{"status":"ok","message-type":"work","message-version":"1.0.0","message":{"indexed":{"date-parts":[[2026,3,4]],"date-time":"2026-03-04T23:29:54Z","timestamp":1772666994092,"version":"3.50.1"},"posted":{"date-parts":[[2026]]},"group-title":"SSRN","reference-count":86,"publisher":"Elsevier BV","content-domain":{"domain":[],"crossmark-restriction":false},"short-container-title":[],"abstract":"<jats:p>Understanding wildlife responses to both anthropogenic pressures and natural environmental gradients requires large-scale and standardized surveys across broad spatial extents, enabling the identification of general patterns to assist tailored management strategies. Here, we aimed to identify the main environmental drivers shaping the occupancy patterns of widespread medium-sized mammals across multiple contrasting regions to support broad-scale inference on species-environment relationships. Using a well-replicated camera-trap dataset from 15 study areas distributed across three macrobioclimatic regions in mainland Portugal, we assessed how species occupancy was influenced by land-use composition and configuration, elevation, human footprint, seasonality, prey availability and hunting regimes, while accounting for spatial dependencies through nested random effects. Hierarchical modelling revealed that species detectability was weakly affected by covariates, whereas occupancy was primarily associated with broad-scale patterns of land-use composition and configuration, with additional effects of elevation for some species. In contrast, human footprint, hunting regimes and seasonality showed little or no detectable influence on occupancy across species at the broad spatial scale considered. Responses to land-use patterns differed among taxa, with lagomorphs showing strong but contrasting occupancy patterns along gradients of agricultural habitat structure, and carnivores exhibiting weaker and more variable responses associated with landscape heterogeneity. Overall, our results revealed consistent species-specific effects of land-use composition and configuration on broad-scale occupancy patterns of medium-sized mammals. These findings demonstrate the value of large-scale monitoring frameworks for moving beyond site-specific inference and informing transferable conservation strategies guided by land-use management across multiple regions and environmental contexts.<\/jats:p>","DOI":"10.2139\/ssrn.6345714","type":"posted-content","created":{"date-parts":[[2026,3,4]],"date-time":"2026-03-04T22:45:46Z","timestamp":1772664346000},"source":"Crossref","is-referenced-by-count":0,"title":["Multi-region occupancy models reveal broad-scale land-use effects on medium-sized mammals"],"prefix":"10.2139","author":[{"ORCID":"https:\/\/orcid.org\/0000-0001-8562-7301","authenticated-orcid":true,"given":"Ricardo","family":"Pita","sequence":"first","affiliation":[]},{"given":"Francisco","family":"Fradinho","sequence":"additional","affiliation":[]},{"ORCID":"https:\/\/orcid.org\/0000-0002-2597-7251","authenticated-orcid":true,"given":"Nuno","family":"Pinto","sequence":"additional","affiliation":[]},{"ORCID":"https:\/\/orcid.org\/0000-0001-8872-3526","authenticated-orcid":true,"given":"Guilherme","family":"Ares Pereira","sequence":"additional","affiliation":[]},{"given":"Carlos","family":"Barroqueiro","sequence":"additional","affiliation":[]},{"given":"Ant\u00f3nio","family":"Mira","sequence":"additional","affiliation":[]},{"given":"Rita","family":"Torres","sequence":"additional","affiliation":[]},{"ORCID":"https:\/\/orcid.org\/0000-0002-9792-9259","authenticated-orcid":true,"given":"Jo\u00e3o","family":"Carvalho","sequence":"additional","affiliation":[]}],"member":"78","reference":[{"key":"ref1","doi-asserted-by":"crossref","DOI":"10.1371\/journal.pone.0051529","article-title":"Past, present and future distributions of an Iberian endemic, Lepus granatensis: ecological and evolutionary clues from species distribution models","volume":"7","author":"P Acevedo","year":"2012","journal-title":"PLoS ONE"},{"key":"ref2","year":"2017","journal-title":"FastPictureViewer Professional: Fast Image Viewer"},{"key":"ref3","doi-asserted-by":"crossref","DOI":"10.1201\/b17115","author":"S Banerjee","year":"2014","journal-title":"Hierarchical modeling and analysis for spatial data"},{"key":"ref4","author":"J M Barea-Azc\ufffdn","year":"2007","journal-title":"Surveying carnivores at large spatial scales: a comparison of four broad-applied methods. Pages 387-404 in Vertebrate conservation and biodiversity"},{"key":"ref5","doi-asserted-by":"crossref","first-page":"636","DOI":"10.1007\/s00267-024-02003-2","article-title":"Do mesocarnivores respond to the seasonality in management practices in an agroforestry landscape?","volume":"74","author":"A L Barros","year":"2024","journal-title":"Environmental Management"},{"key":"ref6","author":"D Bates","year":"2025","journal-title":"2025. lme4: Linear mixed-effects models using 'Eigen' and S4. R package manual"},{"key":"ref7","doi-asserted-by":"crossref","first-page":"488","DOI":"10.1111\/gean.12319","article-title":"R packages for analyzing spatial data: a comparative case study with areal data","volume":"54","author":"R Bivand","year":"2022","journal-title":"Geographical Analysis"},{"key":"ref8","doi-asserted-by":"crossref","first-page":"434","DOI":"10.1080\/10618600.1998.10474787","article-title":"General methods for monitoring convergence of iterative simulations","volume":"7","author":"S P Brooks","year":"1998","journal-title":"Journal of Computational and Graphical Statistics"},{"key":"ref9","author":"K P Burnham","year":"2002","journal-title":"Model selection and multimodel inference: a practical information-theoretic approach"},{"key":"ref10","doi-asserted-by":"crossref","first-page":"675","DOI":"10.1111\/1365-2664.12432","article-title":"Wildlife camera trapping: a review and recommendations for linking surveys to ecological processes","volume":"52","author":"A C Burton","year":"2015","journal-title":"Journal of Applied Ecology"},{"key":"ref11","doi-asserted-by":"crossref","first-page":"399","DOI":"10.1016\/j.mambio.2016.03.003","article-title":"Niche constraints to the northwards expansion of the common genet (Genetta genetta, Linnaeus 1758) in Europe","volume":"81","author":"D Camps","year":"2016","journal-title":"Mammalian Biology"},{"key":"ref12","doi-asserted-by":"crossref","first-page":"328","DOI":"10.1111\/mam.12292","article-title":"Variation in red fox Vulpes vulpes diet in five continents","volume":"52","author":"I Casta\ufffdeda","year":"2022","journal-title":"Mammal Review"},{"key":"ref13","doi-asserted-by":"crossref","first-page":"279","DOI":"10.4098\/AT.arch.94-31","article-title":"Home range, habitat selection and activity of the red fox in a Mediterranean coastal ecotone","volume":"39","author":"P Cavallini","year":"1994","journal-title":"Acta Theriologica"},{"key":"ref14","doi-asserted-by":"crossref","first-page":"24","DOI":"10.17161\/bi.v2i0.5","article-title":"Environmental information: placing biodiversity phenomena in an ecological and environmental context","volume":"2","author":"A D Chapman","year":"2005","journal-title":"Biodiversity Informatics"},{"key":"ref15","doi-asserted-by":"crossref","DOI":"10.3390\/rs14081865","article-title":"Spatially stratified and multi-stage approach for national land cover mapping based on Sentinel-2 data and expert knowledge","volume":"14","author":"H Costa","year":"2022","journal-title":"Remote Sensing"},{"issue":"16","key":"ref16","doi-asserted-by":"crossref","first-page":"280","DOI":"10.1016\/j.agee.2016.12.037","article-title":"Mediterranean mesocarnivores in spatially structured managed landscapes: community organization in time and space","volume":"237","author":"G Curveira-Santos","year":"2017","journal-title":"Agriculture, Ecosystems & Environment"},{"key":"ref17","doi-asserted-by":"crossref","DOI":"10.1371\/journal.pone.0210661","article-title":"Mesocarnivore community structure under predator control: unintended patterns in a conservation context","volume":"14","author":"G Curveira-Santos","year":"2019","journal-title":"PLoS ONE"},{"key":"ref18","doi-asserted-by":"crossref","first-page":"456","DOI":"10.1071\/WR10046","article-title":"A comparison of the effectiveness of camera trapping and live trapping for sampling terrestrial small-mammal communities","volume":"37","author":"N De Bondi","year":"2010","journal-title":"Wildlife Research"},{"key":"ref19","doi-asserted-by":"crossref","first-page":"169","DOI":"10.1017\/S0376892910000214","article-title":"Land-use changes as a critical factor for long-term wild rabbit conservation in the Iberian Peninsula","volume":"37","author":"M Delibes-Mateos","year":"2010","journal-title":"Environmental Conservation"},{"key":"ref20","doi-asserted-by":"crossref","first-page":"59","DOI":"10.1111\/j.1365-2907.2011.00206.x","article-title":"Biogeographical patterns in the diet of an opportunistic predator: the red fox Vulpes vulpes in the Iberian Peninsula","volume":"43","author":"F D\ufffdaz-Ruiz","year":"2013","journal-title":"Mammal Review"},{"key":"ref21","year":"2019","journal-title":"Carta de Uso e Ocupa\ufffd\ufffdo do Solo (COS) de Portugal Continental para 2018. dados.gov.pt dataset"},{"key":"ref22","doi-asserted-by":"crossref","first-page":"27","DOI":"10.1111\/j.1600-0587.2012.07348.x","article-title":"Collinearity: a review of methods to deal with it and a simulation study evaluating their performance","volume":"36","author":"C F Dormann","year":"2013","journal-title":"Ecography"},{"key":"ref23","doi-asserted-by":"crossref","first-page":"1670","DOI":"10.1111\/2041-210X.13897","article-title":"2022. spOccupancy: an R package for singlespecies, multi-species, and integrated spatial occupancy models","volume":"13","author":"J W Doser","journal-title":"Methods in Ecology and Evolution"},{"key":"ref24","doi-asserted-by":"crossref","first-page":"201","DOI":"10.1644\/05-MAMM-A-343R3.1","article-title":"Factors affecting the placement of common genet latrine sites in a Mediterranean landscape in Portugal","volume":"88","author":"C Esp\ufffdrito-Santo","year":"2007","journal-title":"Journal of Mammalogy"},{"key":"ref25","year":"2016","journal-title":"EU-DEM v1.1 digital surface model (DSM)"},{"key":"ref26","doi-asserted-by":"crossref","first-page":"258","DOI":"10.1111\/j.1469-7998.2011.00873.x","article-title":"Effects of human-induced land-use changes on the distribution of the Iberian hare","volume":"286","author":"M A Farf\ufffdn","year":"2012","journal-title":"Journal of Zoology"},{"key":"ref27","doi-asserted-by":"crossref","first-page":"591","DOI":"10.1007\/s10144-015-0510-5","article-title":"Can widespread generalist predators affect keystone prey? A case study with red foxes and European rabbits in their native range","volume":"57","author":"J Fernandez-De-Simon","year":"2015","journal-title":"Popul Ecol"},{"key":"ref28","doi-asserted-by":"crossref","first-page":"487","DOI":"10.1007\/s10344-009-0257-4","article-title":"Influence of habitat management on the abundance and diet of wild rabbit (Oryctolagus cuniculus algirus) populations in Mediterranean ecosystems","volume":"55","author":"C Ferreira","year":"2009","journal-title":"European Journal of Wildlife Research"},{"key":"ref29","doi-asserted-by":"crossref","first-page":"671","DOI":"10.1007\/s11284-008-0538-5","article-title":"The influence of human, livestock, and ecological features on the occurrence of genet (Genetta genetta): a case study on Mediterranean farmland","volume":"24","author":"A Galantinho","year":"2009","journal-title":"Ecological Research"},{"key":"ref30","doi-asserted-by":"crossref","DOI":"10.1002\/ecy.3654","article-title":"Mammals in Portugal: a data set of terrestrial, volant, and marine mammal occurrences in Portugal","volume":"103","author":"C Grilo","year":"2022","journal-title":"Ecology"},{"key":"ref31","doi-asserted-by":"crossref","first-page":"1648","DOI":"10.1111\/ecog.04617","article-title":"landscapemetrics: an open-source R tool to calculate landscape metrics","volume":"42","author":"M H K Hesselbarth","year":"2019","journal-title":"Ecography"},{"key":"ref32","doi-asserted-by":"crossref","first-page":"2951","DOI":"10.1007\/s10531-018-1580-4","article-title":"Drivers of sett site location by European badgers in Portugal","volume":"27","author":"D Hip\ufffdlito","year":"2018","journal-title":"Biodiversity and Conservation"},{"key":"ref33","doi-asserted-by":"crossref","first-page":"127","DOI":"10.1080\/17474230601079241","article-title":"Harmonization of land use class sets to facilitate compatibility and comparability of data across space and time","volume":"1","author":"L J M Jansen","year":"2006","journal-title":"Journal of Land Use Science"},{"key":"ref34","author":"M K\ufffdry","year":"2016","journal-title":"Applied Hierarchical Modeling in Ecology: analysis of distribution, abundance and species richness in R and BUGS: Volume 1: Prelude and Static Models"},{"key":"ref35","doi-asserted-by":"crossref","first-page":"3657","DOI":"10.1007\/s10980-023-01749-0","article-title":"Increasing homogeneity of Mediterranean landscapes limits the co-occurrence of mesocarnivores in space and time","volume":"38","author":"P Linck","year":"2023","journal-title":"Landscape Ecology"},{"key":"ref36","doi-asserted-by":"crossref","first-page":"137","DOI":"10.1007\/s40823-022-00079-2","article-title":"Why we need to invest in large-scale, long-term monitoring programs in landscape ecology and conservation biology","volume":"7","author":"D B Lindenmayer","year":"2022","journal-title":"Current Landscape Ecology Reports"},{"key":"ref37","doi-asserted-by":"crossref","first-page":"26","DOI":"10.1644\/1545-1542(2003)084<0026:HRDIRO>2.0.CO;2","article-title":"Habitat-related differences in rabbit (Oryctolagus cuniculus) abundance, distribution, and activity","volume":"84","author":"L Lombardi","year":"2003","journal-title":"Journal of Mammalogy"},{"key":"ref38","author":"D I Mackenzie","year":"2017","journal-title":"Occupancy estimation and modeling: inferring patterns and dynamics of species occurrence"},{"key":"ref39","author":"M L Mathias","year":"2023","journal-title":"Livro vermelho dos mam\ufffdferos de Portugal continental. FCi\ufffdncias.ID and ICNF"},{"key":"ref40","article-title":"Conditional logit analysis of qualitative choice behavior","volume":"104","author":"D Mcfadden","year":"1974","journal-title":"Frontiers in econometrics"},{"key":"ref41","doi-asserted-by":"crossref","first-page":"4735","DOI":"10.1002\/ece3.1679","article-title":"Estimating occupancy dynamics for large-scale monitoring networks: amphibian breeding occupancy across protected areas in the northeast United States","volume":"5","author":"D A Miller","year":"2015","journal-title":"Ecology and Evolution"},{"key":"ref42","doi-asserted-by":"crossref","first-page":"400","DOI":"10.1002\/joc.4357","article-title":"Bioclimatological mapping tackling uncertainty propagation: application to mainland Portugal","volume":"36","author":"T Monteiro-Henriques","year":"2016","journal-title":"International Journal of Climatology"},{"key":"ref43","doi-asserted-by":"crossref","first-page":"691","DOI":"10.1093\/biomet\/78.3.691","article-title":"A note on a general definition of the coefficient of determination","volume":"78","author":"N J D Nagelkerke","year":"1991","journal-title":"Biometrika"},{"key":"ref44","doi-asserted-by":"crossref","first-page":"45","DOI":"10.1038\/nature14324","article-title":"Global effects of land use on local terrestrial biodiversity","volume":"520","author":"T Newbold","year":"2015","journal-title":"Nature"},{"key":"ref45","doi-asserted-by":"crossref","first-page":"1","DOI":"10.1111\/geb.12236","article-title":"The ecological effects of providing resource subsidies to predators","volume":"24","author":"T M Newsome","year":"2015","journal-title":"Global Ecology and Biogeography"},{"key":"ref46","doi-asserted-by":"crossref","first-page":"27","DOI":"10.1034\/j.1600-0706.2001.11311.x","article-title":"Logic of experiments in ecology: is pseudoreplication a pseudoissue?","volume":"94","author":"L Oksanen","year":"2003","journal-title":"Oikos"},{"key":"ref47","doi-asserted-by":"crossref","first-page":"224","DOI":"10.1078\/1616-5047-00088","article-title":"The negative impact of heavy rains on the abundance of a Mediterranean population of European rabbits","volume":"68","author":"F Palomares","year":"2003","journal-title":"Mammalian Biology"},{"key":"ref48","doi-asserted-by":"crossref","first-page":"752","DOI":"10.2307\/2404253","article-title":"Key habitats for Egyptian mongooses in Do\ufffdana National Park, south-western Spain","volume":"30","author":"F Palomares","year":"1993","journal-title":"Journal of Applied Ecology"},{"key":"ref49","article-title":"Atlas y Libro Rojo de los mam\ufffdferos terrestres de Espa\ufffda","author":"L J Palomo","year":"2007","journal-title":"Direcci\ufffdn General para la Biodiversidad, SECEM-SECEMU"},{"key":"ref50","doi-asserted-by":"crossref","first-page":"439","DOI":"10.32614\/RJ-2018-009","article-title":"Simple features for R: standardized support for spatial vector data","volume":"10","author":"E Pebesma","year":"2018","journal-title":"The R Journal"},{"key":"ref51","doi-asserted-by":"crossref","first-page":"611","DOI":"10.1111\/j.1365-2664.2010.01804.x","article-title":"Conservation value of linear woody remnants for two forest carnivores in a Mediterranean agricultural landscape","volume":"47","author":"M Pereira","year":"2010","journal-title":"Journal of Applied Ecology"},{"key":"ref52","doi-asserted-by":"crossref","first-page":"135","DOI":"10.1007\/978-1-4020-2911-0_10","article-title":"Multifunctionality in Mediterranean landscapes-past and future","author":"T Pinto-Correia","year":"2004","journal-title":"The new dimensions of the European landscape"},{"key":"ref53","first-page":"57","volume":"132","author":"R Pita","year":"2009","journal-title":"Influence of landscape characteristics on carnivore diversity and abundance in Mediterranean farmland. Agriculture"},{"key":"ref54","doi-asserted-by":"crossref","DOI":"10.1016\/j.agee.2019.106721","article-title":"Roads, forestry plantations and hedgerows affect badger occupancy in intensive Mediterranean farmland","volume":"289","author":"R Pita","year":"2020","journal-title":"Agriculture, Ecosystems & Environment"},{"key":"ref55","doi-asserted-by":"crossref","first-page":"1339","DOI":"10.1080\/01621459.2013.829001","article-title":"Bayesian inference for logistic models using Polya-Gamma latent variables","volume":"108","author":"N G Polson","year":"2013","journal-title":"Journal of the American Statistical Association"},{"key":"ref56","doi-asserted-by":"crossref","DOI":"10.3390\/ani12111351","article-title":"The dynamics of Lepus granatensis and Oryctolagus cuniculus in a Mediterranean agrarian area: are hares segregating from rabbit habitats after disease impact?","volume":"12","author":"J Prenda","year":"2022","journal-title":"Animals"},{"key":"ref57","article-title":"Different environmental contexts, different responses: evaluating the drivers of red fox occupancy patterns in Portugal","author":"L","year":"2022","journal-title":"QGIS Geographic Information System (version 3.26.3 \"Buenos Aires\"). Open-Source Geospatial Foundation Project"},{"key":"ref58","article-title":"R: A language and environment for statistical computing","year":"2025","journal-title":"R Foundation for Statistical Computing"},{"key":"ref59","doi-asserted-by":"crossref","first-page":"269","DOI":"10.1016\/S0006-3207(00)00043-4","article-title":"Defining key habitats for low-density populations of Eurasian badgers in Mediterranean environments","volume":"95","author":"E Revilla","year":"2000","journal-title":"Biological Conservation"},{"key":"ref60","doi-asserted-by":"crossref","first-page":"918","DOI":"10.1111\/geb.12600","article-title":"Assessing global patterns in mammalian carnivore occupancy and richness by integrating local camera trap surveys","volume":"26","author":"L N Rich","year":"2017","journal-title":"Global Ecology and Biogeography"},{"key":"ref61","doi-asserted-by":"crossref","first-page":"279","DOI":"10.1016\/j.ecolmodel.2005.03.002","article-title":"The role of environmental generalist species in ecosystem function","volume":"188","author":"C E Richmond","year":"2005","journal-title":"Ecological Modelling"},{"key":"ref62","article-title":"Biogeographic units of the Iberian Peninsula and Balearic Islands to district level: a concise synopsis","volume":"1","author":"S Rivas-Mart\ufffdnez","year":"2017","journal-title":"Pages"},{"key":"ref63","doi-asserted-by":"crossref","first-page":"67","DOI":"10.1111\/j.1365-2907.2008.00134.x","article-title":"Fruit consumption by carnivores in Mediterranean Europe","volume":"39","author":"L M Rosalino","year":"2009","journal-title":"Mammal Review"},{"key":"ref64","doi-asserted-by":"crossref","DOI":"10.1038\/s41598-019-39713-1","article-title":"Climate and landscape changes as driving forces for future range shift in southern populations of the European badger","volume":"9","author":"L M Rosalino","year":"2019","journal-title":"Scientific Reports"},{"key":"ref65","doi-asserted-by":"crossref","first-page":"1493","DOI":"10.1139\/z04-130","article-title":"Spatial structure and land-cover use in a low-density Mediterranean population of Eurasian badgers","volume":"82","author":"L M Rosalino","year":"2004","journal-title":"Canadian Journal of Zoology"},{"key":"ref66","author":"J A Royle","year":"2008","journal-title":"Hierarchical modeling and inference in ecology: the analysis of data from populations, metapopulations and communities"},{"key":"ref67","doi-asserted-by":"crossref","first-page":"1698","DOI":"10.1016\/j.jaridenv.2008.03.006","article-title":"Habitat use by large and small herbivores in a fluctuating Mediterranean ecosystem: implications of seasonal changes","volume":"72","author":"M Rueda","year":"2008","journal-title":"Journal of Arid Environments"},{"key":"ref68","doi-asserted-by":"crossref","DOI":"10.1641\/0006-3568(2002)052[0891:THFATL]2.0.CO;2","article-title":"The human footprint and the last of the wild: The human footprint is a global map of human influence on the land surface, which suggests that human beings are stewards of nature, whether we like it or not","volume":"52","author":"E W Sanderson","year":"2002","journal-title":"BioScience"},{"key":"ref69","doi-asserted-by":"crossref","first-page":"275","DOI":"10.1007\/s10344-009-0317-9","article-title":"Stone marten (Martes foina) habitat in a Mediterranean ecosystem: effects of scale, sex, and interspecific interactions","volume":"56","author":"M J Santos","year":"2010","journal-title":"European Journal of Wildlife Research"},{"key":"ref70","doi-asserted-by":"crossref","first-page":"147","DOI":"10.1007\/0-387-22691-5_7","article-title":"Relationships between stone martens, genets and cork oak woodlands in Portugal","author":"M Santos-Reis","year":"2005","journal-title":"Martens and fishers (Martes) in human-altered environments: an international perspective"},{"key":"ref71","doi-asserted-by":"crossref","first-page":"174","DOI":"10.1016\/j.envsci.2005.11.006","article-title":"The limitations of spatial land use data in environmental analysis","volume":"9","author":"C Schmit","year":"2006","journal-title":"Environmental Science & Policy"},{"key":"ref72","article-title":"Relative effects of land conversion and landuse intensity on terrestrial vertebrate diversity","volume":"13","author":"P Semenchuk","year":"2022","journal-title":"Nature Communications"},{"key":"ref73","doi-asserted-by":"crossref","DOI":"10.1002\/ece3.9328","article-title":"Biogeographic multispecies occupancy models for large-scale survey data","volume":"12","author":"J B Socolar","year":"2022","journal-title":"Ecology and Evolution"},{"key":"ref74","doi-asserted-by":"crossref","first-page":"783","DOI":"10.1111\/2041-210X.12536","article-title":"A multiregion community model for inference about geographic variation in species richness","volume":"7","author":"C Sutherland","year":"2016","journal-title":"Methods in Ecology and Evolution"},{"key":"ref75","doi-asserted-by":"crossref","DOI":"10.5751\/ES-00184-040116","article-title":"Scaling of natal dispersal distances in terrestrial birds and mammals","volume":"4","author":"G D Sutherland","year":"2000","journal-title":"Conservation Ecology"},{"key":"ref76","doi-asserted-by":"crossref","first-page":"73","DOI":"10.4098\/j.at.0001-7051.018.2009","article-title":"Modelling habitat use by Iberian hare Lepus granatensis and European wild rabbit Oryctolagus cuniculus in a mountainous area in northwestern Spain","volume":"55","author":"L Tapia","year":"2009","journal-title":"Acta Theriologica"},{"key":"ref77","doi-asserted-by":"crossref","DOI":"10.3390\/d14020133","article-title":"Mesocarnivore distribution along gradients of anthropogenic disturbance in Mediterranean landscapes","volume":"14","author":"I Torre","year":"2022","journal-title":"Diversity"},{"key":"ref78","author":"R T Torres","year":"2022","journal-title":"Plano Estrat\ufffdgico e de A\ufffd\ufffdo do Javali em Portugal. Relat\ufffdrio Final"},{"key":"ref79","article-title":"Last of the Wild Project","author":"O Venter","year":"2018","journal-title":"NASA SEDAC"},{"key":"ref80","doi-asserted-by":"crossref","DOI":"10.1038\/ncomms12558","article-title":"Sixteen years of change in the global terrestrial human footprint and implications for biodiversity conservation","volume":"7","author":"O Venter","year":"2016","journal-title":"Nature Communications"},{"key":"ref81","doi-asserted-by":"crossref","first-page":"974","DOI":"10.1111\/j.1365-2486.2010.02307.x","article-title":"Challenges in using land use and land cover data for global change studies","volume":"17","author":"P H Verburg","year":"2011","journal-title":"Global Change Biology"},{"key":"ref82","doi-asserted-by":"crossref","first-page":"231","DOI":"10.1016\/S1146-609X(02)01142-6","article-title":"Patch occupancy by stone martens Martes foina in fragmented landscapes of central Spain: the role of fragment size, isolation and habitat structure","volume":"23","author":"E Virg\ufffds","year":"2002","journal-title":"Acta Oecologica"},{"key":"ref83","doi-asserted-by":"crossref","first-page":"113","DOI":"10.1007\/BF03194271","article-title":"Factors shaping European rabbit abundance in continuous and fragmented populations of central Spain","volume":"48","author":"E Virg\ufffds","year":"2003","journal-title":"Acta Theriologica"},{"key":"ref84","doi-asserted-by":"crossref","first-page":"1544","DOI":"10.1139\/z01-114","article-title":"Factors determining \"gaps\" in the distribution of a small carnivore, the common genet (Genetta genetta), in central Spain","volume":"79","author":"E Virg\ufffds","year":"2001","journal-title":"Canadian Journal of Zoology"},{"key":"ref85","first-page":"255","article-title":"Habitat ecology of Martes species in Europe","author":"E Virg\ufffds","year":"2012","journal-title":"Biology and conservation of martens, sables, and fishers: a new synthesis"},{"key":"ref86","article-title":"Last of the Wild Project, Version 2 (LWP-2): Global Human Footprint Dataset (Geographic)","year":"2005","journal-title":"NASA SEDAC"}],"container-title":[],"original-title":[],"deposited":{"date-parts":[[2026,3,4]],"date-time":"2026-03-04T22:47:01Z","timestamp":1772664421000},"score":1,"resource":{"primary":{"URL":"https:\/\/www.ssrn.com\/abstract=6345714"}},"subtitle":[],"short-title":[],"issued":{"date-parts":[[2026]]},"references-count":86,"URL":"https:\/\/doi.org\/10.2139\/ssrn.6345714","relation":{},"subject":[],"published":{"date-parts":[[2026]]},"subtype":"preprint"}}