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The discovery was made about 30 km to the SW of the town of El Calafate (Province of Santa Cruz, Argentina) and consists of a beautifully preserved and articulated skull and jaws, and part of the postcranial skeleton that were preserved encased in a large concretion. This new taxon belongs to the notosuchian clade Peirosauridae, representing the latest and southernmost record for this group of crocodyliforms. The new taxon is recovered as closely related to other robust and broad-snouted peirosaurids that lived by the end of the Cretaceous Period, such as <jats:italic><jats:italic>Colhuehuapisuchus<\/jats:italic><\/jats:italic> from the Maastrichtian of Central Patagonia and <jats:italic><jats:italic>Miadanasuchus oblita<\/jats:italic><\/jats:italic> from the Maastrichtian of Madagascar. The completeness of the new specimen reveals, for the first time, the anatomy and body plan of a large and broad snouted peirosaurid. The new taxon bears large ziphodont teeth, a broad oreinirostral snout that is only slightly longer than 50% the skull length, and a deep adductor chamber in the temporal region and posterior mandibular ramus. The anterior region of its postcranial skeleton is preserved and shows broad scapula and a robust humerus features previously known in large predatorial notosuchians. The new crocodyliform adds to the predatorial component of terrestrial ecosystems at high paleolatitudes by the end of the Cretaceous Period.<\/jats:p>","DOI":"10.1371\/journal.pone.0328561","type":"journal-article","created":{"date-parts":[[2025,8,27]],"date-time":"2025-08-27T17:21:58Z","timestamp":1756315318000},"page":"e0328561","update-policy":"https:\/\/doi.org\/10.1371\/journal.pone.corrections_policy","source":"Crossref","is-referenced-by-count":8,"title":["A new large hypercarnivorous crocodyliform from the Maastrichtian of Southern Patagonia, Argentina"],"prefix":"10.1371","volume":"20","author":[{"given":"Fernando E.","family":"Novas","sequence":"first","affiliation":[],"role":[{"vocabulary":"crossref","role":"author"}]},{"ORCID":"https:\/\/orcid.org\/0000-0002-9690-7517","authenticated-orcid":true,"given":"Diego","family":"Pol","sequence":"additional","affiliation":[],"role":[{"vocabulary":"crossref","role":"author"}]},{"given":"Federico 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Mannion","year":"2019","journal-title":"Palaeontology"},{"key":"pone.0328561.ref024","doi-asserted-by":"crossref","first-page":"354","DOI":"10.1671\/0272-4634(2005)025[0354:FDABSO]2.0.CO;2","article-title":"Femoral dimensions and body size of Alligator mississippiensis: estimating the size of extinct mesoeucrocodylians","volume":"25","author":"JO Farlow","year":"2005","journal-title":"J Vert Paleontol"},{"issue":"1","key":"pone.0328561.ref025","doi-asserted-by":"crossref","first-page":"2373987","DOI":"10.1080\/14772019.2024.2373987","article-title":"A new species of Araripesuchus with durophagous dentition increases the ecological disparity among uruguaysuchid crocodyliforms","volume":"22","author":"ML Fern\u00e1ndez-Dumont","year":"2024","journal-title":"J Syst Palaeontol"},{"issue":"4","key":"pone.0328561.ref026","article-title":"A new notosuchian from the Late Cretaceous of Brazil and the phylogeny of advanced notosuchians","volume":"9","author":"D Pol","year":"2014","journal-title":"PLoS One"},{"issue":"6","key":"pone.0328561.ref027","doi-asserted-by":"crossref","DOI":"10.1080\/02724634.2014.995187","article-title":"The postcranial anatomy of Yacarerani boliviensis and the phylogenetic significance of the notosuchian postcranial skeleton","volume":"35","author":"JM Leardi","year":"2015","journal-title":"J Vert Paleontol"},{"issue":"2","key":"pone.0328561.ref028","doi-asserted-by":"crossref","first-page":"144","DOI":"10.1111\/cla.12524","article-title":"TNTversion 1.6, with a graphical interface forMacOSand Linux, including new routines in parallel","volume":"39","author":"PA Goloboff","year":"2023","journal-title":"Cladistics"},{"issue":"5","key":"pone.0328561.ref029","doi-asserted-by":"crossref","first-page":"515","DOI":"10.1111\/j.1096-0031.2009.00258.x","article-title":"Unstable taxa in cladistic analysis: identification and the assessment of relevant characters","volume":"25","author":"D Pol","year":"2009","journal-title":"Cladistics"},{"key":"pone.0328561.ref030","doi-asserted-by":"crossref","first-page":"93","DOI":"10.1016\/j.ympev.2015.04.003","article-title":"Identifying unstable taxa: efficient implementation of triplet-based measures of stability, and comparison with Phyutility and RogueNaRok","volume":"88","author":"PA Goloboff","year":"2015","journal-title":"Mol Phylogenetics Evol"},{"issue":"4","key":"pone.0328561.ref031","doi-asserted-by":"crossref","first-page":"862","DOI":"10.1111\/brv.12030","article-title":"A new fossil from the Jurassic of Patagonia reveals the early basicranial evolution and the origins of Crocodyliformes","volume":"88","author":"D Pol","year":"2013","journal-title":"Biol Rev Camb Philos Soc"},{"key":"pone.0328561.ref032","first-page":"97","article-title":"The impact of unstable taxa in coelurosaurian phylogeny and resampling support measures for parsimony analyses","volume":"440","author":"D Pol","year":"2020","journal-title":"Bull Am Mus Nat Hist"},{"issue":"4","key":"pone.0328561.ref033","doi-asserted-by":"crossref","first-page":"975","DOI":"10.1016\/S1342-937X(05)71079-0","article-title":"Uberabasuchus terrificus sp. nov., a new Crocodylomorpha from the Bauru Basin (Upper Cretaceous), Brazil","volume":"7","author":"IS Carvalho","year":"2004","journal-title":"Gondwana Res"},{"issue":"1","key":"pone.0328561.ref034","first-page":"35","article-title":"Montealtosuchus arrudacamposi, a new peirosaurid crocodile (Mesoeucrocodylia) from the Late Cretaceous Adamantina Formation of Brazil","volume":"1607","author":"IS Carvalho","year":"2007","journal-title":"Zootaxa"},{"key":"pone.0328561.ref035","doi-asserted-by":"crossref","first-page":"470","DOI":"10.1111\/brv.12666","article-title":"Assessing ontogenetic maturity in extinct saurian reptiles","volume":"96","author":"CT Griffin","year":"2021","journal-title":"Biol 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Anat"},{"issue":"3","key":"pone.0328561.ref039","doi-asserted-by":"crossref","first-page":"255","DOI":"10.1080\/08912960500516112","article-title":"Osteology and phylogeny of a new species of Araripesuchus (Crocodyliformes: Mesoeucrocodylia) from the Late Cretaceous of Madagascar","volume":"18","author":"AH Turner","year":"2006","journal-title":"Hist Biol"},{"issue":"2","key":"pone.0328561.ref040","doi-asserted-by":"crossref","first-page":"328","DOI":"10.1080\/02724634.2012.646833","article-title":"Postcranial anatomy of Sebecus icaeorhinus (Crocodyliformes, Sebecidae) from the Eocene of Patagonia","volume":"32","author":"D Pol","year":"2012","journal-title":"J Vert Paleontol"},{"key":"pone.0328561.ref041","first-page":"317","article-title":"Una nueva familia de cocodrilos zifodontes cret\u00e1cicos de Am\u00e9rica del Sur","volume":"4","author":"Z Gasparini","year":"1981","journal-title":"Actas V Congr Latinoam Geol Buenos Aires"},{"key":"pone.0328561.ref042","first-page":"116","article-title":"Crocodyliformes.","volume-title":"Patagonian mesozoic reptiles","author":"D Pol","year":"2007"},{"issue":"3","key":"pone.0328561.ref043","first-page":"44","article-title":"Phylogenetic nomenclature of Notosuchia (Crocodylomorpha; Crocodyliformes)","volume":"1","author":"JM Leardi","year":"2024","journal-title":"BPN"},{"issue":"12","key":"pone.0328561.ref044","doi-asserted-by":"crossref","first-page":"2280","DOI":"10.1080\/08912963.2022.2139179","article-title":"New Itasuchidae (Sebecia, Ziphosuchia) remains and the radiation of an elusive Mesoeucrocodylia clade","volume":"35","author":"AEP Pinheiro","year":"2023","journal-title":"Hist Biol"},{"key":"pone.0328561.ref045","first-page":"1","article-title":"A new Peirosauridae (Crocodyliformes, Notosuchia) from the Adamantina Formation (Bauru Group, Late Cretaceous), with a revised phylogenetic analysis of Sebecia","author":"JV Ruiz","year":"2024","journal-title":"Anat Rec"},{"issue":"2","key":"pone.0328561.ref046","doi-asserted-by":"crossref","DOI":"10.1002\/spp2.1485","article-title":"A peirosaurid mandible from the Albian\u2013Cenomanian (Lower Cretaceous) of Algeria and the taxonomic content of Hamadasuchus (Crocodylomorpha, Peirosauridae)","volume":"9","author":"Y Pochat\u2010Cottilloux","year":"2023","journal-title":"Pap Palaeontol"},{"key":"pone.0328561.ref047","doi-asserted-by":"crossref","first-page":"191","DOI":"10.1016\/j.cretres.2012.03.017","article-title":"A new peirosaurid from the Upper Cretaceous of Argentina: implications for specimens referred to Peirosaurus torminni Price (Crocodyliformes: Peirosauridae)","volume":"37","author":"A Martinelli","year":"2012","journal-title":"Cretac Res"},{"issue":"6","key":"pone.0328561.ref048","doi-asserted-by":"crossref","first-page":"835","DOI":"10.1080\/08912963.2015.1043999","article-title":"A new peirosaurid (Crocodilyformes) from the Late Cretaceous (Turonian\u2013Coniacian) of Patagonia, Argentina","volume":"28","author":"G Lio","year":"2016","journal-title":"Hist Biol"},{"issue":"3","key":"pone.0328561.ref049","doi-asserted-by":"crossref","first-page":"193","DOI":"10.2992\/007.085.0301","article-title":"A new peirosaurid crocodyliform from the Upper Cretaceous Lago Colhu\u00e9 Huapi Formation of Central Patagonia, Argentina","volume":"85","author":"MC Lamanna","year":"2019","journal-title":"Ann Carnegie Mus"},{"issue":"2","key":"pone.0328561.ref050","doi-asserted-by":"crossref","first-page":"599","DOI":"10.1671\/039.029.0224","article-title":"New material of \u201cTrematochampsa\u201d oblita (Crocodyliformes, Trematochampsidae) from the Late Cretaceous of Madagascar","volume":"29","author":"EL Rasmusson-Simons","year":"2009","journal-title":"J Vert Paleontol"},{"issue":"3","key":"pone.0328561.ref051","doi-asserted-by":"crossref","first-page":"316","DOI":"10.1080\/02724634.1991.10011401","article-title":"A new Senonian peirosaurid (Crocodylomorpha) from Argentina and a synopsis of the South American Cretaceous crocodilians","volume":"11","author":"Z Gasparini","year":"1991","journal-title":"J Vert Paleontol"},{"key":"pone.0328561.ref052","doi-asserted-by":"crossref","first-page":"183","DOI":"10.2113\/gssgfbull.S7-XXI.2.183","article-title":"Un nouveau crocodilien m\u00e9sosuchien dans le Campanien de Madagascar, Trematochampsa oblita, n.sp","volume":"2","author":"E Buffetaut","year":"1979","journal-title":"Bull Soc G\u00e9ol Fr"},{"issue":"4","key":"pone.0328561.ref053","doi-asserted-by":"crossref","first-page":"543","DOI":"10.1111\/pala.12354","article-title":"Evidence for heterochrony in the cranial evolution of fossil crocodyliforms","volume":"61","author":"PL Godoy","year":"2019","journal-title":"Palaeontology"},{"issue":"2","key":"pone.0328561.ref054","doi-asserted-by":"crossref","first-page":"323","DOI":"10.1111\/joa.13192","article-title":"A unique predator in a unique ecosystem: modelling the apex predator within a Late Cretaceous crocodyliform\u2010dominated fauna from Brazil","volume":"237","author":"F Montefeltro","year":"2020","journal-title":"J Anat"},{"issue":"1","key":"pone.0328561.ref055","doi-asserted-by":"crossref","DOI":"10.1111\/pala.12638","article-title":"Phylogenetic structure of the extinction and biotic factors explaining differential survival of terrestrial notosuchians at the Cretaceous\u2013Palaeogene crisis","volume":"66","author":"P Aubier","year":"2023","journal-title":"Palaeontology"},{"key":"pone.0328561.ref056","doi-asserted-by":"crossref","DOI":"10.1111\/j.1096-3642.2011.00713.x","article-title":"Baurusuchid crocodyliforms as theropod mimics: clues from the skull and appendicular morphology of Stratiotosuchus maxhechti (Upper Cretaceous of Brazil)","volume":"163","author":"D Riff","year":"2011","journal-title":"Zool J Linn 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Parrish","year":"1987","journal-title":"Paleobiology"},{"key":"pone.0328561.ref066","first-page":"227","article-title":"The morphology and relationships of a crocodilian, Orthosuchus stormbergi, from the Upper Triassic of Lesotho","volume":"67","author":"D Nash","year":"1975","journal-title":"Ann S Afr Mus"},{"key":"pone.0328561.ref067","first-page":"1","article-title":"A \u201ccurosorial\u201d crocodilian from the Triassic of Lesotho (Basutoland), Southern Africa","volume":"106","author":"KN Whetstone","year":"1983","journal-title":"Occ Pap Mus Nat Hist"},{"issue":"1","key":"pone.0328561.ref068","first-page":"21","article-title":"Postcranial remains of Notosuchus terrestris Woodward (Archosauria: Crocodyliformes) from the upper cretaceous of Patagonia, Argentina","volume":"42","author":"D Pol","year":"2005","journal-title":"Ameghiniana"},{"issue":"2","key":"pone.0328561.ref069","doi-asserted-by":"crossref","first-page":"266","DOI":"10.1002\/ar.25586","article-title":"A review of the 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Palaeoecol"},{"issue":"3","key":"pone.0328561.ref072","doi-asserted-by":"crossref","first-page":"255","DOI":"10.1080\/08912960500516112","article-title":"Osteology and phylogeny of a new species of Araripesuchus (Crocodyliformes: Mesoeucrocodylia) from the Late Cretaceous of Madagascar","volume":"18","author":"AH Turner","year":"2006","journal-title":"Hist Biol"},{"key":"pone.0328561.ref073","first-page":"323","article-title":"A new species of Baurusuchus (Crocodyliformes, Mesoeucrocodylia) from the Upper Cretaceous of Brazil, with the first complete postcranial skeleton described for the family Baurusuchidae","volume":"50","author":"PM Nascimento","year":"2010","journal-title":"Pap Avulsos Zool"},{"key":"pone.0328561.ref074","doi-asserted-by":"crossref","DOI":"10.7717\/peerj.2075","article-title":"Postcranial anatomy of Pissarrachampsa sera (Crocodyliformes, Baurusuchidae) from the Late Cretaceous of Brazil: insights on lifestyle and phylogenetic significance","volume":"4","author":"PL Godoy","year":"2016","journal-title":"PeerJ"},{"issue":"3","key":"pone.0328561.ref075","doi-asserted-by":"crossref","first-page":"1","DOI":"10.18261\/let.56.3.2","article-title":"Is there any evidence for the presence of extensive fibrolamellar complex in Notosuchia? New data on Araripesuchus wegeneri","volume":"56","author":"MG Faure-Brac","year":"2023","journal-title":"Lethaia"},{"key":"pone.0328561.ref076","first-page":"17","article-title":"Los vertebrados f\u00f3siles de la Formaci\u00f3n R\u00edo Colorado, de la ciudad de Neuqu\u00e9n y cercan\u00edas, Cret\u00e1cico superior, Argentina","volume":"4","author":"JF Bonaparte","year":"1991","journal-title":"Rev del Mus Argentino Ciencias Nat"},{"issue":"2","key":"pone.0328561.ref077","doi-asserted-by":"crossref","first-page":"252","DOI":"10.1016\/j.palaeo.2010.08.003","article-title":"Climate\u2019s role in the distribution of the Cretaceous terrestrial Crocodyliformes throughout Gondwana","volume":"297","author":"IS Carvalho","year":"2010","journal-title":"Palaeogeogr Palaeoclimatol Palaeoecol"},{"issue":"60","key":"pone.0328561.ref078","first-page":"149","article-title":"An enigmatic new crocodile from the Upper Cretaceous of Madagascar","volume":"60","author":"GA Buckley","year":"1999","journal-title":"Spec Pap Palaeontol"},{"issue":"2","key":"pone.0328561.ref079","doi-asserted-by":"crossref","first-page":"382","DOI":"10.1671\/0272-4634(2008)28[382:MICMCA]2.0.CO;2","article-title":"Mahajangasuchus insignis (Crocodyliformes: Mesoeucrocodylia) cranial anatomy and new data on the origin of the eusuchian-style palate","volume":"28","author":"AH Turner","year":"2008","journal-title":"J Vert Paleontol"},{"key":"pone.0328561.ref080","doi-asserted-by":"crossref","first-page":"335","DOI":"10.1016\/j.cretres.2018.06.008","article-title":"Bone microanatomy of Pepesuchus deiseae (Mesoeucrocodylia, Peirosauridae) reveals a mature individual from the Upper Cretaceous of Brazil","volume":"90","author":"MV Sena","year":"2018","journal-title":"Cretac Res"},{"issue":"5","key":"pone.0328561.ref081","doi-asserted-by":"crossref","first-page":"803","DOI":"10.1111\/j.2041-210X.2012.00223.x","article-title":"Paleotree: an R package for paleontological and phylogenetic analyses of evolution","volume":"3","author":"DW Bapst","year":"2012","journal-title":"Methods Ecol Evol"}],"container-title":["PLOS One"],"original-title":[],"language":"en","link":[{"URL":"https:\/\/dx.plos.org\/10.1371\/journal.pone.0328561","content-type":"unspecified","content-version":"vor","intended-application":"similarity-checking"}],"deposited":{"date-parts":[[2025,8,27]],"date-time":"2025-08-27T17:22:08Z","timestamp":1756315328000},"score":1,"resource":{"primary":{"URL":"https:\/\/dx.plos.org\/10.1371\/journal.pone.0328561"}},"subtitle":[],"editor":[{"given":"Paolo","family":"Piras","sequence":"first","affiliation":[],"role":[{"vocabulary":"crossref","role":"editor"}]}],"short-title":[],"issued":{"date-parts":[[2025,8,27]]},"references-count":81,"journal-issue":{"issue":"8","published-online":{"date-parts":[[2025,8,27]]}},"URL":"https:\/\/doi.org\/10.1371\/journal.pone.0328561","relation":{},"ISSN":["1932-6203"],"issn-type":[{"value":"1932-6203","type":"electronic"}],"subject":[],"published":{"date-parts":[[2025,8,27]]}}}