{"status":"ok","message-type":"work","message-version":"1.0.0","message":{"indexed":{"date-parts":[[2026,7,15]],"date-time":"2026-07-15T18:06:19Z","timestamp":1784138779505,"version":"3.55.0"},"reference-count":64,"publisher":"Public Library of Science (PLoS)","issue":"7","license":[{"start":{"date-parts":[[2026,7,15]],"date-time":"2026-07-15T00:00:00Z","timestamp":1784073600000},"content-version":"vor","delay-in-days":0,"URL":"http:\/\/creativecommons.org\/licenses\/by\/4.0\/"}],"content-domain":{"domain":["www.plosone.org"],"crossmark-restriction":false},"short-container-title":["PLoS One"],"abstract":"<jats:p>\n                    Identifying the origin of perforating lesions on fossil bone is often difficult, and many are considered tooth traces, in spite of more likely and more parsimonious etiologies. Much of this confusion stems from tooth trace criteria that are ambiguous when the context for the lesions is not considered. Mistaken identification of tooth traces has led to misleading interpretations of animal behavior. This study of tooth traces on fossil bones critically reviews previous criteria and applies them to assessing bones from an\n                    <jats:italic>Edmontosaurus annectens<\/jats:italic>\n                    bonebed within the Lance Formation, Wyoming, USA. Of the 3013 bones examined, thirteen bones had features indicative of tooth traces based on gross appearance. Of these, one bone had perforations determined to have a different etiology. Twelve bones had traces attributed to tooth marks, including four bones with\n                    <jats:italic>Knethichnus parallelum<\/jats:italic>\n                    and\n                    <jats:italic>Linichnus serratus<\/jats:italic>\n                    ichnotaxa.\n                    <jats:italic>Tyrannosaurus rex<\/jats:italic>\n                    was identified as the likely inflictor of traces attributable to both ichnotaxa, by comparison of denticle density of carnivore teeth within the bonebed with striation\/serration density of the traces. The importance of context in the analysis of perforating lesions on fossil bones is shown through the mistaken identification of features such as neurovasculature foramina and lesions associated with pathology as tooth traces. This study contributes to the literature on biting behavior and refines the criteria used to identify perforations caused by bite marks. The application of these refined criteria also proved useful in accurately identifying tooth traces on bone. This, in turn, enhances the guidelines for recognizing perforations as tooth traces and encourages further research on this topic.\n                  <\/jats:p>","DOI":"10.1371\/journal.pone.0351939","type":"journal-article","created":{"date-parts":[[2026,7,15]],"date-time":"2026-07-15T17:22:07Z","timestamp":1784136127000},"page":"e0351939","update-policy":"https:\/\/doi.org\/10.1371\/journal.pone.corrections_policy","source":"Crossref","is-referenced-by-count":0,"title":["Identification of tooth traces from a Cretaceous (Maastrichtian) Edmontosaurus annectens bonebed in the Lance Formation, Wyoming, U.S.A."],"prefix":"10.1371","volume":"21","author":[{"ORCID":"https:\/\/orcid.org\/0009-0009-8456-0227","authenticated-orcid":true,"given":"Bethania","family":"C. T. Siviero","sequence":"first","affiliation":[],"role":[{"vocabulary":"crossref","role":"author"}]},{"given":"Elizabeth","family":"Rega","sequence":"additional","affiliation":[],"role":[{"vocabulary":"crossref","role":"author"}]},{"given":"Matthew A.","family":"McLain","sequence":"additional","affiliation":[],"role":[{"vocabulary":"crossref","role":"author"}]},{"given":"Leonard R.","family":"Brand","sequence":"additional","affiliation":[],"role":[{"vocabulary":"crossref","role":"author"}]},{"given":"David","family":"Nelsen","sequence":"additional","affiliation":[],"role":[{"vocabulary":"crossref","role":"author"}]},{"given":"Art V.","family":"Chadwick","sequence":"additional","affiliation":[],"role":[{"vocabulary":"crossref","role":"author"}]}],"member":"340","published-online":{"date-parts":[[2026,7,15]]},"reference":[{"key":"pone.0351939.ref001","first-page":"167","article-title":"Head-biting behavior in theropod dinosaurs: paleopathological evidence","volume":"15","author":"D Tanke","year":"1998","journal-title":"GAIA"},{"key":"pone.0351939.ref002","doi-asserted-by":"crossref","first-page":"161","DOI":"10.2307\/jj.6047951.11","article-title":"Taphonomic approaches to bite-mark analyses in the fossil record and applications to archosaurian paleobiology.","volume-title":"Ruling reptiles: crocodylian biology and archosaur paleobiology","author":"SK Drumheller","year":"2023"},{"issue":"2","key":"pone.0351939.ref003","doi-asserted-by":"crossref","first-page":"105","DOI":"10.2110\/palo.2011.p11-052r","article-title":"Feeding traces and paleobiology of a Cretaceous (Cenomanian) crocodyliform: example from the Woodbine Formation of Texas","volume":"27","author":"CR Noto","year":"2012","journal-title":"Palaios"},{"key":"pone.0351939.ref004","article-title":"Bite and tooth marks on sauropod dinosaurs from the Morrison Formation","volume":"11","author":"R Lei","year":"2023","journal-title":"PeerJ"},{"issue":"7","key":"pone.0351939.ref005","doi-asserted-by":"crossref","first-page":"922","DOI":"10.1139\/e95-077","article-title":"An azhdarchid pterosaur eaten by a velociraptorine theropod","volume":"32","author":"PJ Currie","year":"1995","journal-title":"Can J Earth Sci"},{"key":"pone.0351939.ref006","first-page":"183","article-title":"Bite marks of the giant crocodylian Deinosuchus on Late Cretaceous (Campanian) bones","volume":"51","author":"DR Schwimmer","year":"2010","journal-title":"N M Mus Nat Hist Sci Bull"},{"issue":"3","key":"pone.0351939.ref007","doi-asserted-by":"crossref","first-page":"387","DOI":"10.5710\/AMGH.v47i3.1","article-title":"Predation bite-marks on a peirosaurid crocodyliform from the Upper Cretaceous of Neuqu\u00e9n Province, Argentina","volume":"47","author":"LE Fiorelli","year":"2010","journal-title":"Ameghiniana"},{"issue":"2","key":"pone.0351939.ref008","article-title":"Crocodyliform feeding traces on juvenile ornithischian dinosaurs from the Upper Cretaceous (Campanian) Kaiparowits Formation, Utah","volume":"8","author":"CA Boyd","year":"2013","journal-title":"PLoS One"},{"key":"pone.0351939.ref009","doi-asserted-by":"crossref","first-page":"304","DOI":"10.1016\/j.cretres.2014.05.006","article-title":"Inferred bite marks on a Late Cretaceous (Santonian) bothremydid turtle and a hylaeochampsid crocodilian from Hungary","volume":"50","author":"G Botfalvai","year":"2014","journal-title":"Cretac Res"},{"issue":"4","key":"pone.0351939.ref010","doi-asserted-by":"crossref","first-page":"703","DOI":"10.1111\/j.1475-4983.2010.00957.x","article-title":"Mammalian tooth marks on the bones of dinosaurs and other Late Cretaceous vertebrates","volume":"53","author":"NR Longrich","year":"2010","journal-title":"Palaeontology"},{"key":"pone.0351939.ref011","doi-asserted-by":"crossref","first-page":"157","DOI":"10.1016\/0031-0182(91)90062-V","article-title":"Prey bone utilization by predatory dinosaurs","volume":"88","author":"AR Fiorillo","year":"1991","journal-title":"Palaeogeogr Palaeoclimatol Palaeoecol"},{"issue":"1","key":"pone.0351939.ref012","doi-asserted-by":"crossref","first-page":"175","DOI":"10.1080\/02724634.1996.10011297","article-title":"Bite marks attributable to Tyrannosaurus rex: preliminary description and implications","volume":"16","author":"GM Erickson","year":"1996","journal-title":"J Vertebr Paleontol"},{"key":"pone.0351939.ref013","first-page":"135","article-title":"Evidence of predatory behavior by carnivorous dinosaurs","volume":"15","author":"K Carpenter","year":"1998","journal-title":"GAIA"},{"key":"pone.0351939.ref014","doi-asserted-by":"crossref","first-page":"325","DOI":"10.2979\/3541.0","article-title":"Evidence for predator-prey relationships: examples for Allosaurus and Stegosaurus.","volume-title":"The carnivorous dinosaurs","author":"K Carpenter","year":"2005"},{"key":"pone.0351939.ref015","first-page":"355","article-title":"An analysis of predator-prey behavior in a head-to-head encounter between Tyrannosaurus rex and Triceratops.","volume-title":"Tyrannosaurus rex the Tyrant king","author":"J Happ","year":"2008"},{"key":"pone.0351939.ref016","doi-asserted-by":"crossref","first-page":"186","DOI":"10.1016\/j.cretres.2012.03.018","article-title":"Tyrannosaur feeding traces on Deinocheirus (Theropoda:? Ornithomimosauria) remains from the Nemegt Formation (Late Cretaceous), Mongolia","volume":"37","author":"PR Bell","year":"2012","journal-title":"Cretac Res"},{"issue":"4","key":"pone.0351939.ref017","doi-asserted-by":"crossref","first-page":"971","DOI":"10.1671\/0272-4634(2004)024[0971:BMOACF]2.0.CO;2","article-title":"Bite marks on a crocodylomorph from the Upper Cretaceous of Brazil: evidence of social behavior?","volume":"24","author":"LS Avilla","year":"2004","journal-title":"J Vertebr Paleontol"},{"issue":"1","key":"pone.0351939.ref018","doi-asserted-by":"crossref","first-page":"12","DOI":"10.1017\/pab.2021.29","article-title":"Intraspecific facial bite marks in tyrannosaurids provide insight into sexual maturity and evolution of bird-like intersexual display","volume":"48","author":"CM Brown","year":"2021","journal-title":"Paleobiology"},{"issue":"6931","key":"pone.0351939.ref019","doi-asserted-by":"crossref","first-page":"515","DOI":"10.1038\/nature01532","article-title":"Cannibalism in the Madagascan dinosaur Majungatholus atopus","volume":"422","author":"RR Rogers","year":"2003","journal-title":"Nature"},{"issue":"5","key":"pone.0351939.ref020","doi-asserted-by":"crossref","DOI":"10.1371\/journal.pone.0233115","article-title":"High frequencies of theropod bite marks provide evidence for feeding, scavenging, and possible cannibalism in a stressed Late Jurassic ecosystem","volume":"15","author":"SK Drumheller","year":"2020","journal-title":"PLoS One"},{"issue":"10","key":"pone.0351939.ref021","doi-asserted-by":"crossref","DOI":"10.1371\/journal.pone.0013419","article-title":"Cannibalism in Tyrannosaurus rex","volume":"5","author":"NR Longrich","year":"2010","journal-title":"PLoS One"},{"issue":"4","key":"pone.0351939.ref022","doi-asserted-by":"crossref","first-page":"164","DOI":"10.2110\/palo.2017.076","article-title":"Tyrannosaur cannibalism: a case of a tooth-traced tyrannosaurid bone in the Lance Formation (Maastrichtian), Wyoming","volume":"33","author":"MA McLain","year":"2018","journal-title":"Palaios"},{"key":"pone.0351939.ref023","first-page":"39","article-title":"New evidence for cannibalism in tyrannosaurid dinosaurs from the Upper Cretaceous (Campanian\/Maastrichtian) San Juan Basin of New Mexico","volume":"82","author":"SG Dalman","year":"2021","journal-title":"N M Mus Nat Hist Sci Bull"},{"key":"pone.0351939.ref024","doi-asserted-by":"crossref","first-page":"35","DOI":"10.1016\/B978-0-12-100036-3.50014-6","article-title":"Patterns of bone modifications produced by nonhuman agents.","volume-title":"Bones: ancient men and modern myths","author":"LR Binford","year":"1981"},{"issue":"2","key":"pone.0351939.ref025","doi-asserted-by":"crossref","first-page":"142","DOI":"10.1016\/j.jhevol.2005.08.008","article-title":"A diagnosis of crocodile feeding traces on larger mammal bone, with fossil examples from the Plio-Pleistocene Olduvai Basin, Tanzania","volume":"50","author":"JK Njau","year":"2006","journal-title":"J Hum Evol"},{"issue":"6","key":"pone.0351939.ref026","doi-asserted-by":"crossref","first-page":"614","DOI":"10.1016\/j.jhevol.2006.11.007","article-title":"Taphonomic analysis of skeletal remains from chimpanzee hunts at Ngogo, Kibale National Park, Uganda","volume":"52","author":"BL Pobiner","year":"2007","journal-title":"J Hum Evol"},{"key":"pone.0351939.ref027","first-page":"373","article-title":"Paleoecological information in predator tooth marks","volume":"6","author":"B Pobiner","year":"2008","journal-title":"J Taphon"},{"issue":"3","key":"pone.0351939.ref028","doi-asserted-by":"crossref","first-page":"113","DOI":"10.1080\/10420940600850729","article-title":"Three new ichnogenera of biting and gnawing traces on reptilian and mammalian bones: a case study from the Miocene of the Czech Republic","volume":"13","author":"R Mikul\u00e1\u0161","year":"2006","journal-title":"Ichnos"},{"issue":"4","key":"pone.0351939.ref029","first-page":"373","article-title":"New ichnotaxa based on tooth impressions on dinosaur and whale bones","volume":"53","author":"AR Jacobsen","year":"2009","journal-title":"Geol Q"},{"key":"pone.0351939.ref030","first-page":"287","article-title":"Tyrannosaurid pathologies as clues to nature and nurture in the Cretaceous.","volume-title":"Tyrannosaurus rex, the tyrant king","author":"BM Rothschild","year":"2008"},{"issue":"3","key":"pone.0351939.ref031","first-page":"92","article-title":"Paleopathology of a mature Tyrannosaurus rex skeleton","volume":"21","author":"EA Rega","year":"2001","journal-title":"J Vertebr Paleontol"},{"issue":"9","key":"pone.0351939.ref032","article-title":"Common avian infection plagued the tyrant dinosaurs","volume":"4","author":"EDS Wolff","year":"2009","journal-title":"PLoS One"},{"key":"pone.0351939.ref033","first-page":"666","article-title":"Disease in dinosaurs.","volume-title":"The complete dinosaur","author":"EA. Rega","year":"2012"},{"issue":"4","key":"pone.0351939.ref034","article-title":"Analysis of pseudopathologies in Edmontosaurus annectens bones: taphonomic implications from biogenetic and diagenetic bone alterations from a Cretaceous bonebed in the Lance Formation, Wyoming","volume":"45","author":"BCT Siviero","year":"2025","journal-title":"J Vertebr Paleontol"},{"issue":"4","key":"pone.0351939.ref035","doi-asserted-by":"crossref","first-page":"201","DOI":"10.2110\/palo.2019.079","article-title":"Skeletal trauma with implications for intratail mobility in Edmontosaurus annectens from a monodominant bonebed, Lance Formation (Maastrichtian), Wyoming USA","volume":"35","author":"BC Siviero","year":"2020","journal-title":"Palaios"},{"issue":"5","key":"pone.0351939.ref036","doi-asserted-by":"crossref","DOI":"10.1371\/journal.pone.0233182","article-title":"Over 13,000 elements from a single bonebed help elucidate disarticulation and transport of an Edmontosaurus thanatocoenosis","volume":"15","author":"K Snyder","year":"2020","journal-title":"PLoS One"},{"issue":"1","key":"pone.0351939.ref037","doi-asserted-by":"crossref","first-page":"1","DOI":"10.3390\/fossils2010001","article-title":"Taxonomic status of Nanotyrannus lancensis (Dinosauria: Tyrannosauroidea)\u2014a distinct taxon of small-bodied tyrannosaur","volume":"2","author":"NR Longrich","year":"2024","journal-title":"Foss Stud"},{"issue":"8093","key":"pone.0351939.ref038","doi-asserted-by":"crossref","first-page":"357","DOI":"10.1038\/s41586-025-09801-6","article-title":"Nanotyrannus and Tyrannosaurus coexisted at the close of the Cretaceous","volume":"648","author":"LE Zanno","year":"2025","journal-title":"Nature"},{"issue":"6782","key":"pone.0351939.ref039","doi-asserted-by":"crossref","first-page":"300","DOI":"10.1126\/science.adx8706","article-title":"A diminutive tyrannosaur lived alongside Tyrannosaurus rex","volume":"391","author":"CT Griffin","year":"2026","journal-title":"Science"},{"issue":"1","key":"pone.0351939.ref040","doi-asserted-by":"crossref","first-page":"79","DOI":"10.1666\/09079.1","article-title":"Using striated tooth marks on bone to predict body size in theropod dinosaurs: a model based on feeding observations of Varanus komodoensis, the Komodo monitor","volume":"38","author":"DC D\u2019Amore","year":"2012","journal-title":"Paleobiology"},{"issue":"2","key":"pone.0351939.ref041","doi-asserted-by":"crossref","first-page":"699","DOI":"10.1002\/ar.a.20206","article-title":"Dental morphology and variation in theropod dinosaurs: implications for the taxonomic identification of isolated teeth","volume":"285","author":"JB Smith","year":"2005","journal-title":"Anat Rec A Discov Mol Cell Evol Biol"},{"issue":"1","key":"pone.0351939.ref042","first-page":"1","article-title":"Standardizing terms for crocodile-induced bite marks on bone surfaces in light of the frequent bone modification equifinality found to result from crocodile feeding behavior, stone tool modification, and trampling","volume":"3","author":"J Njau","year":"2016","journal-title":"FOROST Occas Publ"},{"key":"pone.0351939.ref043","doi-asserted-by":"crossref","first-page":"438","DOI":"10.1525\/california\/9780520242098.003.0023","article-title":"Hadrosauridae.","volume-title":"The Dinosauria","author":"HR Horner","year":"2004","edition":"2"},{"issue":"2","key":"pone.0351939.ref044","doi-asserted-by":"crossref","first-page":"79","DOI":"10.1016\/S0196-0644(80)80335-0","article-title":"Mammalian bite wounds","volume":"9","author":"RV Aghababian","year":"1980","journal-title":"Ann Emerg Med"},{"key":"pone.0351939.ref045","doi-asserted-by":"crossref","first-page":"90","DOI":"10.1016\/j.quaint.2012.12.035","article-title":"Osteomyelitis in Quaternary mammal from the Rio Grande do Norte State, Brazil","volume":"299","author":"FH de Souza Barbosa","year":"2013","journal-title":"Quat Int"},{"issue":"1","key":"pone.0351939.ref046","doi-asserted-by":"crossref","first-page":"1","DOI":"10.1016\/0021-9975(88)90101-6","article-title":"Comparative pathology of inflammation in the higher vertebrates (reptiles, birds and mammals)","volume":"99","author":"RJ Montali","year":"1988","journal-title":"J Comp Pathol"},{"key":"pone.0351939.ref047","first-page":"149","article-title":"Osteomyelitis in reptiles","author":"N Antinoff","year":"1997","journal-title":"Proc ARAV"},{"issue":"1","key":"pone.0351939.ref048","doi-asserted-by":"crossref","first-page":"30","DOI":"10.1647\/2003-007","article-title":"Avian mycobacteriosis in free-living raptors in California: 6 cases (1997\u20132001)","volume":"18","author":"LA Tell","year":"2004","journal-title":"J Avian Med Surg"},{"issue":"1","key":"pone.0351939.ref049","doi-asserted-by":"crossref","first-page":"76","DOI":"10.1671\/0272-4634(2002)022[0076:MIAIIA]2.0.CO;2","article-title":"Multiple injury and infection in a sub-adult theropod dinosaur Allosaurus fragilis with comparisons to allosaur pathology in the Cleveland-Lloyd Dinosaur Quarry Collection","volume":"22","author":"RR Hanna","year":"2002","journal-title":"J Vertebr Paleontol"},{"issue":"1","key":"pone.0351939.ref050","doi-asserted-by":"crossref","first-page":"5045","DOI":"10.1038\/s41598-018-23451-x","article-title":"Possible bite-induced abscess and osteomyelitis in Lufengosaurus (Dinosauria: sauropodomorph) from the Lower Jurassic of the Yimen Basin, China","volume":"8","author":"L Xing","year":"2018","journal-title":"Sci Rep"},{"key":"pone.0351939.ref051","first-page":"171","article-title":"Osteochondrosis in Late Cretaceous Hadrosauria: a manifestation of ontologic failure.","volume-title":"Horns and beaks: ceratopsian and ornithopod dinosaurs","author":"BM Rothschild","year":"2007"},{"issue":"31","key":"pone.0351939.ref052","doi-asserted-by":"crossref","first-page":"12560","DOI":"10.1073\/pnas.1216534110","article-title":"Physical evidence of predatory behavior in Tyrannosaurus rex","volume":"110","author":"RA DePalma","year":"2013","journal-title":"Proc Natl Acad Sci U S A"},{"issue":"1","key":"pone.0351939.ref053","doi-asserted-by":"crossref","first-page":"195","DOI":"10.1666\/11-058","article-title":"Ichnotaxobases for bioerosion trace fossils in bones","volume":"88","author":"CA Pirrone","year":"2014","journal-title":"J Paleontol"},{"issue":"6","key":"pone.0351939.ref054","doi-asserted-by":"crossref","first-page":"565","DOI":"10.1669\/0883-1351(2004)019<0565:BBIPCA>2.0.CO;2","article-title":"Bivalve borings in phosphatic coprolites and bone, Cretaceous-Paleogene, Northeastern Mali","volume":"19","author":"L Tapanila","year":"2004","journal-title":"PALAIOS"},{"key":"pone.0351939.ref055","doi-asserted-by":"crossref","first-page":"350","DOI":"10.1016\/B978-044452949-7\/50146-7","article-title":"Macroborings and the evolution of marine bioerosion.","volume-title":"Trace fossils: concepts, problems, prospects","author":"MA. Wilson","year":"2007"},{"key":"pone.0351939.ref056","author":"FCGD Lima","year":"2017","journal-title":"Altera\u00e7\u00f5es ante mortem e pseudopatologias em exoesqueleto de cingulados pleistoc\u00eanicos de grande porte da regi\u00e3o intertropical brasileira"},{"key":"pone.0351939.ref057","first-page":"1","article-title":"Osteopathology and insect traces in the Australopithecus africanus skeleton StW 431","volume":"113","author":"EJ Odes","year":"2017","journal-title":"S Afr J Sci"},{"issue":"2","key":"pone.0351939.ref058","doi-asserted-by":"crossref","first-page":"122","DOI":"10.1097\/PAF.0000000000000449","article-title":"Depressions of Dermestes maculatus (Coleoptera: Dermestidae) on bones could be pupation chambers","volume":"40","author":"NI Zanetti","year":"2019","journal-title":"Am J Forensic Med Pathol"},{"issue":"6056","key":"pone.0351939.ref059","doi-asserted-by":"crossref","first-page":"768","DOI":"10.1038\/319768a0","article-title":"Trampling as a cause of bone surface damage and pseudo-cutmarks","volume":"319","author":"AK Behrensmeyer","year":"1986","journal-title":"Nature"},{"issue":"5","key":"pone.0351939.ref060","doi-asserted-by":"crossref","first-page":"535","DOI":"10.1016\/0305-4403(88)90081-7","article-title":"Surface modification on bone: trampling versus butchery","volume":"15","author":"SL Olsen","year":"1988","journal-title":"J Archaeol Sci"},{"key":"pone.0351939.ref061","first-page":"61","article-title":"An experimental study of trampling: implications for the fossil record.","volume-title":"Bone modification","author":"AR Fiorillo","year":"1989"},{"issue":"12","key":"pone.0351939.ref062","doi-asserted-by":"crossref","first-page":"2643","DOI":"10.1016\/j.jas.2009.07.017","article-title":"A new protocol to differentiate trampling marks from butchery cut marks","volume":"36","author":"M Dom\u00ednguez-Rodrigo","year":"2009","journal-title":"J Archaeol Sci"},{"issue":"1","key":"pone.0351939.ref063","doi-asserted-by":"crossref","first-page":"39","DOI":"10.4000\/quaternaire.8593","article-title":"Trampling experiments on bones in fine and soft sediments","volume":"29","author":"L Rozada","year":"2018","journal-title":"Quaternaire"},{"key":"pone.0351939.ref064","first-page":"303249","article-title":"Refinement of tooth trace criteria through experimentation and literature review","volume":"49","author":"BT Siviero","year":"2017","journal-title":"Geol Soc Am Abstr Programs"}],"container-title":["PLOS One"],"original-title":[],"language":"en","link":[{"URL":"https:\/\/dx.plos.org\/10.1371\/journal.pone.0351939","content-type":"unspecified","content-version":"vor","intended-application":"similarity-checking"}],"deposited":{"date-parts":[[2026,7,15]],"date-time":"2026-07-15T17:22:19Z","timestamp":1784136139000},"score":1,"resource":{"primary":{"URL":"https:\/\/dx.plos.org\/10.1371\/journal.pone.0351939"}},"subtitle":[],"editor":[{"given":"Furqan A.","family":"Shah","sequence":"first","affiliation":[],"role":[{"vocabulary":"crossref","role":"editor"}]}],"short-title":[],"issued":{"date-parts":[[2026,7,15]]},"references-count":64,"journal-issue":{"issue":"7","published-online":{"date-parts":[[2026,7,15]]}},"URL":"https:\/\/doi.org\/10.1371\/journal.pone.0351939","relation":{},"ISSN":["1932-6203"],"issn-type":[{"value":"1932-6203","type":"electronic"}],"subject":[],"published":{"date-parts":[[2026,7,15]]}}}