{"status":"ok","message-type":"work","message-version":"1.0.0","message":{"indexed":{"date-parts":[[2026,7,10]],"date-time":"2026-07-10T18:18:51Z","timestamp":1783707531125,"version":"3.55.0"},"reference-count":95,"publisher":"Public Library of Science (PLoS)","issue":"7","license":[{"start":{"date-parts":[[2026,7,10]],"date-time":"2026-07-10T00:00:00Z","timestamp":1783641600000},"content-version":"vor","delay-in-days":0,"URL":"http:\/\/creativecommons.org\/licenses\/by\/4.0\/"}],"funder":[{"DOI":"10.13039\/501100004633","name":"Mount Allison University","doi-asserted-by":"publisher","award":["JEA Crake Foundation Grant"],"award-info":[{"award-number":["JEA Crake Foundation Grant"]}],"id":[{"id":"10.13039\/501100004633","id-type":"DOI","asserted-by":"publisher"}]},{"DOI":"10.13039\/501100004633","name":"Mount Allison University","doi-asserted-by":"publisher","award":["Part-time Research Stipend"],"award-info":[{"award-number":["Part-time Research Stipend"]}],"id":[{"id":"10.13039\/501100004633","id-type":"DOI","asserted-by":"publisher"}]},{"DOI":"10.13039\/501100000038","name":"Natural Sciences and Engineering Research Council of Canada","doi-asserted-by":"publisher","award":["Discovery Grant"],"award-info":[{"award-number":["Discovery Grant"]}],"id":[{"id":"10.13039\/501100000038","id-type":"DOI","asserted-by":"publisher"}]}],"content-domain":{"domain":["www.plosone.org"],"crossmark-restriction":false},"short-container-title":["PLoS One"],"abstract":"<jats:p>\n                    Biological invasions provide unique opportunities to examine how populations cope with rapid ecological change, as well as the influence of human activity on our natural world. The island of Newfoundland, Canada, has no native amphibians; however, five species have established non-native populations, including one caudatan, the Eastern Red-backed Salamander (\n                    <jats:italic>Plethodon cinereus<\/jats:italic>\n                    ). We posit that this salamander\u2019s successful colonisation may have been mediated by dietary shifts that allowed them to capitalize on a greater variety of prey, as well as by exploiting pre-existing invasive prey communities \u2013 both these mechanisms could limit the likelihood of competitive exclusion and promote establishment in a novel environment. To examine this, we identified stomach contents of Newfoundland Eastern Red-backed salamanders, characterised the prey as native or invasive, and compared the dietary composition of the invasive population to that of conspecifics in the native range through a systematic literature review. As predicted, the invasive salamander population\u2019s diet is more generalised (i.e., a broader dietary niche) than the native salamanders, and invasive invertebrates comprised two thirds of the volume of prey eaten by invasive salamanders. Our research provides insight into the Niche Breadth Invasion Hypothesis as well as the Invasional Meltdown Hypothesis, which are mechanisms that may have allowed this species to establish a population in an island ecosystem. Furthermore, our research suggests that prior invasions of a diverse invertebrate community to Newfoundland may have bolstered the invasive potential of this novel, non-native salamander.\n                  <\/jats:p>","DOI":"10.1371\/journal.pone.0352970","type":"journal-article","created":{"date-parts":[[2026,7,10]],"date-time":"2026-07-10T17:54:08Z","timestamp":1783706048000},"page":"e0352970","update-policy":"https:\/\/doi.org\/10.1371\/journal.pone.corrections_policy","source":"Crossref","is-referenced-by-count":0,"title":["In the belly of the beast: How dietary changes and preexisting invasive prey may have promoted the success of a novel invasive amphibian"],"prefix":"10.1371","volume":"21","author":[{"given":"Maya J.","family":"Williams","sequence":"first","affiliation":[],"role":[{"vocabulary":"crossref","role":"author"}]},{"given":"Julia L.","family":"Riley","sequence":"additional","affiliation":[],"role":[{"vocabulary":"crossref","role":"author"}]},{"ORCID":"https:\/\/orcid.org\/0000-0002-1283-8893","authenticated-orcid":true,"given":"James","family":"Baxter-Gilbert","sequence":"additional","affiliation":[],"role":[{"vocabulary":"crossref","role":"author"}]}],"member":"340","published-online":{"date-parts":[[2026,7,10]]},"reference":[{"issue":"9","key":"pone.0352970.ref001","doi-asserted-by":"crossref","first-page":"470","DOI":"10.1016\/j.tree.2004.07.005","article-title":"Are invasive species a major cause of extinctions?","volume":"19","author":"J Gurevitch","year":"2004","journal-title":"Trends Ecol Evol"},{"key":"pone.0352970.ref002","doi-asserted-by":"crossref","first-page":"1","DOI":"10.1016\/bs.aecr.2016.10.001","article-title":"Impacts of invasive species on food webs: A review of empirical data","volume":"56","author":"P David","year":"2017","journal-title":"Advances in Ecological 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cane toad: The history and ecology of a successful colonist","author":"C Lever","year":"2001"},{"key":"pone.0352970.ref007","volume-title":"100 of the world\u2019s worst invasive alien species: A selection from the global invasive species database","author":"S Lowe","year":"2000"},{"issue":"3","key":"pone.0352970.ref008","doi-asserted-by":"crossref","first-page":"253","DOI":"10.1086\/655116","article-title":"The ecological impact of invasive cane toads (Bufo marinus) in Australia","volume":"85","author":"R Shine","year":"2010","journal-title":"Q Rev Biol"},{"issue":"1","key":"pone.0352970.ref009","doi-asserted-by":"crossref","first-page":"1","DOI":"10.1111\/aec.12066","article-title":"A review of ecological interactions between native frogs and invasive cane toads in Australia","volume":"39","author":"R Shine","year":"2013","journal-title":"Austral 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salamander, plethodon cinereus","volume":"43","author":"H Heatwole","year":"1962","journal-title":"Ecology"},{"key":"pone.0352970.ref046","doi-asserted-by":"crossref","first-page":"186","DOI":"10.1016\/j.foreco.2006.04.034","article-title":"Effects of structural complexity enhancement on eastern red-backed salamander (Plethodon cinereus) populations in northern hardwood forests","volume":"230","author":"HC McKenny","year":"2006","journal-title":"For Ecol Manage"},{"issue":"1","key":"pone.0352970.ref047","doi-asserted-by":"crossref","first-page":"30","DOI":"10.1674\/0003-0031-168.1.30","article-title":"Interactions among forest-floor guild members in structurally simple microhabitats","volume":"168","author":"CA Hickerson","year":"2012","journal-title":"Am Midl Nat"},{"key":"pone.0352970.ref048","article-title":"Eastern Red-backed salamanders regulate top-down effects in a temperate forest-floor community","author":"C-AM Hickerson","year":"2017","journal-title":"Herpetologica"},{"issue":"3","key":"pone.0352970.ref049","doi-asserted-by":"crossref","first-page":"433","DOI":"10.1007\/BF00317613","article-title":"Territories of male and female terrestrial salamanders: Costs, benefits, and intersexual spatial associations","volume":"86","author":"A Mathis","year":"1991","journal-title":"Oecologia"},{"issue":"2","key":"pone.0352970.ref050","doi-asserted-by":"crossref","first-page":"270","DOI":"10.2307\/1443413","article-title":"The effectiveness of antipredator secretions and behavior of selected salamanders against shrews","volume":"1979","author":"ED Brodie","year":"1979","journal-title":"Copeia"},{"issue":"4","key":"pone.0352970.ref051","doi-asserted-by":"crossref","DOI":"10.21425\/F59433282","article-title":"Potential change in the distribution of an abundant and wide-ranging forest salamander in a context of climate change","volume":"9","author":"JD Moore","year":"2018","journal-title":"Front 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CRAN. 2019. 235\u20136."},{"issue":"1","key":"pone.0352970.ref076","doi-asserted-by":"crossref","first-page":"1","DOI":"10.18637\/jss.v033.i01","article-title":"Regularization paths for generalized linear models via coordinate descent","volume":"33","author":"J Friedman","year":"2010","journal-title":"J Stat Softw"},{"issue":"61","key":"pone.0352970.ref077","doi-asserted-by":"crossref","first-page":"3167","DOI":"10.21105\/joss.03167","article-title":"Visualizations with statistical details: The \u201cggstatsplot\u201d approach","volume":"6","author":"I Patil","year":"2021","journal-title":"JOSS"},{"issue":"9","key":"pone.0352970.ref078","doi-asserted-by":"crossref","first-page":"2633","DOI":"10.1007\/s10530-017-1473-6","article-title":"Let\u2019s eat out, there\u2019s crayfish for dinner: American bullfrog niche shifts inside and outside native ranges and the effect of introduced crayfish","volume":"19","author":"AM Bissattini","year":"2017","journal-title":"Biol Invasions"},{"issue":"1","key":"pone.0352970.ref079","first-page":"34","article-title":"Diet and prey selection of the invasive American bullfrog (Lithobates catesbeianus) in southwestern China","volume":"6","author":"X Liu","year":"2015","journal-title":"Asian Herpetol Res"},{"issue":"10","key":"pone.0352970.ref080","doi-asserted-by":"crossref","DOI":"10.1002\/ecs2.3789","article-title":"Of olives and carp: Interactive effects of an aquatic and a terrestrial invader on a stream\u2010riparian ecosystem","volume":"12","author":"KK Heinrich","year":"2021","journal-title":"Ecosphere"},{"key":"pone.0352970.ref081","volume-title":"Invasion meltdown: investigating mutual facilitation across ecosystem boundaries","author":"CA Cheek","year":"2019"},{"key":"pone.0352970.ref082","unstructured":"Massicotte P, South A, Hufkens K. rnaturalearth: World map data from natural earth. R package version. 2023. https:\/\/CRAN.R-project.org\/package=rnaturalearth"},{"key":"pone.0352970.ref083","doi-asserted-by":"crossref","first-page":"189","DOI":"10.1007\/978-3-319-24277-4_9","article-title":"Data analysis","volume-title":"ggplot2: elegant graphics for data analysis","author":"H Wickham","year":"2016"},{"key":"pone.0352970.ref084","unstructured":"NatureServe and IUCN (International Union for Conservation of Nature). 2022.Plethodon cinereus. IUCN Red List of Threatened Species. https:\/\/www.iucnredlist.org. Accessed 2023 December 18."},{"key":"pone.0352970.ref085","unstructured":"Yang C. Island life of salamanders: the genetic origin and behavioural traits of the introduced eastern red-backed salamander (Plethodon cinereus) in Newfoundland, Canada. 2026."},{"issue":"2","key":"pone.0352970.ref086","doi-asserted-by":"crossref","first-page":"281","DOI":"10.1643\/0045-8511(2007)7[281:DOTIFE]2.0.CO;2","article-title":"Diet of the Invasive Frog, Eleutherodactylus Coqui, in Hawaii","volume":"2007","author":"KH Beard","year":"2007","journal-title":"Copeia"},{"issue":"1","key":"pone.0352970.ref087","doi-asserted-by":"crossref","first-page":"168","DOI":"10.1111\/aje.12814","article-title":"Toad\u2010kill: Prey diversity and preference of invasive guttural toads (Sclerophrys gutturalis) in Mauritius","volume":"59","author":"J Baxter\u2010Gilbert","year":"2021","journal-title":"Afr J Ecol"},{"key":"pone.0352970.ref088","volume-title":"Trophic interactions of the Guttural Toad (Sclerophrys gutturalis, Power 1927) along an invasion gradient","author":"ST Peta","year":"2022"},{"issue":"1","key":"pone.0352970.ref089","doi-asserted-by":"crossref","first-page":"32","DOI":"10.1111\/j.1365-2486.2010.02247.x","article-title":"What makes an urban bird?","volume":"17","author":"KL Evans","year":"2010","journal-title":"Global Change Biology"},{"issue":"1","key":"pone.0352970.ref090","doi-asserted-by":"crossref","first-page":"42","DOI":"10.1111\/ibi.12732","article-title":"Urban exploiters have broader dietary niches than urban avoiders","volume":"162","author":"FX Palacio","year":"2020","journal-title":"Ibis"},{"key":"pone.0352970.ref091","doi-asserted-by":"crossref","DOI":"10.1007\/978-3-030-34721-5","volume-title":"The ecology of invasions by animals and plants","author":"CS Elton","year":"2020"},{"issue":"4","key":"pone.0352970.ref092","doi-asserted-by":"crossref","first-page":"627","DOI":"10.1046\/j.1365-294X.2002.01471.x","article-title":"Molecular identification of prey in predator diets","volume":"11","author":"WOC Symondson","year":"2002","journal-title":"Mol Ecol"},{"issue":"5","key":"pone.0352970.ref093","doi-asserted-by":"crossref","first-page":"1302","DOI":"10.1111\/1365-2664.14141","article-title":"Hotspots of pest\u2010induced US urban tree death, 2020\u20132050","volume":"59","author":"EJ Hudgins","year":"2022","journal-title":"J Appl Ecol"},{"issue":"3","key":"pone.0352970.ref094","doi-asserted-by":"crossref","first-page":"273","DOI":"10.1016\/j.ecolecon.2004.10.002","article-title":"Update on the environmental and economic costs associated with alien-invasive species in the United States","volume":"52","author":"D Pimentel","year":"2005","journal-title":"Ecol Econ"},{"key":"pone.0352970.ref095","doi-asserted-by":"crossref","first-page":"3557","DOI":"10.1007\/s10530-017-1596-9","article-title":"How do invasive species travel to and through urban environments?","volume":"19","author":"AL Padayachee","year":"2017","journal-title":"Biol Invasions"}],"container-title":["PLOS One"],"original-title":[],"language":"en","link":[{"URL":"https:\/\/dx.plos.org\/10.1371\/journal.pone.0352970","content-type":"unspecified","content-version":"vor","intended-application":"similarity-checking"}],"deposited":{"date-parts":[[2026,7,10]],"date-time":"2026-07-10T17:54:26Z","timestamp":1783706066000},"score":1,"resource":{"primary":{"URL":"https:\/\/dx.plos.org\/10.1371\/journal.pone.0352970"}},"subtitle":[],"editor":[{"given":"Alexandre Ribeiro","family":"da Silva","sequence":"first","affiliation":[],"role":[{"vocabulary":"crossref","role":"editor"}]}],"short-title":[],"issued":{"date-parts":[[2026,7,10]]},"references-count":95,"journal-issue":{"issue":"7","published-online":{"date-parts":[[2026,7,10]]}},"URL":"https:\/\/doi.org\/10.1371\/journal.pone.0352970","relation":{},"ISSN":["1932-6203"],"issn-type":[{"value":"1932-6203","type":"electronic"}],"subject":[],"published":{"date-parts":[[2026,7,10]]}}}