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However, in fermentation processes that depend on bacterial activity, bacteriophages can pose a major threat by infecting and disrupting key microbial populations, leading to delayed or failed fermentations and resulting in economic losses. While the impact of bacteriophages on fermented dairy foods, such as yogurt and cheese, is well-documented, such knowledge is comparatively limited for fermented soybean foods (FSFs). Most FSFs (such as Cheonggukjang, Miso, Thua Nao, and soy sauce) are traditionally produced using natural microbiota under less controlled conditions. This makes them more susceptible to unpredictable microbial dynamics, including phage interference. In this review, we examine the diversity and occurrence of bacteriophages associated with FSFs. Finally, the potential roles of bacteriophages in FSFs\u00a0\u2013\u00a0ranging from fermentation disruption to possible microbial modulation\u00a0\u2013\u00a0are also discussed.<\/jats:p>","DOI":"10.1515\/bmc-2025-0061","type":"journal-article","created":{"date-parts":[[2026,7,8]],"date-time":"2026-07-08T15:56:09Z","timestamp":1783526169000},"source":"Crossref","is-referenced-by-count":0,"title":["Research highlights in bacteriophages of fermented soybean foods"],"prefix":"10.1515","volume":"17","author":[{"given":"Ekachai","family":"Chukeatirote","sequence":"first","affiliation":[{"name":"School of Science , Mae Fah Luang University , Chiang Rai 57100 , Thailand"}],"role":[{"vocabulary":"crossref","role":"author"}]},{"ORCID":"https:\/\/orcid.org\/0000-0001-8935-8807","authenticated-orcid":false,"given":"Dong-Qin","family":"Dai","sequence":"additional","affiliation":[{"name":"Center for Yunnan Plateau Biological Resources Protection and Utilization, College of Biological Resource and Food Engineering , Qujing Normal University , Qujing 655011 , China"}],"role":[{"vocabulary":"crossref","role":"author"}]},{"given":"Samantha C.","family":"Karunarathna","sequence":"additional","affiliation":[{"name":"Center for Yunnan Plateau Biological Resources Protection and Utilization, College of Biological Resource and Food Engineering , Qujing Normal University , Qujing 655011 , China"}],"role":[{"vocabulary":"crossref","role":"author"}]},{"given":"Saowaluck","family":"Tibpromma","sequence":"additional","affiliation":[{"name":"Center for Yunnan Plateau Biological Resources Protection and Utilization, College of Biological Resource and Food Engineering , Qujing Normal University , Qujing 655011 , China"}],"role":[{"vocabulary":"crossref","role":"author"}]}],"member":"374","published-online":{"date-parts":[[2026,7,8]]},"reference":[{"key":"2026070815560228532_j_bmc-2025-0061_ref_001","doi-asserted-by":"crossref","unstructured":"Mushegian, AR. 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Sci Rep 2023;13:7573. https:\/\/doi.org\/10.1038\/s41598-023-34818-0.","DOI":"10.1038\/s41598-023-34818-0"},{"key":"2026070815560228532_j_bmc-2025-0061_ref_047","doi-asserted-by":"crossref","unstructured":"Wicaksono, WA, Akinyemi, OE, Wassermann, B, Bickel, S, Suwanto, A, Berg, G. The terroir of Tempeh: strong region-specific signatures in the bacterial community structures across Indonesia. Curr Res Microb Sci 2024;7:100287. https:\/\/doi.org\/10.1016\/j.crmicr.2024.100287.","DOI":"10.1016\/j.crmicr.2024.100287"},{"key":"2026070815560228532_j_bmc-2025-0061_ref_048","unstructured":"d\u2019Herelle, F. Sur un microbe invisible antagoniste des bacilles dysent\u00e9riques. CR Acad Sci Paris 1917;165:373\u20135."},{"key":"2026070815560228532_j_bmc-2025-0061_ref_049","doi-asserted-by":"crossref","unstructured":"Twort, FW. An investigation on the nature of ultramicroscopic viruses. 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Extensive host range determination and improved efficacy of the bacteriophage JBP901 in the presence of divalent cations for control of Bacillus cereus in Cheonggukjang. Food Sci Biotechnol 2014;23:499\u2013504. https:\/\/doi.org\/10.1007\/s10068-014-0068-6.","DOI":"10.1007\/s10068-014-0068-6"},{"key":"2026070815560228532_j_bmc-2025-0061_ref_105","doi-asserted-by":"crossref","unstructured":"Kumar, J, Sharma, N, Kaushal, G, Samurailatpam, S, Sahoo, D, Rai, AK, et al.. Metagenomic insights into the taxonomic and functional features of Kinema, a traditional fermented soybean product of Sikkim Himalaya. Front Microbiol 2019;10:1744. https:\/\/doi.org\/10.3389\/fmicb.2019.01744.","DOI":"10.3389\/fmicb.2019.01744"},{"key":"2026070815560228532_j_bmc-2025-0061_ref_106","doi-asserted-by":"crossref","unstructured":"Tan, G, Qi, S, Wang, Y, Li, X, Li, X, Li, M, et al.. 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Appl Environ Microbiol 2011;77:2264\u201374. https:\/\/doi.org\/10.1128\/AEM.02157-10.","DOI":"10.1128\/AEM.02157-10"},{"key":"2026070815560228532_j_bmc-2025-0061_ref_118","doi-asserted-by":"crossref","unstructured":"Park, WJ, Kong, SJ, Park, JH. Kimchi bacteriophages of lactic acid bacteria: population, characteristics, and their role in watery kimchi. Food Sci Biotechnol 2021;30:949\u201357. https:\/\/doi.org\/10.1007\/s10068-021-00930-y.","DOI":"10.1007\/s10068-021-00930-y"},{"key":"2026070815560228532_j_bmc-2025-0061_ref_119","doi-asserted-by":"crossref","unstructured":"Ledormand, P, Desmasures, N, Bernay, B, Goux, D, Rue, O, Midoux, C, et al.. Molecular approaches to uncover phage\u2013lactic acid bacteria interactions in a model community simulating fermented beverages. 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