{"status":"ok","message-type":"work","message-version":"1.0.0","message":{"indexed":{"date-parts":[[2026,9,22]],"date-time":"2026-09-22T18:06:17Z","timestamp":1790100377816,"version":"4.0.1"},"reference-count":62,"publisher":"Public Library of Science (PLoS)","issue":"9","license":[{"start":{"date-parts":[[2026,9,22]],"date-time":"2026-09-22T00:00:00Z","timestamp":1790035200000},"content-version":"vor","delay-in-days":0,"URL":"http:\/\/creativecommons.org\/licenses\/by\/4.0\/"}],"funder":[{"DOI":"10.13039\/501100001659","name":"Deutsche Forschungsgemeinschaft","doi-asserted-by":"publisher","award":["EI438\/12-4"],"award-info":[{"award-number":["EI438\/12-4"]}],"id":[{"id":"10.13039\/501100001659","id-type":"DOI","asserted-by":"publisher"}]}],"content-domain":{"domain":["www.plosone.org"],"crossmark-restriction":false},"short-container-title":["PLoS One"],"abstract":"<jats:p>Since the beginning of excavations at G\u00f6beklitepe, a Pre-Pottery Neolithic (PPN) site in southeastern T\u00fcrkiye, fragmented human bones have been recovered from the fill of the special (monumental) buildings. These fragments derive from males and females of all age categories and exhibit various taphonomic alterations. Compared with contemporaneous Anatolian burial customs, certain parallels exist; however, the seemingly random distribution of human bone fragments at G\u00f6beklitepe remains enigmatic. Recently, the first two intact burials were discovered in Pre-Pottery Neolithic B (8,700\u20138,000 BCE) contexts. The taphonomic evidence from these burials could explain the presence of the dispersed human bone fragments. It appears that lower-lying parts of the site became inundated by slope deposits from higher areas of the mound, which contained human remains from originally intact burials. Nonetheless, this explanation does not rule out the possibility of additional mortuary practices, thereby underscoring the considerable diversity of burial rituals in Neolithic Anatolia.<\/jats:p>","DOI":"10.1371\/journal.pone.0358979","type":"journal-article","created":{"date-parts":[[2026,9,22]],"date-time":"2026-09-22T17:30:18Z","timestamp":1790098218000},"page":"e0358979","update-policy":"https:\/\/doi.org\/10.1371\/journal.pone.corrections_policy","source":"Crossref","is-referenced-by-count":0,"title":["First burials give insights into mortuary rites at G\u00f6beklitepe, Early Neolithic T\u00fcrkiye"],"prefix":"10.1371","volume":"21","author":[{"ORCID":"https:\/\/orcid.org\/0000-0003-3493-2757","authenticated-orcid":true,"given":"Julia","family":"Gresky","sequence":"first","affiliation":[{"name":"Georg-August University","place":["G\u00f6ttingen, Germany"],"department":["Faculty of Biology and Psychology"]},{"name":"German Archaeological Institute","place":["Berlin, Germany"],"department":["Division of Natural Sciences"]}],"role":[{"vocabulary":"crossref","role":"author"}]},{"ORCID":"https:\/\/orcid.org\/0000-0001-8187-2078","authenticated-orcid":true,"given":"Lee","family":"Clare","sequence":"additional","affiliation":[{"name":"German Archaeological Institute","place":["Istanbul, T\u00fcrkiye"],"department":["Istanbul Department"]}],"role":[{"vocabulary":"crossref","role":"author"}]}],"member":"340","published-online":{"date-parts":[[2026,9,22]]},"reference":[{"key":"pone.0358979.ref001","first-page":"522","article-title":"G\u00f6beklitepe.","volume-title":"UNESCO World Heritage in Turkey","author":"L Clare","year":"2019"},{"key":"pone.0358979.ref002","first-page":"239","article-title":"G\u00f6beklitepe.","volume-title":"The Neolithic in T\u00fcrkiye. 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Copenhagen; 2023."},{"key":"pone.0358979.ref013","unstructured":"Gresky J, Clare L. Human body parts from the monumental special buildings at Pre-Pottery Neolithic (PPN) G\u00f6beklitepe, Southeast T\u00fcrkiye. 89th Annual Meeting of the Society for American Archaeology; 2024."},{"key":"pone.0358979.ref014","unstructured":"Gresky J, Karul N, Clare L. Un-intentional modifications of the human bone fragments from G\u00f6beklitepe. World Neolithic Congress; 2024 Nov 4\u20138; \u015eanl\u0131urfa, T\u00fcrkiye. 587 p."},{"key":"pone.0358979.ref015","unstructured":"Gresky J, Clare L, Karul N. Intentional modifications of the human bone fragments from G\u00f6beklitepe. 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Lectures in archaeothanatology. 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Archaeologica Euphratica 5","author":"J Peters","year":"2025"},{"key":"pone.0358979.ref040","first-page":"735","article-title":"A bioarchaeological and forensic re-assessment of vulture defleshing and mortuary practices at Neolithic \u00c7atalh\u00f6y\u00fck","volume":"10","author":"MA Pilloud","year":"2016","journal-title":"J Archaeol Sci Rep"},{"key":"pone.0358979.ref041","first-page":"153","article-title":"On the Intersection of the Euphrates and Tigris Basins: Sefertepe.","volume-title":"The Neolithic in T\u00fcrkiye. Vol. 7, The Eastern Taurus","author":"E. G\u00fcldo\u011fan","year":"2025"},{"key":"pone.0358979.ref042","author":"M \u00d6zbek","journal-title":"\u00c7ay\u00f6n\u00fc\u2019de \u0130nsan. Istanbul: Arkeoloji ve Sanat Yay\u0131nlar\u0131"},{"key":"pone.0358979.ref043","first-page":"35","article-title":"\u00c7ay\u00f6n\u00fc.","volume-title":"Neolithic in Turkey: The Cradle of Civilization","author":"A. \u00d6zdo\u011fan","year":"1999"},{"key":"pone.0358979.ref044","unstructured":"Erdal YS. Extensive incisions and delayed burials in the early Neolithic settlements of Upper Euphrates. World Neolithic Congress; 2024 Nov 4\u20138; \u015eanl\u0131urfa, T\u00fcrkiye. 319 p."},{"issue":"1","key":"pone.0358979.ref045","doi-asserted-by":"crossref","first-page":"4","DOI":"10.1179\/1461957114Y.0000000072","article-title":"Bone or flesh: Defleshing and post-depositional treatments at K\u00f6rtik Tepe (Southeastern Anatolia, PPNA period)","volume":"18","author":"YS Erdal","year":"2015","journal-title":"Eur J Archaeol"},{"key":"pone.0358979.ref046","first-page":"73","article-title":"\u201cDomesticating\u201d death: the burial contexts.","volume-title":"Death in Ba\u2019ja: Sepulchral Identity and Symbolism in an Early Neolithic Community of the Transjordanian Highlands. Household and Death in Ba\u2019ja 2","author":"M Benz","year":"2023"},{"key":"pone.0358979.ref047","article-title":"Anthropological and paleopathological investigations.","volume-title":"The Late Neolithic Site of Shir, Syria. Volume I","author":"K. Bartl","year":"2018"},{"issue":"2","key":"pone.0358979.ref048","doi-asserted-by":"crossref","first-page":"203","DOI":"10.1016\/S0047-2484(85)80007-5","article-title":"Cannibalism and burial at Krapina","volume":"14","author":"E Trinkaus","year":"1985","journal-title":"J Hum Evol"},{"key":"pone.0358979.ref049","doi-asserted-by":"crossref","first-page":"44","DOI":"10.4312\/dp.51.23","article-title":"Saybur\u00e7 a mid-9th millennium BC site in the foothills of the Eastern Taurus","volume":"51","author":"E \u00d6zdo\u011fan","year":"2024","journal-title":"Doc Praeh"},{"key":"pone.0358979.ref050","doi-asserted-by":"crossref","first-page":"52","DOI":"10.5744\/bi.2017.1002","article-title":"Skull retrieval and secondary burial practices in the Neolithic Near East: recent insights from \u00c7atalh\u00f6y\u00fck, Turkey","volume":"1","author":"SD Haddow","year":"2017","journal-title":"BI"},{"key":"pone.0358979.ref051","volume-title":"Standards for data collection from human skeletal remains","author":"JE Buikstra","year":"1994"},{"key":"pone.0358979.ref052","volume-title":"History of human life span and mortality","author":"GY Acs\u00e1di","year":"1970"},{"key":"pone.0358979.ref053","first-page":"1","article-title":"Empfehlungen f\u00fcr die Alters- und Geschlechtsdiagnose am Skelett","volume":"30","author":"D Ferembach","year":"1979","journal-title":"Homo"},{"key":"pone.0358979.ref054","first-page":"444","article-title":"Geschlechtsdiagnose am Skelett.","volume-title":"Anthropologie. Handbuch der vergleichenden Biologie des Menschen. Band 1, Teil 1","author":"T. Sj\u00f8vold","year":"1988"},{"key":"pone.0358979.ref055","volume-title":"Digging up bones: the excavation, treatment, and study of human skeletal remains","author":"DR Brothwell","year":"1981"},{"key":"pone.0358979.ref056","first-page":"369","article-title":"Diachronic dental research on human skeletal remains excavated in the Netherlands 1: Dorestad\u2019s cemetery on the Heul","volume":"31","author":"WRK Perizonius","year":"1981","journal-title":"BROB"},{"issue":"1","key":"pone.0358979.ref057","doi-asserted-by":"crossref","first-page":"57","DOI":"10.1002\/ajpa.1330680106","article-title":"Ectocranial suture closure: a revised method for the determination of skeletal age at death based on the lateral-anterior sutures","volume":"68","author":"RS Meindl","year":"1985","journal-title":"Am J Phys Anthropol"},{"key":"pone.0358979.ref058","first-page":"499","article-title":"La dur\u00e9e de la vie chez l\u2019homme fossile","volume":"47","author":"HV Vallois","year":"1937","journal-title":"L\u2019Anthropologie"},{"key":"pone.0358979.ref059","first-page":"421","article-title":"Altersdiagnose am Skelett.","volume-title":"Anthropologie. Band I, 1. Teil","author":"J. Szilvassy","year":"1988"},{"issue":"1","key":"pone.0358979.ref060","first-page":"53","article-title":"Die L\u00e4nge der L\u00e4ngsknochen altslawischer Bev\u00f6lkerungen unter besonderer Ber\u00fccksichtigung von Wachstumsfragen","volume":"29","author":"M Stloukal","year":"1978","journal-title":"Homo"},{"key":"pone.0358979.ref061","article-title":"Developmental juvenile osteology","author":"L Scheuer","year":"2000","journal-title":"Elsevier"},{"key":"pone.0358979.ref062","volume-title":"Human skeletal remains: excavation, analysis, interpretation","author":"DH Ubelaker","year":"1978"}],"container-title":["PLOS One"],"original-title":[],"language":"en","link":[{"URL":"https:\/\/dx.plos.org\/10.1371\/journal.pone.0358979","content-type":"unspecified","content-version":"vor","intended-application":"similarity-checking"}],"deposited":{"date-parts":[[2026,9,22]],"date-time":"2026-09-22T17:30:30Z","timestamp":1790098230000},"score":1,"resource":{"primary":{"URL":"https:\/\/dx.plos.org\/10.1371\/journal.pone.0358979"}},"subtitle":[],"editor":[{"given":"Olga","family":"Spekker","sequence":"first","affiliation":[{"name":"University of Szeged Institute of Biology: Szegedi Tudomanyegyetem Biologia Intezet","place":["HUNGARY"]}],"role":[{"vocabulary":"crossref","role":"editor"}]}],"short-title":[],"issued":{"date-parts":[[2026,9,22]]},"references-count":62,"journal-issue":{"issue":"9","published-online":{"date-parts":[[2026,9,22]]}},"URL":"https:\/\/doi.org\/10.1371\/journal.pone.0358979","relation":{},"ISSN":["1932-6203"],"issn-type":[{"value":"1932-6203","type":"electronic"}],"subject":[],"published":{"date-parts":[[2026,9,22]]}}}