{"status":"ok","message-type":"work","message-version":"1.0.0","message":{"indexed":{"date-parts":[[2026,7,30]],"date-time":"2026-07-30T13:01:42Z","timestamp":1785416502557,"version":"3.56.0"},"reference-count":70,"publisher":"Frontiers Media SA","license":[{"start":{"date-parts":[[2026,7,30]],"date-time":"2026-07-30T00:00:00Z","timestamp":1785369600000},"content-version":"vor","delay-in-days":0,"URL":"https:\/\/creativecommons.org\/licenses\/by\/4.0\/"}],"funder":[{"DOI":"10.13039\/501100007129","name":"Natural Science Foundation of Shandong Province","doi-asserted-by":"publisher","id":[{"id":"10.13039\/501100007129","id-type":"DOI","asserted-by":"publisher"}]},{"DOI":"10.13039\/501100001809","name":"National Natural Science Foundation of China","doi-asserted-by":"publisher","id":[{"id":"10.13039\/501100001809","id-type":"DOI","asserted-by":"publisher"}]}],"content-domain":{"domain":["frontiersin.org"],"crossmark-restriction":true},"short-container-title":["Front. Plant Sci."],"abstract":"<jats:p>\n                    Drought is a decisive abiotic constraint limiting the growth and yield of jujube, its alleviation is orchestrated by a complex hormonal network that governs the tree\u2019s drought-resistance responses. Based on preliminary evidence that abscisic acid (ABA) and cytokinin play coordinated roles in drought adaptation, this study aimed to clarify their individual and combined effects on regulating drought resistance in sour jujube. Phenotypic responses, anatomical alterations, endogenous hormone profiles, and the expression levels of genes involved in the zeatin biosynthesis pathway after exogenous hormone spraying under drought stress were comprehensively analyzed. The results demonstrated that ABA and cytokinin significantly modulated drought adaptation of sour jujube across multiple aspects. Compared with the control group, ABA spraying xylem vessels number in leaf veins and roots, shortened stem internodes, and elevated the content of salicylic acid (SA), thereby enhancing drought resistance. In contrast, spraying cytokinin treatment increased drought sensitivity, expanded root xylem vessel diameter, and induced a slender stem and increased auxin content. Combined application of ABA and cytokinin was found to further increase vessel number and diameter in leaves and roots, improve branching density, and significantly increase SA, ABA, and jasmonic acid (JA) levels compared with single application of ABA and cytokinin. These factors may collectively enhance the drought resistance of sour jujube. Through weighted gene co-expression network analysis (WGCNA), candidate genes such as\n                    <jats:italic>CGLD27<\/jats:italic>\n                    ,\n                    <jats:italic>GNAT10<\/jats:italic>\n                    , and\n                    <jats:italic>LOC107415885<\/jats:italic>\n                    were obtained. Further analysis found that, both ABA and cytokinin inhibited zeatin biosynthesis, while accelerated trans-zeatin metabolism and delayed cis-zeatin turnover. Spraying of cytokinin enhanced isopentenyladenine degradation, which may reduce resistance, whereas downregulation of\n                    <jats:italic>ZjZOG1<\/jats:italic>\n                    ,\n                    <jats:italic>ZjZOG2<\/jats:italic>\n                    , and\n                    <jats:italic>ZjZOG3<\/jats:italic>\n                    may promote resilience. These findings provide a preliminary mechanistic framework for the role of ABA and cytokinin in drought adaptation of sour jujube.\n                  <\/jats:p>","DOI":"10.3389\/fpls.2026.1901882","type":"journal-article","created":{"date-parts":[[2026,7,30]],"date-time":"2026-07-30T12:22:56Z","timestamp":1785414176000},"update-policy":"https:\/\/doi.org\/10.3389\/crossmark-policy","source":"Crossref","is-referenced-by-count":0,"title":["Exogenous ABA combined with cytokinin enhances drought resistance in Ziziphus jujuba Mill. var. spinosa by regulating xylem development and zeatin signaling pathway"],"prefix":"10.3389","volume":"17","author":[{"given":"Shuhui","family":"Zhou","sequence":"first","affiliation":[{"name":"College of Resources and Environment, Linyi University","place":["Linyi, China"]}],"role":[{"vocabulary":"crossref","role":"author"}]},{"given":"Meng","family":"Li","sequence":"additional","affiliation":[{"name":"College of Resources and Environment, Linyi University","place":["Linyi, China"]}],"role":[{"vocabulary":"crossref","role":"author"}]},{"given":"Mingyu","family":"Jiang","sequence":"additional","affiliation":[{"name":"College of Resources and Environment, Linyi University","place":["Linyi, China"]}],"role":[{"vocabulary":"crossref","role":"author"}]},{"given":"Wei","family":"Zheng","sequence":"additional","affiliation":[{"name":"College of Resources and Environment, Linyi University","place":["Linyi, China"]}],"role":[{"vocabulary":"crossref","role":"author"}]},{"given":"Jin","family":"Yang","sequence":"additional","affiliation":[{"name":"College of Resources and Environment, Linyi University","place":["Linyi, China"]}],"role":[{"vocabulary":"crossref","role":"author"}]},{"given":"Chenghui","family":"Li","sequence":"additional","affiliation":[{"name":"College of Resources and Environment, Linyi University","place":["Linyi, China"]}],"role":[{"vocabulary":"crossref","role":"author"}]},{"given":"Ruixue","family":"Wang","sequence":"additional","affiliation":[{"name":"Shandong Provincial Forestry Protection and Development Service Center","place":["Jinan, China"]}],"role":[{"vocabulary":"crossref","role":"author"}]},{"given":"Yao","family":"Zhao","sequence":"additional","affiliation":[{"name":"College of Resources and Environment, Linyi University","place":["Linyi, China"]}],"role":[{"vocabulary":"crossref","role":"author"}]}],"member":"1965","published-online":{"date-parts":[[2026,7,30]]},"reference":[{"key":"B1","doi-asserted-by":"publisher","first-page":"119","DOI":"10.1002\/csc2.20319","article-title":"Evaluation of durum wheat lines derived from interspecific crosses under drought and heat stress","volume":"61","author":"Aberkane","year":"2021","journal-title":"Crop Sci."},{"key":"B2","doi-asserted-by":"publisher","first-page":"148","DOI":"10.1016\/j.biosystemseng.2017.01.011","article-title":"Leaf thickness to predict plant water status","volume":"156","author":"Afzal","year":"2017","journal-title":"Biosyst. Eng."},{"key":"B3","doi-asserted-by":"publisher","first-page":"2061","DOI":"10.1111\/pce.14597","article-title":"Pepper homeobox abscisic acid signalling-related transcription factor 1, CaHAT1, plays a positive role in drought response","volume":"46","author":"Baek","year":"2023","journal-title":"Plant Cell Environ."},{"key":"B4","doi-asserted-by":"publisher","DOI":"10.3389\/fpls.2020.605069","article-title":"A rapid method for quantifying RNA and phytohormones from a small amount of plant tissue","volume":"11","author":"Cao","year":"2020","journal-title":"Front. Plant Sci."},{"key":"B5","doi-asserted-by":"publisher","first-page":"1733","DOI":"10.1016\/j.molp.2023.09.010","article-title":"TBtools-II: A \"one for all, all for one\" bioinformatics platform for biological big-data mining","volume":"16","author":"Chen","year":"2023","journal-title":"Mol. Plant"},{"key":"B6","doi-asserted-by":"publisher","first-page":"25","DOI":"10.1111\/jipb.12899","article-title":"Abscisic acid dynamics, signaling, and functions in plants","volume":"62","author":"Chen","year":"2020","journal-title":"J. Integr. Plant Biol."},{"key":"B7","doi-asserted-by":"publisher","first-page":"878","DOI":"10.1111\/pbi.13595","article-title":"Cytokinin glucosyl transferases, key regulators of cytokinin homeostasis, have potential value for wheat improvement","volume":"19","author":"Chen","year":"2021","journal-title":"Plant Biotechnol. J."},{"key":"B8","doi-asserted-by":"publisher","first-page":"68","DOI":"10.1038\/s41477-021-01040-7","article-title":"Phosphorylation of SWEET sucrose transporters regulates plant root:shoot ratio under drought","volume":"8","author":"Chen","year":"2022","journal-title":"Nat. Plants"},{"key":"B9","doi-asserted-by":"publisher","first-page":"389","DOI":"10.1007\/s11816-023-00828-7","article-title":"Whole-genome identification and salt- and ABA-induced expression trends of the Nicotiana tabacum CKX gene family","volume":"17","author":"Chen","year":"2023","journal-title":"Plant Biotechnol. Rep."},{"key":"B10","doi-asserted-by":"publisher","first-page":"556","DOI":"10.1186\/s12870-020-02772-y","article-title":"Combined proteomics, metabolomics and physiological analyses of rice growth and grain yield with heavy nitrogen application before and after drought","volume":"20","author":"Du","year":"2020","journal-title":"BMC Plant Biol."},{"key":"B11","doi-asserted-by":"publisher","first-page":"1388","DOI":"10.1093\/pcp\/pcv054","article-title":"HAG3, a histone acetyltransferase, affects UV-B responses by negatively regulating the expression of DNA repair enzymes and sunscreen content in Arabidopsis thaliana","volume":"56","author":"Fina","year":"2015","journal-title":"Plant Cell Physiol."},{"key":"B12","doi-asserted-by":"publisher","first-page":"1388","DOI":"10.1093\/jxb\/erw079","article-title":"Cross-tolerance to biotic and abiotic stresses in plants: a focus on resistance to aphid infestation","volume":"67","author":"Foyer","year":"2016","journal-title":"J. Exp. Bot."},{"key":"B13","doi-asserted-by":"publisher","DOI":"10.1016\/j.scienta.2022.111709","article-title":"The effect of plant growth regulators on xylem differentiation, water and nutrient transport, and bitter pit susceptibility of apple","volume":"31","author":"Griffith","year":"2023","journal-title":"Sci. Hortic."},{"key":"B14","doi-asserted-by":"publisher","first-page":"496","DOI":"10.1186\/s12870-025-06543-5","article-title":"Integrative physiological and transcriptomic analysis provides insights on the molecular basis of ABA-enhanced drought tolerance in pear (Pyrus betulaefolia)","volume":"25","author":"Guo","year":"2025","journal-title":"BMC Plant Biol."},{"key":"B15","doi-asserted-by":"publisher","first-page":"266","DOI":"10.1126\/science.aaz7614","article-title":"The physiology of plant responses to drought","volume":"368","author":"Gupta","year":"2020","journal-title":"Science"},{"key":"B16","doi-asserted-by":"publisher","DOI":"10.1093\/plphys\/kiaf081","article-title":"CYTOKININ DEHYDROGENASE suppression increases intrinsic water-use efficiency and photosynthesis in cotton under drought","volume":"197","author":"Hu","year":"2025","journal-title":"Plant Physiol."},{"key":"B17","doi-asserted-by":"publisher","first-page":"7871","DOI":"10.1021\/acs.jafc.2c01905","article-title":"Wild Jujube (Ziziphus jujuba var. spinosa): A review of its phytonutrients, health benefits, metabolism, and applications","volume":"70","author":"Hua","year":"2022","journal-title":"J. Agric. Food. Chem"},{"key":"B18","doi-asserted-by":"publisher","first-page":"494","DOI":"10.1016\/j.plaphy.2017.07.016","article-title":"Wheat and rye genome confer specific phytohormone profile features and interplay under water stress in two phenotypes of triticale","volume":"118","author":"Hura","year":"2017","journal-title":"Plant Physiol. Biochem."},{"key":"B19","doi-asserted-by":"publisher","first-page":"1252","DOI":"10.1093\/pcp\/pcaa050","article-title":"PsbQ-like protein 3 functions as an assembly factor for the chloroplast NADH dehydrogenase-like complex in Arabidopsis","volume":"61","author":"Ishikawa","year":"2020","journal-title":"Plant Cell Physiol."},{"key":"B20","doi-asserted-by":"publisher","first-page":"549","DOI":"10.1093\/pcp\/pcad017","article-title":"Loss of chloroplast GNAT acetyltransferases results in distinct metabolic phenotypes in Arabidopsis","volume":"64","author":"Ivanauskaite","year":"2023","journal-title":"Plant Cell Physiol."},{"key":"B21","doi-asserted-by":"publisher","first-page":"991","DOI":"10.1007\/s00425-012-1818-4","article-title":"Overexpression of glucosyltransferase UGT85A1 influences trans-zeatin homeostasis and trans-zeatin responses likely through O-glucosylation","volume":"237","author":"Jin","year":"2013","journal-title":"Planta"},{"key":"B22","doi-asserted-by":"publisher","first-page":"645","DOI":"10.1007\/s00425-008-0846-6","article-title":"Role of AUX1 in the control of organ identity during in vitro organogenesis and in mediating tissue specific auxin and cytokinin interaction in Arabidopsis","volume":"229","author":"Kakani","year":"2009","journal-title":"Planta"},{"key":"B23","doi-asserted-by":"publisher","first-page":"605","DOI":"10.1146\/annurev.arplant.54.031902.134802","article-title":"Perception and signal transduction of cytokinins","volume":"54","author":"Kakimoto","year":"2003","journal-title":"Annu. Rev. Plant Biol."},{"key":"B24","doi-asserted-by":"publisher","DOI":"10.1242\/dev.149344","article-title":"Cytokinin signaling in plant development","volume":"145","author":"Kieber","year":"2018","journal-title":"Development"},{"key":"B25","doi-asserted-by":"publisher","first-page":"419","DOI":"10.1007\/s00299-016-2091-y","article-title":"Auxin synthesis gene tms1 driven by tuber-specific promoter alters hormonal status of transgenic potato plants and their responses to exogenous phytohormones","volume":"36","author":"Kolachevskaya","year":"2017","journal-title":"Plant Cell Rep."},{"key":"B26","doi-asserted-by":"publisher","first-page":"956","DOI":"10.1038\/s41893-022-00962-0","article-title":"Anticipating drought-related food security changes","volume":"5","author":"Krishnamurthy","year":"2022","journal-title":"Nat. Sus."},{"key":"B27","doi-asserted-by":"publisher","DOI":"10.1371\/journal.pone.0116819","article-title":"The CRE1 cytokinin pathway is differentially recruited depending on Medicago truncatula root environments and negatively regulates resistance to a pathogen","volume":"10","author":"Laffont","year":"2015","journal-title":"PloS One"},{"key":"B28","doi-asserted-by":"publisher","DOI":"10.1093\/hr\/uhae071","article-title":"Haplotype-resolved T2T reference genomes for wild and domesticated accessions shed new insights into the domestication of jujube","volume":"11","author":"Li","year":"2024","journal-title":"Hortic. Res."},{"key":"B29","doi-asserted-by":"publisher","DOI":"10.1186\/1471-2105-12-323","article-title":"RSEM: accurate transcript quantification from RNA-Seq data with or without a reference genome","volume":"12","author":"Li","year":"2011","journal-title":"BMC Bioinf."},{"key":"B30","doi-asserted-by":"publisher","DOI":"10.3390\/horticulturae8090786","article-title":"Application of morphological and physiological markers for study of drought tolerance in Lilium varieties","volume":"8","author":"Li","year":"2022","journal-title":"Horticulturae"},{"key":"B31","doi-asserted-by":"publisher","DOI":"10.3389\/fpls.2022.1050132","article-title":"The interaction of ABA and ROS in plant growth and stress resistances","volume":"13","author":"Li","year":"2022","journal-title":"Front. Plant Sci."},{"key":"B32","doi-asserted-by":"publisher","DOI":"10.1016\/j.xplc.2024.101133","article-title":"Jasmonic acid plays an important role in mediating retrograde signaling under mitochondrial translational stress to balance plant growth and defense","volume":"6","author":"Li","year":"2025","journal-title":"Plant Commun."},{"key":"B33","doi-asserted-by":"publisher","first-page":"119","DOI":"10.1186\/s13578-021-00633-1","article-title":"Multiple responses contribute to the enhanced drought tolerance of the autotetraploid Ziziphus jujuba Mill. var. spinosa","volume":"11","author":"Li","year":"2021","journal-title":"Cell. Biosci."},{"key":"B34","doi-asserted-by":"publisher","DOI":"10.1016\/j.plaphy.2026.111466","article-title":"ABA-cytokinin crosstalk regulate drought tolerance in Ziziphus jujuba var. spinosa through the phenylpropanoid pathway: insights from physiological and multi-omics integration","volume":"236","author":"Li","year":"2026","journal-title":"Plant Physiol. Biochem."},{"key":"B35","doi-asserted-by":"publisher","first-page":"5457","DOI":"10.1111\/pce.15080","article-title":"Rewatering after drought: Unravelling the drought thresholds and function recovery-limiting factors in maize leaves","volume":"47","author":"Liu","year":"2024","journal-title":"Plant Cell Environ."},{"key":"B36","doi-asserted-by":"publisher","first-page":"119","DOI":"10.1038\/s41438-020-00346-5","article-title":"The historical and current research progress on jujube-a superfruit for the future","volume":"7","author":"Liu","year":"2020","journal-title":"Hortic. Res."},{"key":"B37","doi-asserted-by":"publisher","first-page":"1346","DOI":"10.1016\/s2095-3119(21)63786-3","article-title":"Phenotype and mechanism analysis of plant dwarfing in pear regulated by abscisic acid","volume":"21","author":"Liu","year":"2022","journal-title":"J. Integr. Agric."},{"key":"B38","doi-asserted-by":"publisher","first-page":"550","DOI":"10.1186\/s13059-014-0550-8","article-title":"Moderated estimation of fold change and dispersion for RNA-seq data with DESeq2","volume":"15","author":"Love","year":"2014","journal-title":"Genome Biol."},{"key":"B39","doi-asserted-by":"publisher","first-page":"901","DOI":"10.1016\/j.cj.2020.08.012","article-title":"Exogenous abscisic acid coordinating leaf senescence and transport of assimilates into wheat grains under drought stress by regulating hormones homeostasis","volume":"9","author":"Luo","year":"2021","journal-title":"Crop J."},{"key":"B40","doi-asserted-by":"publisher","DOI":"10.1093\/jpe\/rtae099","article-title":"Variation of six local poplar clones in growth and eco-physiological traits in two types of arid valleys","volume":"18","author":"Mao","year":"2025","journal-title":"J. Plant Ecol."},{"key":"B41","doi-asserted-by":"publisher","first-page":"1189","DOI":"10.1111\/pce.12913","article-title":"Silicon promotes cytokinin biosynthesis and delays senescence in Arabidopsis and Sorghum","volume":"40","author":"Markovich","year":"2017","journal-title":"Plant Cell Environ."},{"key":"B42","doi-asserted-by":"publisher","first-page":"5922","DOI":"10.1073\/pnas.101128798","article-title":"A maize cytokinin gene encoding an O-glucosyltransferase specific to cis-zeatin","volume":"98","author":"Martin","year":"2001","journal-title":"PNAS"},{"key":"B43","doi-asserted-by":"publisher","first-page":"199","DOI":"10.1111\/tpj.13646","article-title":"Interaction network of ABA receptors in grey poplar","volume":"92","author":"Papacek","year":"2017","journal-title":"Plant J."},{"key":"B44","doi-asserted-by":"publisher","DOI":"10.3389\/fpls.2020.00570","article-title":"Coping with water limitation: Hormones that modify plant root xylem development","volume":"11","author":"Ramachandran","year":"2020","journal-title":"Front. Plant Sci."},{"key":"B45","doi-asserted-by":"publisher","first-page":"270","DOI":"10.1007\/s00299-024-03289-6","article-title":"Cytokinin oxidase2-deficient mutants improve panicle and grain architecture through cytokinin accumulation and enhance drought tolerance in indica rice","volume":"43","author":"Rashid","year":"2024","journal-title":"Plant Cell Rep."},{"key":"B46","doi-asserted-by":"publisher","DOI":"10.3389\/fpls.2013.00276","article-title":"The transcriptional response of Arabidopsis leaves to Fe deficiency","volume":"4","author":"Rodriguez-Celma","year":"2013","journal-title":"Front. Plant Sci."},{"key":"B47","doi-asserted-by":"publisher","DOI":"10.1126\/science.adu1470","article-title":"Redox-regulated Aux\/IAA multimerization modulates auxin responses","volume":"389","author":"Roy","year":"2025","journal-title":"Science"},{"key":"B48","doi-asserted-by":"publisher","first-page":"3830","DOI":"10.1016\/j.febslet.2014.09.018","article-title":"Arabidopsis STAYGREEN-LIKE (SGRL) promotes abiotic stress-induced leaf yellowing during vegetative growth","volume":"588","author":"Sakuraba","year":"2014","journal-title":"FEBS Lett."},{"key":"B49","doi-asserted-by":"publisher","first-page":"1299","DOI":"10.1007\/s10068-016-0204-6","article-title":"Phenolic composition, enzyme inhibitory, and antioxidant activity of Bituminaria bituminosa","volume":"25","author":"Sarikurkcu","year":"2016","journal-title":"Food Sci. Biotechnol."},{"key":"B50","doi-asserted-by":"publisher","first-page":"486","DOI":"10.1111\/ppl.12611","article-title":"The role of Euglena gracilis paramylon in modulating xylem hormone levels, photosynthesis and water-use efficiency in Solanum lycopersicum L","volume":"161","author":"Scartazza","year":"2017","journal-title":"Physiol. Plant"},{"key":"B51","doi-asserted-by":"publisher","DOI":"10.3390\/plants10020259","article-title":"Drought stress impacts on plants and different approaches to alleviate its adverse effects","volume":"10","author":"Seleiman","year":"2021","journal-title":"Plants-Basel"},{"key":"B52","doi-asserted-by":"publisher","DOI":"10.1016\/j.indcrop.2024.118808","article-title":"Exogenous ascorbic acid application ameliorates drought stress through improvement in morpho-physiology, nutrient dynamics, stress metabolite production and antioxidant activities recovering cellulosic fibre production in jute (Corchorus olitorius L.)","volume":"217","author":"Sharma","year":"2024","journal-title":"Ind. Crops Prod."},{"key":"B53","doi-asserted-by":"publisher","first-page":"956","DOI":"10.1038\/s41422-019-0254-4","article-title":"Defying gravity: a plant's quest for moisture","volume":"29","author":"Sinclair","year":"2019","journal-title":"Cell Res."},{"key":"B54","doi-asserted-by":"publisher","DOI":"10.1016\/j.plaphy.2023.108237","article-title":"Voltage-dependent anion channel 3 (VDAC3) mediates P. syringae induced ABA-SA signaling crosstalk in Arabidopsis thaliana","volume":"206","author":"Singh","year":"2024","journal-title":"Plant Physiol. Biochem."},{"key":"B55","doi-asserted-by":"publisher","DOI":"10.3390\/ijms18081734","article-title":"Genome-wide organization and expression profiling of the SBP-Box gene family in Chinese jujube (Ziziphus jujuba Mill.)","volume":"18","author":"Song","year":"2017","journal-title":"Int. J. Mol. Sci."},{"key":"B56","doi-asserted-by":"publisher","first-page":"92","DOI":"10.1016\/j.ijfoodmicro.2016.12.015","article-title":"Development of quantitative real-time PCR for detection and enumeration of Enterobacteriaceae","volume":"246","author":"Takahashi","year":"2017","journal-title":"Int. J. Food Microbiol."},{"key":"B57","first-page":"1401","article-title":"Comparison on growth, physiology and medicinal components of Dendrobium huoshanense hybrid and its parents","volume":"31","author":"Wang","year":"2006","journal-title":"Zhongguo. Zhong. Yao. Za. Zhi. = Zhongguo. Zhongyao. Zazhi. = China J. Chin. Materia. Med."},{"key":"B58","doi-asserted-by":"publisher","first-page":"15","DOI":"10.1186\/s43897-022-00036-x","article-title":"A positive feedback regulatory loop, SA-AtNAP-SAG202\/SARD1-ICS1-SA, in SA biosynthesis involved in leaf senescence but not defense response","volume":"2","author":"Wang","year":"2022","journal-title":"Mol. Hortic."},{"key":"B59","doi-asserted-by":"publisher","first-page":"6619","DOI":"10.1093\/jxb\/erad346","article-title":"Wheat cis-zeatin-O-glucosyltransferase cZOGT1 interacts with the Ca2+-dependent lipid binding protein TaZIP to regulate senescence","volume":"74","author":"Wang","year":"2023","journal-title":"J. Exp. Bot."},{"key":"B60","doi-asserted-by":"publisher","first-page":"1","DOI":"10.1093\/plphys\/kiae569","article-title":"The transcription factor TaFDL2-1A functions in auxin metabolism mediated by abscisic acid to regulate shoot growth in wheat","volume":"197","author":"Wang","year":"2024","journal-title":"Plant Physiol."},{"key":"B61","doi-asserted-by":"publisher","first-page":"101","DOI":"10.1007\/s10311-016-0584-0","article-title":"Salicylic acid to decrease plant stress","volume":"15","author":"Wani","year":"2017","journal-title":"Environ. Chem. Lett."},{"key":"B62","doi-asserted-by":"publisher","DOI":"10.1016\/j.ecolind.2022.108934","article-title":"The effects of climate change on the quality of Ziziphus jujuba var. Spinosa in China","volume":"139","author":"Wu","year":"2022","journal-title":"Ecol. Indic."},{"key":"B63","doi-asserted-by":"publisher","DOI":"10.3390\/agriculture11060477","article-title":"Efficacy of root zone temperature increase in root and shoot development and hormone changes in different maize genotypes","volume":"11","author":"Xia","year":"2021","journal-title":"Agriculture-Basel"},{"key":"B64","doi-asserted-by":"publisher","first-page":"5952","DOI":"10.1111\/pce.15575","article-title":"Unlocking the synergy: ABA seed priming enhances drought tolerance in seedlings of sweet sorghum through ABA-IAA crosstalk","volume":"48","author":"Yao","year":"2025","journal-title":"Plant Cell Environ."},{"key":"B65","doi-asserted-by":"publisher","DOI":"10.3390\/plants11182376","article-title":"Structural and functional responses of the heteromorphic leaves of different tree heights on Populus euphratica Oliv. to different soil moisture conditions","volume":"11","author":"Zhai","year":"2022","journal-title":"Plants-Basel"},{"key":"B66","doi-asserted-by":"publisher","first-page":"135","DOI":"10.1016\/j.sjbs.2020.09.022","article-title":"Gibberellin disturbs the balance of endogenesis hormones and inhibits adventitious root development of Pseudostellaria heterophylla through regulating gene expression related to hormone synthesis","volume":"28","author":"Zhang","year":"2021","journal-title":"Saudi. J. Biol. Sci."},{"key":"B67","doi-asserted-by":"publisher","first-page":"2809","DOI":"10.1111\/pbi.70112","article-title":"Natural variation of CsUGT71A60 determines growth and cold tolerance via regulating cytokinin glycosylation in Camellia sinensis","volume":"23","author":"Zhao","year":"2025","journal-title":"Plant Biotechnol. J."},{"key":"B68","doi-asserted-by":"publisher","DOI":"10.1016\/j.xplc.2024.100936","article-title":"Spatially distributed cytokinins: Metabolism signaling, and transport","volume":"5","author":"Zhao","year":"2024","journal-title":"Plant Commun."},{"key":"B69","doi-asserted-by":"publisher","first-page":"1067","DOI":"10.17957\/ijab\/15.0212","article-title":"Involvement of endogenous abscisic acid and cytokinin in photosynthetic performance of different stay green inbred lines of maize under drought","volume":"18","author":"Zhou","year":"2016","journal-title":"Int. J. Agric. Biol."},{"key":"B70","doi-asserted-by":"publisher","DOI":"10.1016\/j.plaphy.2024.108679","article-title":"Integrated anatomical structure, physiological, and transcriptomic analyses to identify differential cold tolerance responses of Ziziphus jujuba mill. 'Yueguang' and its autotetraploid 'Hongguang'","volume":"211","author":"Zhu","year":"2024","journal-title":"Plant Physiol. Biochem."}],"container-title":["Frontiers in Plant Science"],"original-title":[],"link":[{"URL":"https:\/\/www.frontiersin.org\/articles\/10.3389\/fpls.2026.1901882\/full","content-type":"unspecified","content-version":"vor","intended-application":"similarity-checking"}],"deposited":{"date-parts":[[2026,7,30]],"date-time":"2026-07-30T12:22:58Z","timestamp":1785414178000},"score":1,"resource":{"primary":{"URL":"https:\/\/www.frontiersin.org\/articles\/10.3389\/fpls.2026.1901882\/full"}},"subtitle":[],"short-title":[],"issued":{"date-parts":[[2026,7,30]]},"references-count":70,"alternative-id":["10.3389\/fpls.2026.1901882"],"URL":"https:\/\/doi.org\/10.3389\/fpls.2026.1901882","relation":{},"ISSN":["1664-462X"],"issn-type":[{"value":"1664-462X","type":"electronic"}],"subject":[],"published":{"date-parts":[[2026,7,30]]},"article-number":"1901882"}}