsets and codas important in processing letter position? A comparison of TL effects in English and Korean","volume":"60","author":"CH Lee","year":"2009","journal-title":"Journal of Memory and Language"},{"issue":"1","key":"pone.0249111.ref022","doi-asserted-by":"crossref","first-page":"129","DOI":"10.3758\/s13423-010-0028-y","article-title":"Subsyllabic structure reflected in letter confusability effects in Korean word recognition","volume":"18","author":"CH Lee","year":"2011","journal-title":"Psychonomic Bulletin & Review"},{"issue":"4","key":"pone.0249111.ref023","first-page":"458","article-title":"No flexibility in letter position coding in Korean","volume":"45","author":"K Rastle","year":"2019","journal-title":"Journal of Experimental Psychology: Human Perception and Performance"},{"issue":"3","key":"pone.0249111.ref024","doi-asserted-by":"crossref","first-page":"309","DOI":"10.1007\/s10936-015-9353-7","article-title":"Syllable transposition effects in Korean word recognition","volume":"44","author":"CH Lee","year":"2015","journal-title":"Journal of Psycholinguistic Research"},{"issue":"3","key":"pone.0249111.ref025","doi-asserted-by":"crossref","first-page":"261","DOI":"10.22172\/cogbio.2018.30.3.004","article-title":"Syllable transposition effect on processing the morphologically complex Korean noun Eojeol","volume":"30","author":"J Kim","year":"2018","journal-title":"The Korean Journal of Cognitive and Biological Psychology"},{"issue":"4","key":"pone.0249111.ref026","doi-asserted-by":"crossref","first-page":"498","DOI":"10.1006\/jmla.1998.2625","article-title":"Positional specificity of radicals in Chinese character recognition","volume":"40","author":"M Taft","year":"1999","journal-title":"Journal of Memory and Language"},{"issue":"2","key":"pone.0249111.ref027","doi-asserted-by":"crossref","first-page":"e0149583","DOI":"10.1371\/journal.pone.0149583","article-title":"Functional anatomy of recognition of Chinese multi-character words: Convergent evidence from effects of transposable nonwords, lexicality, and word frequency","volume":"11","author":"N Lin","year":"2016","journal-title":"PloS One"},{"issue":"1","key":"pone.0249111.ref028","doi-asserted-by":"crossref","first-page":"67","DOI":"10.1080\/01690960802053924","article-title":"Beyond alphabetic orthographies: The role of form and phonology in transposition effects in Katakana","volume":"24","author":"M Perea","year":"2009","journal-title":"Language and Cognitive Processes"},{"issue":"3","key":"pone.0249111.ref029","first-page":"914","article-title":"Transposed letter priming with horizontal and vertical text in Japanese and English readers","volume":"37","author":"N Witzel","year":"2011","journal-title":"Journal of Experimental Psychology: Human Perception and Performance"},{"issue":"2","key":"pone.0249111.ref030","first-page":"510","article-title":"Seeing stems everywhere: Position-independent identification of stem morphemes","volume":"39","author":"D Crepaldi","year":"2013","journal-title":"Journal of Experimental Psychology: Human Perception and Performance"},{"issue":"6","key":"pone.0249111.ref031","first-page":"1327","article-title":"Letter transpositions within and across morphemes","volume":"31","author":"K Christianson","year":"2005","journal-title":"Journal of Experimental Psychology: Learning, Memory, and Cognition"},{"issue":"3","key":"pone.0249111.ref032","doi-asserted-by":"crossref","first-page":"691","DOI":"10.1016\/j.cognition.2006.12.001","article-title":"Do transposed-letter similarity effects occur at a morpheme level? Evidence for morpho-orthographic decomposition","volume":"105","author":"JA Du\u00f1abeitia","year":"2007","journal-title":"Cognition"},{"issue":"6","key":"pone.0249111.ref033","doi-asserted-by":"crossref","first-page":"1557","DOI":"10.3758\/s13423-014-0609-2","article-title":"Revisiting letter transpositions within and across morphemic boundaries","volume":"21","author":"JA Du\u00f1abeitia","year":"2014","journal-title":"Psychonomic Bulletin & Review"},{"issue":"7\u20138","key":"pone.0249111.ref034","doi-asserted-by":"crossref","first-page":"942","DOI":"10.1080\/01690960802069730","article-title":"Morphological decomposition based on the analysis of orthography","volume":"23","author":"K Rastle","year":"2008","journal-title":"Language and Cognitive Processes"},{"issue":"6","key":"pone.0249111.ref035","doi-asserted-by":"crossref","first-page":"1090","DOI":"10.3758\/BF03196742","article-title":"The broth in my brother\u2019s brothel: Morpho-orthographic segmentation in visual word recognition","volume":"11","author":"K Rastle","year":"2004","journal-title":"Psychonomic Bulletin & Review"},{"issue":"6","key":"pone.0249111.ref036","doi-asserted-by":"crossref","first-page":"895","DOI":"10.3758\/MC.37.6.895","article-title":"Semantic transparency and masked morphological priming: The case of prefixed words","volume":"37","author":"K Diependaele","year":"2009","journal-title":"Memory & Cognition"},{"issue":"7","key":"pone.0249111.ref037","doi-asserted-by":"crossref","first-page":"1610","DOI":"10.1162\/jocn_a_00180","article-title":"Differential sensitivity of letters, numbers, and symbols to character transpositions","volume":"24","author":"JA Du\u00f1abeitia","year":"2012","journal-title":"Journal of Cognitive Neuroscience"},{"issue":"9","key":"pone.0249111.ref038","doi-asserted-by":"crossref","first-page":"1798","DOI":"10.1080\/17470218.2012.672437","article-title":"Parallel processing of whole words and morphemes in visual word recognition","volume":"65","author":"E Beyersmann","year":"2012","journal-title":"The Quarterly Journal of Experimental Psychology"},{"key":"pone.0249111.ref039","doi-asserted-by":"crossref","first-page":"895","DOI":"10.3758\/MC.37.6.895","article-title":"Semantic transparency and masked morphological priming: The case of prefixed words","volume":"37","author":"K Diependaele","year":"2009","journal-title":"Memory & Cognition"},{"issue":"4\u20136","key":"pone.0249111.ref040","doi-asserted-by":"crossref","first-page":"482","DOI":"10.1080\/01690965.2010.500020","article-title":"On the interaction of letter transpositions and morphemic boundaries","volume":"26","author":"JG Rueckl","year":"2011","journal-title":"Language and Cognitive Processes"},{"issue":"5","key":"pone.0249111.ref041","doi-asserted-by":"crossref","first-page":"988","DOI":"10.3758\/s13423-013-0425-0","article-title":"Letter transpositions within and across morphemic boundaries: Is there a cross-language difference?","volume":"20","author":"C S\u00e1nchez-Guti\u00e9rrez","year":"2013","journal-title":"Psychonomic Bulletin & Review"},{"issue":"12","key":"pone.0249111.ref042","doi-asserted-by":"crossref","first-page":"4748","DOI":"10.1093\/cercor\/bhu163","article-title":"Orthographic coding: Brain activation for letters, symbols, and digits","volume":"25","author":"M Carreiras","year":"2014","journal-title":"Cerebral Cortex"},{"key":"pone.0249111.ref043","unstructured":"Korean Word Database. 21st century Sejong Project Corpus. Seoul: Korea: The National Institute of the Korean Language. 2001."},{"issue":"4","key":"pone.0249111.ref044","doi-asserted-by":"crossref","first-page":"373","DOI":"10.22172\/cogbio.2018.30.4.004","article-title":"Lexical factors that influence the Korean Eojeol recognition","volume":"30","author":"J Kim","year":"2018","journal-title":"The Korean Journal of Cognitive and Biological Psychology"},{"key":"pone.0249111.ref045","unstructured":"Brett M, Anton JL, Valabregue R, Poline JB. Region of interest analysis using the MarsBar toolbox for SPM 99 [abstract]. Paper Presented at the 8th International Conference on Functional Mapping of the Human Brain, Sendai. 2002."},{"issue":"1","key":"pone.0249111.ref046","doi-asserted-by":"crossref","first-page":"90","DOI":"10.1016\/j.neuroimage.2007.04.042","article-title":"A component based noise correction method (CompCor) for BOLD and perfusion based fMRI","volume":"37","author":"Y Behzadi","year":"2007","journal-title":"Neuroimage"},{"issue":"12","key":"pone.0249111.ref047","doi-asserted-by":"crossref","first-page":"1983","DOI":"10.1162\/jocn.2007.19.12.1983","article-title":"Neural correlates of morphological decomposition during visual word recognition","volume":"19","author":"BT Gold","year":"2007","journal-title":"Journal of Cognitive Neuroscience"},{"issue":"1","key":"pone.0249111.ref048","doi-asserted-by":"crossref","first-page":"66","DOI":"10.1016\/S0165-0173(99)00005-3","article-title":"A neuronal model of attentional spotlight: Parietal guiding the temporal","volume":"30","author":"TR Vidyasagar","year":"1999","journal-title":"Brain Research Reviews"},{"key":"pone.0249111.ref049","doi-asserted-by":"crossref","first-page":"297","DOI":"10.1016\/S0079-6123(01)34020-7","article-title":"From attentional gating in macaque primary visual cortex to dyslexia in humans","volume":"134","author":"TR Vidyasagar","year":"2001","journal-title":"Progress in Brain Research"},{"issue":"1","key":"pone.0249111.ref050","doi-asserted-by":"crossref","first-page":"20","DOI":"10.1016\/0166-2236(92)90344-8","article-title":"Separate visual pathways for perception and action","volume":"15","author":"MA Goodale","year":"1992","journal-title":"Trends in Neurosciences"},{"issue":"4853","key":"pone.0249111.ref051","doi-asserted-by":"crossref","first-page":"740","DOI":"10.1126\/science.3283936","article-title":"Segregation of form, color, movement, and depth: Anatomy, physiology, and perception","volume":"240","author":"M Livingstone","year":"1988","journal-title":"Science"},{"issue":"4","key":"pone.0249111.ref052","doi-asserted-by":"crossref","first-page":"364","DOI":"10.1038\/7274","article-title":"Involvement of striate and extrastriate visual cortical areas in spatial attention","volume":"2","author":"A Martinez","year":"1999","journal-title":"Nature Neuroscience"},{"issue":"6700","key":"pone.0249111.ref053","doi-asserted-by":"crossref","first-page":"376","DOI":"10.1038\/26475","article-title":"Object-based attention in the primary visual cortex of the macaque monkey","volume":"395","author":"PR Roelfsema","year":"1998","journal-title":"Nature"},{"key":"pone.0249111.ref054","volume-title":"The standard Korean grammar","author":"K. S. Nam","year":"1985"},{"key":"pone.0249111.ref055","doi-asserted-by":"crossref","first-page":"319","DOI":"10.1146\/annurev.neuro.22.1.319","article-title":"Space and attention in parietal cortex","volume":"22","author":"C.E. Colby","year":"1999","journal-title":"Annual Review of Neuroscience"},{"issue":"6984","key":"pone.0249111.ref056","doi-asserted-by":"crossref","first-page":"751","DOI":"10.1038\/nature02466","article-title":"Capacity limit of visual short-term memory in human posterior parietal cortex","volume":"428","author":"JJ Todd","year":"2004","journal-title":"Nature"},{"issue":"5","key":"pone.0249111.ref057","doi-asserted-by":"crossref","first-page":"465","DOI":"10.1093\/cercor\/7.5.465","article-title":"Bilateral Activation of Fronto-parietal Networks by Incrementing Demand in a Working Memory Task","volume":"7","author":"T Klingberg","year":"1997","journal-title":"Cerebral Cortex"},{"issue":"2","key":"pone.0249111.ref058","doi-asserted-by":"crossref","first-page":"532","DOI":"10.1093\/cercor\/bhs333","article-title":"Task Difficulty Manipulation Reveals Multiple Demand Activity but no Frontal Lobe Hierarchy","volume":"24","author":"BM Crittenden","year":"2014","journal-title":"Cerebral Cortex"},{"issue":"12","key":"pone.0249111.ref059","doi-asserted-by":"crossref","first-page":"1021","DOI":"10.1111\/j.1467-9280.2006.01821.x","article-title":"The Time Course of and Phonological Orthographic Code Activation","volume":"17","author":"J Grainger","year":"2006","journal-title":"Psychological Science"},{"issue":"7","key":"pone.0249111.ref060","doi-asserted-by":"crossref","first-page":"1610","DOI":"10.1162\/jocn_a_00180","article-title":"Differential sensitivity of letters, numbers, and symbols to character transpositions","volume":"24","author":"JA Du\u00f1abeitia","year":"2012","journal-title":"Journal of Cognitive Neuroscience"},{"issue":"1","key":"pone.0249111.ref061","doi-asserted-by":"crossref","first-page":"11","DOI":"10.1016\/j.bandl.2012.12.009","article-title":"ERP correlates of letter identity and letter position are modulated by lexical frequency","volume":"125","author":"M Vergara-Mart\u00ednez","year":"2013","journal-title":"Brain and Language"},{"issue":"4","key":"pone.0249111.ref062","doi-asserted-by":"crossref","first-page":"381","DOI":"10.1097\/WNR.0b013e3283249b1c","article-title":"N250 effects for letter transpositions depend on lexicality: \u201cCasual\u201d or \u201ccausal\u201d?","volume":"20","author":"JA Du\u00f1abeitia","year":"2009","journal-title":"NeuroReport"},{"issue":"2","key":"pone.0249111.ref063","doi-asserted-by":"crossref","first-page":"264","DOI":"10.1093\/cercor\/bhs007","article-title":"A meta-analysis of executive components of working memory","volume":"23","author":"DE Nee","year":"2012","journal-title":"Cerebral Cortex"},{"issue":"1","key":"pone.0249111.ref064","doi-asserted-by":"crossref","first-page":"74","DOI":"10.1162\/jocn_a_00281","article-title":"Intrinsic architecture underlying the relations among the default, dorsal attention, and frontoparietal control networks of the human brain","volume":"25","author":"RN Spreng","year":"2013","journal-title":"Journal of Cognitive Neuroscience"},{"issue":"2","key":"pone.0249111.ref065","doi-asserted-by":"crossref","first-page":"150","DOI":"10.1177\/1073858413494269","article-title":"Dorsal and ventral attention systems: Distinct neural circuits but collaborative roles","volume":"20","author":"S Vossel","year":"2014","journal-title":"The Neuroscientist"},{"issue":"4","key":"pone.0249111.ref066","doi-asserted-by":"crossref","first-page":"172","DOI":"10.1016\/j.tics.2010.01.004","article-title":"The multiple-demand (MD) system of the primate brain: Mental programs for intelligent behaviour","volume":"14","author":"J Duncan","year":"2010","journal-title":"Trends in Cognitive Sciences"},{"key":"pone.0249111.ref067","doi-asserted-by":"crossref","first-page":"9","DOI":"10.1016\/j.neuron.2006.12.009","article-title":"From thought to action: The parietal cortex as a bridge between perception, action, and cognition","volume":"53","author":"J Gottlieb","year":"2007","journal-title":"Neuron"},{"key":"pone.0249111.ref068","doi-asserted-by":"crossref","first-page":"91","DOI":"10.1038\/35039043","article-title":"Visual attention: Insights from brain imaging","volume":"1","author":"N Kanwisher","year":"2000","journal-title":"Nature Reviews Neuroscience"},{"issue":"5831","key":"pone.0249111.ref069","doi-asserted-by":"crossref","first-page":"1612","DOI":"10.1126\/science.1139140","article-title":"Neural mechanisms of visual attention: How top-down feedback highlights relevant locations","volume":"316","author":"YB Saalmann","year":"2007","journal-title":"Science"},{"key":"pone.0249111.ref070","doi-asserted-by":"crossref","first-page":"233","DOI":"10.1007\/s10548-006-0001-2","article-title":"FMRI of ventral and dorsal processing streams in basic reading processes: Insular sensitivity to phonology","volume":"18","author":"R Borowsky","year":"2006","journal-title":"Brain Topography"},{"key":"pone.0249111.ref071","doi-asserted-by":"crossref","first-page":"353","DOI":"10.1016\/j.neuroimage.2007.11.036","article-title":"Reading normal and degraded words: Contribution of the dorsal and ventral pathways","volume":"40","author":"L Cohen","year":"2008","journal-title":"Neuroimage"},{"issue":"14","key":"pone.0249111.ref072","doi-asserted-by":"crossref","first-page":"2926","DOI":"10.1016\/j.neuropsychologia.2006.06.028","article-title":"Attentional shifting and the role of the dorsal pathway in visual word recognition","volume":"44","author":"K Pammer","year":"2006","journal-title":"Neuropsychologia"},{"issue":"6","key":"pone.0249111.ref073","first-page":"479","article-title":"Neurobiological studies of reading and reading disability","volume":"34","author":"KR Pugh","year":"2001","journal-title":"Journal of Communicable Diseases"},{"issue":"14","key":"pone.0249111.ref074","doi-asserted-by":"crossref","first-page":"2181","DOI":"10.1016\/S0042-6989(98)00016-9","article-title":"Coherent motion detection and letter position encoding","volume":"38","author":"PL Cornelissen","year":"1998","journal-title":"Vision Research"},{"issue":"6586","key":"pone.0249111.ref075","doi-asserted-by":"crossref","first-page":"66","DOI":"10.1038\/382066a0","article-title":"Abnormal processing of visual motion in dyslexia revealed by functional brain imaging","volume":"382","author":"GF Eden","year":"1996","journal-title":"Nature"},{"issue":"6","key":"pone.0249111.ref076","doi-asserted-by":"crossref","first-page":"1057","DOI":"10.1016\/j.neuron.2012.12.002","article-title":"The role of medial prefrontal cortex in memory and decision making","volume":"76","author":"DR Euston","year":"2012","journal-title":"Neuron"},{"issue":"5680","key":"pone.0249111.ref077","doi-asserted-by":"crossref","first-page":"96","DOI":"10.1126\/science.1098180","article-title":"Sites of neocortical reorganization critical for remote spatial memory","volume":"305","author":"T Maviel","year":"2004","journal-title":"Science"},{"issue":"2","key":"pone.0249111.ref078","doi-asserted-by":"crossref","first-page":"325","DOI":"10.1016\/j.bbr.2010.12.009","article-title":"The role of the ventromedial prefrontal cortex in memory consolidation","volume":"218","author":"IL Nieuwenhuis","year":"2011","journal-title":"Behavioural Brain Research"},{"key":"pone.0249111.ref079","doi-asserted-by":"crossref","first-page":"433","DOI":"10.1146\/annurev-neuro-071013-014030","article-title":"The brain\u2019s default mode network","volume":"38","author":"ME Raichle","year":"2015","journal-title":"Annual Review of Neuroscience"},{"issue":"8","key":"pone.0249111.ref080","doi-asserted-by":"crossref","first-page":"3047","DOI":"10.1002\/hbm.22827","article-title":"Specific default mode subnetworks support mentalizing as revealed through opposing network recruitment by social and semantic FMRI tasks","volume":"36","author":"CJ Hyatt","year":"2015","journal-title":"Human Brain Mapping"},{"issue":"1","key":"pone.0249111.ref081","doi-asserted-by":"crossref","first-page":"80","DOI":"10.1162\/089892999563265","article-title":"Conceptual processing during the conscious resting state: A functional MRI study","volume":"11","author":"JR Binder","year":"1999","journal-title":"Journal of Cognitive Neuroscience"},{"issue":"12","key":"pone.0249111.ref082","doi-asserted-by":"crossref","first-page":"2767","DOI":"10.1093\/cercor\/bhp055","article-title":"Where is the semantic system? A critical review and meta-analysis of 120 functional neuroimaging studies","volume":"19","author":"JR Binder","year":"2009","journal-title":"Cerebral Cortex"},{"issue":"1","key":"pone.0249111.ref083","doi-asserted-by":"crossref","first-page":"42","DOI":"10.1038\/nrn.2016.150","article-title":"The neural and computational bases of semantic cognition","volume":"18","author":"MAL Ralph","year":"2017","journal-title":"Nature Reviews Neuroscience"},{"key":"pone.0249111.ref084","article-title":"The neural bases of resilient cognitive systems: Evidence of variable neuro-displacement in the semantic system","author":"J Jung","year":"2019","journal-title":"bioRxiv"},{"issue":"3","key":"pone.0249111.ref085","doi-asserted-by":"crossref","first-page":"206","DOI":"10.1177\/1534582302001003002","article-title":"Semantic retrieval, mnemonic control, and prefrontal cortex","volume":"1","author":"D Badre","year":"2002","journal-title":"Behavioral and Cognitive Neuroscience Reviews"},{"issue":"11","key":"pone.0249111.ref086","doi-asserted-by":"crossref","first-page":"1824","DOI":"10.1162\/jocn_a_00442","article-title":"Going beyond inferior prefrontal involvement in semantic control: Evidence for the additional contribution of dorsal angular gyrus and posterior middle temporal cortex","volume":"25","author":"KA Noonan","year":"2013","journal-title":"Journal of Cognitive Neuroscience"},{"issue":"26","key":"pone.0249111.ref087","doi-asserted-by":"crossref","first-page":"14792","DOI":"10.1073\/pnas.94.26.14792","article-title":"Role of left inferior prefrontal cortex in retrieval of semantic knowledge: A reevaluation","volume":"94","author":"SL Thompson-Schill","year":"1997","journal-title":"Proceedings of the National Academy of Sciences"},{"issue":"8","key":"pone.0249111.ref088","doi-asserted-by":"crossref","first-page":"1228","DOI":"10.1162\/jocn_a_00959","article-title":"Spatiotemporal dynamics of morphological processing in visual word recognition","volume":"28","author":"E Cavalli","year":"2016","journal-title":"Journal of Cognitive Neuroscience"},{"issue":"1","key":"pone.0249111.ref089","doi-asserted-by":"crossref","first-page":"128","DOI":"10.1111\/j.1749-818X.2008.00121.x","article-title":"Watching the word go by: On the time\u2010course of component processes in visual word recognition","volume":"3","author":"J Grainger","year":"2009","journal-title":"Language and Linguistics Compass"},{"issue":"4","key":"pone.0249111.ref090","doi-asserted-by":"crossref","first-page":"1383","DOI":"10.1016\/j.neuroimage.2005.11.048","article-title":"The time course of visual word recognition as revealed by linear regression analysis of ERP data","volume":"30","author":"O Hauk","year":"2006","journal-title":"Neuroimage"},{"issue":"3","key":"pone.0249111.ref091","doi-asserted-by":"crossref","first-page":"374","DOI":"10.1016\/j.biopsycho.2006.09.008","article-title":"Early semantic context integration and lexical access as revealed by event-related brain potentials","volume":"74","author":"B Penolazzi","year":"2007","journal-title":"Biological Psychology"},{"issue":"1","key":"pone.0249111.ref092","doi-asserted-by":"crossref","first-page":"201","DOI":"10.1046\/j.0953-816X.2000.01380.x","article-title":"Neuromagnetic evidence for early semantic access in word recognition","volume":"13","author":"F Pulverm\u00fcller","year":"2001","journal-title":"European Journal of Neuroscience"},{"key":"pone.0249111.ref093","doi-asserted-by":"crossref","first-page":"11","DOI":"10.3389\/fnhum.2012.00011","article-title":"The time course of semantic richness effects in visual word recognition","volume":"6","author":"M Rabovsky","year":"2012","journal-title":"Frontiers in Human Neuroscience"},{"issue":"3","key":"pone.0249111.ref094","doi-asserted-by":"crossref","first-page":"173","DOI":"10.1016\/S0926-6410(97)00033-5","article-title":"Evoked potential correlates of semantic meaning\u2014A brain mapping study","volume":"6","author":"W Skrandies","year":"1998","journal-title":"Cognitive Brain Research"},{"key":"pone.0249111.ref095","doi-asserted-by":"crossref","first-page":"439","DOI":"10.1016\/j.neuroimage.2019.05.002","article-title":"How meaning unfolds in neural time: Embodied reactivations can precede multimodal semantic effects during language processing","volume":"197","author":"AM Garc\u00eda","year":"2019","journal-title":"Neuroimage"},{"issue":"1","key":"pone.0249111.ref096","doi-asserted-by":"crossref","first-page":"e0169269","DOI":"10.1371\/journal.pone.0169269","article-title":"Oscillatory dynamics supporting semantic cognition: MEG evidence for the contribution of the anterior temporal lobe hub and modality-specific spokes","volume":"12","author":"G Mollo","year":"2017","journal-title":"PloS One"},{"key":"pone.0249111.ref097","doi-asserted-by":"crossref","first-page":"1","DOI":"10.1016\/j.cortex.2016.02.015","article-title":"The \u2018when\u2019 and \u2018where\u2019 of semantic coding in the anterior temporal lobe: Temporal representational similarity analysis of electrocorticogram data","volume":"79","author":"Y Chen","year":"2016","journal-title":"Cortex"},{"issue":"1","key":"pone.0249111.ref098","doi-asserted-by":"crossref","first-page":"93","DOI":"10.1006\/nimg.1998.0350","article-title":"Brain activation during word identification and word recognition","volume":"8","author":"TL Jernigan","year":"1998","journal-title":"NeuroImage"},{"issue":"5","key":"pone.0249111.ref099","doi-asserted-by":"crossref","first-page":"535","DOI":"10.1038\/nn.2303","article-title":"Circular analysis in systems neuroscience\u2013the dangers of double dipping","volume":"12","author":"N Kriegeskorte","year":"2009","journal-title":"Nature Neuroscience"},{"issue":"2","key":"pone.0249111.ref100","doi-asserted-by":"crossref","first-page":"90","DOI":"10.1016\/j.tics.2013.11.005","article-title":"The what, when, where, and how of visual word recognition","volume":"18","author":"M Carreiras","year":"2014","journal-title":"Trends in Cognitive Sciences"}],"container-title":["PLOS ONE"],"original-title":[],"language":"en","link":[{"URL":"https:\/\/dx.plos.org\/10.1371\/journal.pone.0249111","content-type":"unspecified","content-version":"vor","intended-application":"similarity-checking"}],"deposited":{"date-parts":[[2021,4,15]],"date-time":"2021-04-15T18:08:12Z","timestamp":1618510092000},"score":1,"resource":{"primary":{"URL":"https:\/\/dx.plos.org\/10.1371\/journal.pone.0249111"}},"subtitle":[],"editor":[{"given":"Manuel","family":"Perea","sequence":"first","affiliation":[],"role":[{"vocabulary":"crossref","role":"editor"}]}],"short-title":[],"issued":{"date-parts":[[2021,4,15]]},"references-count":100,"journal-issue":{"issue":"4","published-online":{"date-parts":[[2021,4,15]]}},"URL":"https:\/\/doi.org\/10.1371\/journal.pone.0249111","relation":{},"ISSN":["1932-6203"],"issn-type":[{"value":"1932-6203","type":"electronic"}],"subject":[],"published":{"date-parts":[[2021,4,15]]}}}