{"status":"ok","message-type":"work","message-version":"1.0.0","message":{"indexed":{"date-parts":[[2026,4,30]],"date-time":"2026-04-30T15:17:27Z","timestamp":1777562247297,"version":"3.51.4"},"reference-count":109,"publisher":"Public Library of Science (PLoS)","issue":"3","license":[{"start":{"date-parts":[[2017,3,8]],"date-time":"2017-03-08T00:00:00Z","timestamp":1488931200000},"content-version":"vor","delay-in-days":0,"URL":"http:\/\/creativecommons.org\/licenses\/by\/4.0\/"}],"content-domain":{"domain":["www.plosone.org"],"crossmark-restriction":false},"short-container-title":["PLoS ONE"],"DOI":"10.1371\/journal.pone.0171919","type":"journal-article","created":{"date-parts":[[2017,3,8]],"date-time":"2017-03-08T18:28:54Z","timestamp":1488997734000},"page":"e0171919","update-policy":"https:\/\/doi.org\/10.1371\/journal.pone.corrections_policy","source":"Crossref","is-referenced-by-count":13,"title":["Two new ootaxa from the late Jurassic: The oldest record of crocodylomorph eggs, from the Lourinh\u00e3 Formation, Portugal"],"prefix":"10.1371","volume":"12","author":[{"ORCID":"https:\/\/orcid.org\/0000-0001-7730-1395","authenticated-orcid":true,"given":"Jo\u00e3o","family":"Russo","sequence":"first","affiliation":[]},{"given":"Oct\u00e1vio","family":"Mateus","sequence":"additional","affiliation":[]},{"given":"Marco","family":"Marzola","sequence":"additional","affiliation":[]},{"given":"Ausenda","family":"Balbino","sequence":"additional","affiliation":[]}],"member":"340","published-online":{"date-parts":[[2017,3,8]]},"reference":[{"key":"ref1","doi-asserted-by":"crossref","first-page":"357","DOI":"10.1146\/annurev.earth.31.100901.141308","article-title":"Phylogenetic approaches toward crocodylian history","volume":"31","author":"CA Brochu","year":"2003","journal-title":"Annu Rev Earth Planet Sci"},{"key":"ref2","first-page":"8","article-title":"Reproductive characteristics of the order Crocodylia","author":"JB Thorbjarnarson","year":"1996","journal-title":"Herpetologica"},{"key":"ref3","doi-asserted-by":"crossref","DOI":"10.1038\/ncomms9438","article-title":"Climate constrains the evolutionary history and biodiversity of crocodylians","volume":"6","author":"PD Mannion","year":"2015","journal-title":"Nat Commun"},{"key":"ref4","doi-asserted-by":"crossref","first-page":"140385","DOI":"10.1098\/rsos.140385","article-title":"Diversification events and the effects of mass extinctions on Crocodyliformes evolutionary history","volume":"2","author":"M Bronzati","year":"2015","journal-title":"R Soc Open Sci"},{"key":"ref5","doi-asserted-by":"crossref","first-page":"20152840","DOI":"10.1098\/rspb.2015.2840","article-title":"Environmental drivers of crocodyliform extinction across the Jurassic\/Cretaceous transition","author":"JP Tennant","year":"2016","journal-title":"Proc R Soc B. The Royal Society"},{"key":"ref6","doi-asserted-by":"crossref","first-page":"470","DOI":"10.1017\/S009483730002011X","article-title":"Crocodilian diversity in space and time: the role of climate in paleoecology and its implication for understanding K\/T extinctions","volume":"24","author":"PJ Markwick","year":"1998","journal-title":"Paleobiology"},{"key":"ref7","doi-asserted-by":"crossref","first-page":"309","DOI":"10.1111\/j.1475-4983.2010.01028.x","article-title":"Crocodylomorph eggs and eggshells from the Adamantina Formation (Bauru Group), Upper Cretaceous of Brazil","volume":"54","author":"CE Oliveira","year":"2011","journal-title":"Palaeontology"},{"key":"ref8","unstructured":"Moreno-Azanza M. Los huevos f\u00f3siles de amniotas del Cret\u00e1cico de la Pen\u00ednsula Ib\u00e9rica: tafonom\u00eda, formaci\u00f3n de la c\u00e1scara y sistem\u00e1tica. PhD Diss Univ Zaragoza. 2014;"},{"key":"ref9","doi-asserted-by":"crossref","first-page":"115","DOI":"10.1080\/08912963.2013.871009","article-title":"Identification and comparison of modern and fossil crocodilian eggs and eggshell structures","volume":"27","author":"M Marzola","year":"2015","journal-title":"Hist Biol"},{"key":"ref10","doi-asserted-by":"crossref","first-page":"446","DOI":"10.2110\/palo.2014.062","article-title":"Rare in situ preservation of adult crocodylian with eggs from the Middle Eocene of Geiseltal, Germany","volume":"30","author":"AK Hastings","year":"2015","journal-title":"Palaios"},{"key":"ref11","doi-asserted-by":"crossref","first-page":"e0144369","DOI":"10.1371\/journal.pone.0144369","article-title":"Crocodilian nest in a Late Cretaceous sauropod hatchery from the type Lameta Ghat locality, Jabalpur, India","volume":"10","author":"R Srivastava","year":"2015","journal-title":"PloS One"},{"key":"ref12","doi-asserted-by":"crossref","first-page":"513","DOI":"10.1016\/j.crpv.2015.05.006","article-title":"Testudoid and crocodiloid eggshells from the Upper Cretaceous Deccan Intertrappean Beds of Central India","volume":"14","author":"GV Prasad","year":"2015","journal-title":"Comptes Rendus Palevol"},{"key":"ref13","article-title":"Classification of fossil eggshells of amniotic vertebrates","volume":"36","author":"KE Mikhailov","year":"1991","journal-title":"Acta Palaeontol Pol"},{"key":"ref14","author":"KE Mikhailov","year":"1997"},{"key":"ref15","unstructured":"Marzola M, Mateus O, Russo J, Mil\u00e1n J. Comparison of modern and fossil Crocodylomorpha eggs and contribution to the oophylogeny of Amniota. XII Annual Meeting of the European Association of Vertebrate Palaeontologists, Museo Regionale di Scienze Naturali, Regione Piemonte, Torino. Delfino, M., Carnevale, G., Pavia, M.; 2014. p. 109."},{"key":"ref16","first-page":"55","article-title":"Fossile Eischalenreste aus der eoz\u00e4nen Braunkohle des Geiseltales bei Halle-Saale","volume":"1","author":"F Heller","year":"1931","journal-title":"Nova Acta Leopoldina N F"},{"key":"ref17","doi-asserted-by":"crossref","first-page":"17","DOI":"10.1007\/BF00421674","article-title":"Beitr\u00e4ge zur kenntnis der krokodileier, insbesondere ihrer kalkschale","volume":"40","author":"WJ Schmidt","year":"1943","journal-title":"Z F\u00fcr Morphol \u00d6kol Tiere"},{"key":"ref18","unstructured":"Erben HK, M\u00fcnzenburg HK. Ultrastrukturen und Mineralisation rezenter und fossiler Eischalen bei V\u00f6geln und Reptilien. Schattaur; 1970."},{"key":"ref19","first-page":"32","article-title":"Kristalline bausteine und mineralbestand von kalkigen eischalen","volume":"6","author":"HK Erben","year":"1972","journal-title":"Biomineralisation"},{"key":"ref20","doi-asserted-by":"crossref","first-page":"259","DOI":"10.1080\/10292389509380524","article-title":"Reflections on the origin of the amniote egg in the light of reproductive strategies and shell structure","volume":"10","author":"RR Kohring","year":"1995","journal-title":"Hist Biol"},{"key":"ref21","author":"K Carpenter","year":"1999"},{"key":"ref22","doi-asserted-by":"crossref","first-page":"291","DOI":"10.1016\/B978-012676460-4\/50010-X","article-title":"Morphology and evolution of the egg of oviparous amniotes","author":"JR Stewart","year":"1997","journal-title":"Amniote Orig"},{"key":"ref23","doi-asserted-by":"crossref","first-page":"67","DOI":"10.1080\/02724634.1996.10011285","article-title":"Crocodilian and avian eggshells from the Middle Eocene of the Geiseltal, Eastern Germany","volume":"16","author":"R Kohring","year":"1996","journal-title":"J Vertebr Paleontol"},{"key":"ref24","first-page":"136","article-title":"Structure of eggshells and water relations of reptilian eggs","author":"MJ Packard","year":"1982","journal-title":"Herpetologica"},{"key":"ref25","first-page":"329","article-title":"Reproductive biology and embryology of the crocodilians","volume":"14","author":"MW Ferguson","year":"1985","journal-title":"Biol Reptil"},{"key":"ref26","first-page":"531","article-title":"Fossil crocodilian eggs from the Eocene of Colorado","author":"KF Hirsch","year":"1985","journal-title":"J Paleontol"},{"key":"ref27","doi-asserted-by":"crossref","first-page":"53","DOI":"10.1017\/CBO9780511585739.006","article-title":"Eggshell structure and formation in eggs of oviparous reptiles","author":"MJ Packard","year":"1991","journal-title":"Egg Incubation Its Eff Embryonic Dev Birds Reptil Camb Univ Press Camb"},{"key":"ref28","doi-asserted-by":"crossref","first-page":"763","DOI":"10.1080\/02724634.1996.10011364","article-title":"Parataxonomy of fossil egg remains (Veterovata): principles and applications","volume":"16","author":"KE Mikhailov","year":"1996","journal-title":"J Vertebr Paleontol"},{"key":"ref29","first-page":"105","article-title":"Microstructure of the dinosaurian eggshells of Nanxiong, Guangdong, and the problems in dinosaur egg classification","volume":"13","author":"Z Zhao","year":"1975","journal-title":"Vertebr Palasiat"},{"key":"ref30","doi-asserted-by":"crossref","first-page":"e1185432","DOI":"10.1080\/02724634.2016.1185432","article-title":"Fossil egg and eggshells from the Upper Cretaceous Hell Creek Formation, Montana","volume":"36","author":"FD Jackson","year":"2016","journal-title":"J Vertebr Paleontol"},{"key":"ref31","doi-asserted-by":"crossref","first-page":"617","DOI":"10.1016\/j.cretres.2015.06.019","article-title":"Enigmatic Early Cretaceous ootaxa from Western Europe with signals of extrinsic eggshell degradation","volume":"56","author":"M Moreno-Azanza","year":"2015","journal-title":"Cretac Res"},{"key":"ref32","first-page":"79","article-title":"Reptile eggshell from the Purbeck Limestone Group of Dorset, southern England","author":"PC Ensom","year":"1997","journal-title":"Proceedings of the Dorset Natural History and Archaeological Society"},{"key":"ref33","first-page":"221","article-title":"Reptile eggshell, tiny vertebrate remains and globular calcified cartilage from the Purbeck Limestone Group of southern England","volume":"68","author":"PC Ensom","year":"2002","journal-title":"Spec Pap Palaeontol"},{"key":"ref34","article-title":"Fossile Krokodil-Eischalen aus der Unteren S\u00fc\\s swasser-Molasse (Untermioz\u00e4n) von Ulm (Ulm-Westtangente, Baden-W\u00fcrttemberg): Fossil Crocodilian eggshells from the Lower Freshwater Molasse (Early Miocene) of Ulm (Ulm-Westtangente, Baden-W\u00fcrttemberg)","author":"R Kohring","year":"1992","journal-title":"Staatl. Museum f\u00fcr Naturkunde"},{"key":"ref35","doi-asserted-by":"crossref","first-page":"195","DOI":"10.1080\/08912963.2013.786067","article-title":"A re-evaluation of aff. Megaloolithidae eggshell fragments from the uppermost Cretaceous of the Pyrenees and implications for crocodylomorph eggshell structure","volume":"26","author":"M Moreno-Azanza","year":"2014","journal-title":"Hist Biol"},{"key":"ref36","doi-asserted-by":"crossref","first-page":"275","DOI":"10.1016\/S0016-6995(87)80038-4","article-title":"Pr\u00e9sence d\u2019oeufs de crocodiliens dans le Rognacien inf\u00e9rieur (Maastrichtien sup\u00e9rieur) du bassin d\u2019Aix-en-Provence (Bouches-du-Rh\u00f4ne, France). Note pr\u00e9liminaire","volume":"20","author":"P Kerourio","year":"1987","journal-title":"Geobios"},{"key":"ref37","doi-asserted-by":"crossref","first-page":"329","DOI":"10.1007\/BF02985723","article-title":"Fossile Reptil-Eischalen (Chelonia, Crocodilia, Dinosauria) aus dem unteren Barremium von Galve (Provinz Teruel, SE-Spanien)","volume":"64","author":"R Kohring","year":"1990","journal-title":"Pal\u00e4ontol Z"},{"key":"ref38","first-page":"253","article-title":"Histostructure of fossil crocodilian eggshell fragments from the Lower Cretaceous of Galve (Spain)","volume":"3","author":"R Kohring","year":"1992","journal-title":"Berl Geowiss Abh"},{"key":"ref39","article-title":"Restos indirectos de dinosaurios del registro espa\u00f1ol Paleoicnolog\u00eda de la Cuenca de Cameros (Jur\u00e1sico Superior-Cret\u00e1cico Inferior) y Paleoolog\u00eda de Cret\u00e1cico Superior","author":"JJM Garc\u00eda","year":"1993"},{"key":"ref40","doi-asserted-by":"crossref","first-page":"113","DOI":"10.1016\/S0016-6995(00)80155-2","article-title":"Diversit\u00e9 des coquilles \u201cMinces\u201d d\u2019oeufs fossiles du Cr\u00e9tac\u00e9 sup\u00e9rieur du Sud de la France","volume":"33","author":"G Garcia","year":"2000","journal-title":"Geobios"},{"key":"ref41","doi-asserted-by":"crossref","first-page":"687","DOI":"10.1016\/j.cretres.2008.02.004","article-title":"The vertebrate assemblage of Buenache de la Sierra (Upper Barremian of Serrania de Cuenca, Spain) with insights into its taphonomy and palaeoecology","volume":"29","author":"AD Buscalioni","year":"2008","journal-title":"Cretac Res"},{"key":"ref42","unstructured":"Grellet-Tinner G, Pouech J, Mazin J-M. Exquisitely preserved reptile eggshell fragments from the Berriasian site of Cherves-de-Cognac (Charente): paleobiological implications. Doc Lab G\u00e9ologie Lyon. 2008;164."},{"key":"ref43","doi-asserted-by":"crossref","first-page":"205","DOI":"10.5209\/rev_JIGE.2010.v36.n2.8","article-title":"La Cantalera: an exceptional window onto the vertebrate biodiversity of the Hauterivian-Barremian transition in the Iberian Peninsula\/La Cantalera: una excepcional ventana a la biodiversidad del tr\u00e1nsito Hauteriviense-Barremiense en la Pen\u00ednsula Ib\u00e9rica","volume":"36","author":"JI Canudo","year":"2010","journal-title":"J Iber Geol"},{"key":"ref44","doi-asserted-by":"crossref","first-page":"491","DOI":"10.1080\/02724634.1990.10011832","article-title":"Eggs and eggshell fragments from the Upper Cretaceous Two Medicine Formation of Montana","volume":"10","author":"KF Hirsch","year":"1990","journal-title":"J Vertebr Paleontol"},{"key":"ref45","doi-asserted-by":"crossref","first-page":"1142","DOI":"10.1080\/02724634.2010.483537","article-title":"Fossil eggs and eggshell from the lowermost Two Medicine Formation of western Montana, Sevenmile Hill locality","volume":"30","author":"FD Jackson","year":"2010","journal-title":"J Vertebr Paleontol"},{"key":"ref46","doi-asserted-by":"crossref","first-page":"41","DOI":"10.1080\/08912963.2010.499171","article-title":"Fossil eggshells from the Upper Cretaceous (Campanian) Fruitland Formation, New Mexico","volume":"23","author":"K Tanaka","year":"2011","journal-title":"Hist Biol"},{"key":"ref47","doi-asserted-by":"crossref","first-page":"780","DOI":"10.1671\/0272-4634(2000)020[0780:ACCEFT]2.0.CO;2","article-title":"A complete crocodiloid egg from the Lower Cretaceous (Albian) Glen Rose Formation, central Texas","volume":"20","author":"JV Rogers","year":"2001","journal-title":"J Vertebr Paleontol"},{"key":"ref48","unstructured":"Erickson BR. An amniote egg from the Upper Cretaceous of Wyoming. 1978."},{"key":"ref49","doi-asserted-by":"crossref","first-page":"59","DOI":"10.1080\/02724634.1992.10011431","article-title":"Crocodilian eggs from the Middle Eocene Bridger Formation, Wyoming","volume":"12","author":"KF Hirsch","year":"1992","journal-title":"J Vertebr Paleontol"},{"key":"ref50","first-page":"137","article-title":"Upper Jurassic eggshells from the western interior of North America","author":"KF Hirsch","year":"1994","journal-title":"Carpent K Hirsch KF Horner JR Ed Dinosaur Eggs Babies"},{"key":"ref51","unstructured":"Ribeiro CMM, Carvalho IDS, Nava WR. Ovos de crocodilomorfos da Forma\u00e7\u00e3o Ara\u00e7atuba (Bacia Bauru, Cret\u00e1ceo Superior), Brasil. Gallo V Brito PM Silva HMA Figueiredo FJ Ed Paleontol Vertebr Gd Temas E Contrib Cient\u00edficas Rio Jan Ed Interci\u00eancia. 2006; 285\u2013292."},{"key":"ref52","unstructured":"Ribeiro CMM, Carvalho IDS, Freitas IF, Andrade JAFG. Ovos de crocodilomorfos da Forma\u00e7\u00e3o Ara\u00e7atuba (Bacia Bauru, Cret\u00e1ceo Superior), Brasil. Gallo V Brito PM Silva HMA Figueiredo FJ Ed Paleontol Vertebr Gd Temas E Contrib Cient\u00edficas Rio Jan Ed Interci\u00eancia. 2011; 663\u2013668."},{"key":"ref53","doi-asserted-by":"crossref","first-page":"1316","DOI":"10.1671\/039.029.0409","article-title":"Bizarre notosuchian crocodyliform with associated eggs from the Upper Cretaceous of Bolivia","volume":"29","author":"FE Novas","year":"2009","journal-title":"J Vertebr Paleontol"},{"key":"ref54","first-page":"91","article-title":"Fossil crocodile from the Upper Siwaliks of India","volume":"33","author":"R Patnaik","year":"1993","journal-title":"Mitteilungen Bayer Staatssam Palaeont Hist Geol"},{"key":"ref55","article-title":"A complete crocodylian egg from the Upper Miocene (Chinji Beds) of Pakistan and its palaeobiographical implications","volume":"6","author":"I Panad\u00e8s","year":"2009","journal-title":"PalArchs J Vertebr Palaeontol"},{"key":"ref56","first-page":"307","article-title":"Reptilia from the intertrappean beds of Bombay (India)","volume":"4","author":"SD Singh","year":"1998","journal-title":"Ver\u00f6ff Aus Dem Fuhlrott-Mus"},{"key":"ref57","unstructured":"Russo J, Mateus O, Balbino A, Marzola M. Crocodylomorph eggs and eggshells from the Lourinh\u00e3 Fm.(Upper Jurassic), Portugal. 2014; Available: <ext-link xmlns:xlink=\"http:\/\/www.w3.org\/1999\/xlink\" ext-link-type=\"uri\" xlink:href=\"http:\/\/www.rdpc.uevora.pt\/handle\/10174\/14251\" xlink:type=\"simple\">http:\/\/www.rdpc.uevora.pt\/handle\/10174\/14251<\/ext-link>"},{"key":"ref58","first-page":"83","article-title":"Upper Jurassic dinosaur and crocodile eggs from Paimogo nesting site (Lourinha-Portugal)","volume":"37","author":"MT Antunes","year":"1998","journal-title":"Mem Acad Ci\u00eanc Lisb"},{"key":"ref59","first-page":"71","article-title":"Couv\u00e9e, oeufs et embryons d\u2019un dinosaure th\u00e9ropode du Jurassique sup\u00e9rieur de Lourinh\u00e3 (Portugal)","volume":"325","author":"I Mateus","year":"1997","journal-title":"Comptes Rendus Acad\u00e9mie Sci-Ser IIA-Earth Planet Sci"},{"key":"ref60","article-title":"Upper Jurassic theropod dinosaur embryos from Lourinh\u00e3 (Portugal)","volume":"37","author":"I Mateus","year":"1998","journal-title":"Mem Acad Ci\u00eanc Lisb"},{"key":"ref61","first-page":"647","article-title":"Histomorphogenesis of embryos of Upper Jurassic theropods from Lourinh\u00e3 (Portugal)","volume":"332","author":"A de Ricql\u00e8s","year":"2001","journal-title":"Comptes Rendus Acad\u00e9mie Sci-Ser IIA-Earth Planet Sci"},{"issue":"3","key":"ref62","article-title":"Dinosaur eggshell and embryo localities in Lourinha Formation, Late Jurassic, Portugal","volume":"29","author":"R Castanhinha","year":"2009","journal-title":"J Vertebr Paleontol"},{"key":"ref63","doi-asserted-by":"crossref","unstructured":"Martins RM, Beckmann F, Castanhinha R, Mateus O, Pranzas PK. Dinosaur and crocodile fossils from the Mesozoic of Portugal: neutron tomography and synchrotron-radiation based micro-computed tomography. MRS Proceedings. Cambridge Univ Press; 2011. pp. mrsf10\u20131319. Available: <ext-link xmlns:xlink=\"http:\/\/www.w3.org\/1999\/xlink\" ext-link-type=\"uri\" xlink:href=\"http:\/\/journals.cambridge.org\/abstract_S1946427411007949\" xlink:type=\"simple\">http:\/\/journals.cambridge.org\/abstract_S1946427411007949<\/ext-link>","DOI":"10.1557\/opl.2011.794"},{"key":"ref64","doi-asserted-by":"crossref","first-page":"206","DOI":"10.1080\/08912963.2013.807254","article-title":"Two new theropod egg sites from the Late Jurassic Lourinh\u00e3 Formation, Portugal","volume":"26","author":"V Ribeiro","year":"2014","journal-title":"Hist Biol"},{"key":"ref65","first-page":"223","article-title":"Late Jurassic dinosaurs from the Morrison Formation (USA), the Lourinha and Alcoba\u00e7a formations (Portugal), and the Tendaguru Beds (Tanzania): a comparison","volume":"36","author":"O Mateus","year":"2006","journal-title":"N M Mus Nat Hist Sci Bull"},{"key":"ref66","article-title":"The sedimentology and lithostratigraphy of the Upper Jurassic Lourinha formation, Lusitanian Basin, Portugal","author":"G Hill","year":"1988"},{"key":"ref67","article-title":"Upper Jurassic to Lowermost Cretaceous of the Lusitanian Basin, Portugal-landscapes where dinosaurs walked","author":"O Mateus","year":"2014","journal-title":"Ci\u00eanc Terra Spec"},{"key":"ref68","doi-asserted-by":"crossref","unstructured":"Salminen J, Dinis J, Mateus O. Preliminary Magnetostratigraphy for the Jurassic\u2013Cretaceous Transition in Porto da Calada, Portugal. In: Rocha R, Pais J, Kullberg JC, Finney S, editors. STRATI 2013: First International Congress on Stratigraphy At the Cutting Edge of Stratigraphy. Cham: Springer International Publishing; 2014. pp. 873\u2013877.","DOI":"10.1007\/978-3-319-04364-7_165"},{"key":"ref69","author":"JCR Kullberg","year":"2000"},{"key":"ref70","unstructured":"Kullberg JC, Rocha RB, Soares AF, Rey J, Terrinha P, Callapez P, et al. A Bacia Lusitaniana: estratigrafia, paleogeografia e tect\u00f3nica. 2006; Available: <ext-link xmlns:xlink=\"http:\/\/www.w3.org\/1999\/xlink\" ext-link-type=\"uri\" xlink:href=\"http:\/\/run.unl.pt\/handle\/10362\/1487\" xlink:type=\"simple\">http:\/\/run.unl.pt\/handle\/10362\/1487<\/ext-link>"},{"key":"ref71","doi-asserted-by":"crossref","first-page":"1949","DOI":"10.1007\/s00531-008-0359-3","article-title":"Sr-isotope stratigraphy of the Upper Jurassic of central Portugal (Lusitanian Basin) based on oyster shells","volume":"98","author":"S Schneider","year":"2009","journal-title":"Int J Earth Sci"},{"key":"ref72","first-page":"123","article-title":"The large theropod fauna of the Lourinh\u00e3 Formation (Portugal) and its similarity to the Morrison Formation, with a description of a new species of <italic>Allosaurus<\/italic>","volume":"36","author":"O Mateus","year":"2006","journal-title":"N M Mus Nat Hist Sci Bull"},{"key":"ref73","doi-asserted-by":"crossref","first-page":"e88905","DOI":"10.1371\/journal.pone.0088905","article-title":"Torvosaurus gurneyi n. sp., the largest terrestrial predator from Europe, and a proposed terminology of the maxilla anatomy in nonavian theropods","volume":"9","author":"C Hendrickx","year":"2014","journal-title":"PloS One"},{"key":"ref74","first-page":"13","article-title":"A new diplodocid, <italic>Dinheirosaurus lourinhanensis<\/italic> gen. et sp. nov., from the Late Jurassic beds of Portugal","volume":"5","author":"JF Bonaparte","year":"1999","journal-title":"Rev Mus Argent Cienc Nat"},{"key":"ref75","doi-asserted-by":"crossref","first-page":"98","DOI":"10.1111\/zoj.12029","article-title":"Osteology of the Late Jurassic Portuguese sauropod dinosaur <italic>Lusotitan atalaiensis<\/italic> (Macronaria) and the evolutionary history of basal titanosauriforms","volume":"168","author":"PD Mannion","year":"2013","journal-title":"Zool J Linn Soc"},{"key":"ref76","doi-asserted-by":"crossref","first-page":"e857","DOI":"10.7717\/peerj.857","article-title":"A specimen-level phylogenetic analysis and taxonomic revision of Diplodocidae (Dinosauria, Sauropoda)","volume":"3","author":"E Tschopp","year":"2015","journal-title":"PeerJ"},{"key":"ref77","first-page":"1","article-title":"Les dinosauriens du Portugal","volume":"2","author":"AF de Lapparent","year":"1957","journal-title":"Memoires Serv G\u00e9ologiques Port"},{"key":"ref78","article-title":"Les dinosauriens du Portugal","author":"AF de Lapparent","year":"1957"},{"key":"ref79","doi-asserted-by":"crossref","first-page":"77","DOI":"10.1016\/S1631-0683(03)00003-4","article-title":"Dinosaurs of Portugal","volume":"2","author":"MT Antunes","year":"2003","journal-title":"Comptes Rendus Palevol"},{"key":"ref80","first-page":"1","article-title":"Biomineralisation und Diagenese prolatocanaliculater Dinosaurier-Eischalen aus oberjurassisch\/unterkretazischen Ablagerungen von Porto Pinheiro (Portugal)","volume":"9","author":"R Kohring","year":"1993","journal-title":"Berl Geowiss Abh"},{"key":"ref81","first-page":"57","article-title":"Late Jurassic theropod embryos from Porto das Barcas, Lourinh\u00e3 Formation, Portugal","author":"R Ara\u00fajo","year":"2012","journal-title":"Journal of Vertebrate Paleontology"},{"key":"ref82","doi-asserted-by":"crossref","DOI":"10.1038\/srep01924","article-title":"Filling the gaps of dinosaur eggshell phylogeny: Late Jurassic Theropod clutch with embryos from Portugal","volume":"3","author":"R Ara\u00fajo","year":"2013","journal-title":"Sci Rep"},{"key":"ref83","doi-asserted-by":"crossref","first-page":"128","DOI":"10.1080\/02724634.1990.10011796","article-title":"Upper jurassic chelonian eggshell fragments from the Guimarota mine (central Portugal)","volume":"10","author":"R Kohring","year":"1990","journal-title":"J Vertebr Paleontol"},{"key":"ref84","first-page":"87","article-title":"Eggshells from the Guimarota mine","author":"R Kohring","year":"2000","journal-title":"Guimarota Jurass Ecosyst M\u00fcnch Pfeil"},{"key":"ref85","doi-asserted-by":"crossref","first-page":"99","DOI":"10.1111\/j.1096-3642.1982.tb00064.x","article-title":"The structure and composition of the eggshell and embryonic membranes of Alligator mississippiensis","volume":"36","author":"MW Ferguson","year":"1982","journal-title":"Trans Zool Soc Lond"},{"key":"ref86","author":"TA Bryan","year":"2005"},{"key":"ref87","first-page":"531","article-title":"Fossil crocodilian eggs from the Eocene of Colorado","author":"KF Hirsch","year":"1985","journal-title":"J Paleontol"},{"key":"ref88","doi-asserted-by":"crossref","first-page":"67","DOI":"10.1080\/02724634.1996.10011285","article-title":"Crocodilian and avian eggshells from the Middle Eocene of the Geiseltal, Eastern Germany","volume":"16","author":"R Kohring","year":"1996","journal-title":"J Vertebr Paleontol"},{"key":"ref89","doi-asserted-by":"crossref","first-page":"431","DOI":"10.1590\/S1519-69842001000300012","article-title":"Allometry of reproduction in broad-snouted caiman (Caiman latirostris)","volume":"61","author":"LM Verdade","year":"2001","journal-title":"Braz J Biol"},{"key":"ref90","doi-asserted-by":"crossref","first-page":"268","DOI":"10.1643\/CH-11-136","article-title":"Can reproductive allometry assess population marginality in crocodilians? A comparative analysis of Gulf Coast American Alligator (Alligator mississippiensis) populations","volume":"2013","author":"CM Murray","year":"2013","journal-title":"Copeia"},{"key":"ref91","doi-asserted-by":"crossref","first-page":"301","DOI":"10.1670\/145-03A","article-title":"Allometry of reproduction in wild broad-snouted caimans (Caiman latirostris)","volume":"38","author":"A Larriera","year":"2004","journal-title":"J Herpetol"},{"key":"ref92","doi-asserted-by":"crossref","first-page":"185","DOI":"10.1111\/1475-4983.00233","article-title":"A new species of Goniopholis from the Upper Jurassic of Portugal","volume":"45","author":"D Schwarz","year":"2002","journal-title":"Palaeontology"},{"key":"ref93","doi-asserted-by":"crossref","first-page":"1259","DOI":"10.1139\/e04-059","article-title":"Lusitanisuchus, a new generic name for Lisboasaurus mitracostatus (Crocodylomorpha: Mesoeucrocodylia), with a description of new remains from the Upper Jurassic (Kimmeridgian) and Lower Cretaceous (Berriasian) of Portugal","volume":"41","author":"D Schwarz","year":"2004","journal-title":"Can J Earth Sci"},{"key":"ref94","doi-asserted-by":"crossref","first-page":"264","DOI":"10.1671\/0272-4634(2008)28[264:TFDOLC]2.0.CO;2","article-title":"The first dentary of lisboasaurus (crocodylomorpha,? mesoeucrocodylia) from the Lower Cretaceous (Barremian) of Una, Cuenca Province, Spain","volume":"28","author":"D Schwarz","year":"2008","journal-title":"J Vertebr Paleontol"},{"key":"ref95","doi-asserted-by":"crossref","first-page":"140222","DOI":"10.1098\/rsos.140222","article-title":"Revision of the Late Jurassic teleosaurid genus Machimosaurus (Crocodylomorpha, Thalattosuchia)","volume":"1","author":"MT Young","year":"2014","journal-title":"R Soc Open Sci"},{"key":"ref96","doi-asserted-by":"crossref","first-page":"e599","DOI":"10.7717\/peerj.599","article-title":"Revision of the Late Jurassic crocodyliform Alligatorellus, and evidence for allopatric speciation driving high diversity in western European atoposaurids","volume":"2","author":"JP Tennant","year":"2014","journal-title":"PeerJ"},{"key":"ref97","unstructured":"Krebs B. Le crocodilien Machimosaurus. 1968."},{"key":"ref98","unstructured":"Seiffert J. Upper Jurassic lizards from central Portugal. 1973."},{"key":"ref99","first-page":"572","article-title":"Mittheilungen an Professor Bronn gerichtet","volume":"1837","author":"H von Meyer","year":"1837","journal-title":"Neues Jahrb F\u00fcr Mineral Geogn Geol Petrefaktenkunde"},{"key":"ref100","unstructured":"Mateus O. Checklist for Late Jurassic reptiles and amphibians from Portugal. Livro de Resumos do X Congresso Luso-Espanhol de Herpetologia. 2008. Available: <ext-link xmlns:xlink=\"http:\/\/www.w3.org\/1999\/xlink\" ext-link-type=\"uri\" xlink:href=\"http:\/\/docentes.fct.unl.pt\/omateus\/publications\/checklist-late-jurassic-reptiles-and-amphibians-portugal?page=1\" xlink:type=\"simple\">http:\/\/docentes.fct.unl.pt\/omateus\/publications\/checklist-late-jurassic-reptiles-and-amphibians-portugal?page=1<\/ext-link>"},{"key":"ref101","first-page":"161","article-title":"Chronology of the Late Jurassic dinosaur faunas, and other reptilian faunas, from Portugal","author":"V Ribeiro","year":"2012","journal-title":"Journal of Vertebrate Paleontology, Program and Abstracts"},{"key":"ref102","doi-asserted-by":"crossref","first-page":"779","DOI":"10.1016\/j.geobios.2004.04.005","article-title":"A new species of Theriosuchus (Atoposauridae, Crocodylomorpha) from the Late Jurassic (Kimmeridgian) of Guimarota, Portugal","volume":"38","author":"D Schwarz","year":"2005","journal-title":"Geobios"},{"key":"ref103","unstructured":"Mateus O. Crocodylomorphs from the Mesozoic of Portugal and a new skull of eusuchian from the Late Cretaceous. Abstract Book of Hwaseong International Dinosaurs Expedition Symposium, South Korea. 2013. pp. 66\u201368."},{"key":"ref104","first-page":"59","article-title":"The Late Jurassic crocodiles of the Langenberg near Oker, Lower Saxony (Germany), and description of related materials (with remarks on the history of quarrying the \u201cLangenberg Limestone\u201d and \u201cObernkirchen Sandstone\u201d)","volume":"5","author":"H-V Karl","year":"2006","journal-title":"Clausthal Geowiss"},{"key":"ref105","doi-asserted-by":"crossref","first-page":"46","DOI":"10.1007\/BF02998548","article-title":"Der Jura-Krokodilier Machimosaurus H. v. Meyer","volume":"41","author":"B Krebs","year":"1967","journal-title":"Pal\u00e4ontol Z"},{"key":"ref106","doi-asserted-by":"crossref","first-page":"801","DOI":"10.1111\/j.1096-3642.2009.00571.x","article-title":"The evolution of Metriorhynchoidea (Mesoeucrocodylia, Thalattosuchia): an integrated approach using geometric morphometrics, analysis of disparity, and biomechanics","volume":"158","author":"MT Young","year":"2010","journal-title":"Zool J Linn Soc"},{"key":"ref107","unstructured":"Moreno-Azanza M, Canudo JI, Gasca JM. Was crocodylomorph eggshell structure always as conservative as it is in extant crocodilians? XIII Annual Meeting of the European Association of Vertebrate Palaeontologists, Opole, Poland. 2015. p. 97. Available: <ext-link xmlns:xlink=\"http:\/\/www.w3.org\/1999\/xlink\" ext-link-type=\"uri\" xlink:href=\"http:\/\/www.sciencedirect.com\/science\/article\/pii\/S0195667115300239\" xlink:type=\"simple\">http:\/\/www.sciencedirect.com\/science\/article\/pii\/S0195667115300239<\/ext-link>"},{"key":"ref108","article-title":"First definitive association between embryonic Allosaurus bones and Prismatoolithus eggs in the Morrison Formation (Upper Jurassic, Wyoming, USA)","volume":"101","author":"M Carrano","year":"2013","journal-title":"J Vertebr Paleontol Program Abstr"},{"key":"ref109","doi-asserted-by":"crossref","first-page":"23","DOI":"10.1655\/HERPMONOGRAPHS-D-11-00001","article-title":"The role of predation in shaping crocodilian natural history","volume":"27","author":"R Somaweera","year":"2013","journal-title":"Herpetol Monogr"}],"container-title":["PLOS ONE"],"original-title":[],"language":"en","link":[{"URL":"http:\/\/dx.plos.org\/10.1371\/journal.pone.0171919","content-type":"unspecified","content-version":"vor","intended-application":"similarity-checking"}],"deposited":{"date-parts":[[2022,7,25]],"date-time":"2022-07-25T17:44:40Z","timestamp":1658771080000},"score":1,"resource":{"primary":{"URL":"https:\/\/dx.plos.org\/10.1371\/journal.pone.0171919"}},"subtitle":[],"editor":[{"given":"Matthew","family":"Shawkey","sequence":"first","affiliation":[]}],"short-title":[],"issued":{"date-parts":[[2017,3,8]]},"references-count":109,"journal-issue":{"issue":"3","published-online":{"date-parts":[[2017,3,8]]}},"URL":"https:\/\/doi.org\/10.1371\/journal.pone.0171919","relation":{},"ISSN":["1932-6203"],"issn-type":[{"value":"1932-6203","type":"electronic"}],"subject":[],"published":{"date-parts":[[2017,3,8]]}}}