{"status":"ok","message-type":"work","message-version":"1.0.0","message":{"indexed":{"date-parts":[[2026,8,31]],"date-time":"2026-08-31T05:18:19Z","timestamp":1788153499023,"version":"build-2803163510"},"reference-count":72,"publisher":"Frontiers Media SA","license":[{"start":{"date-parts":[[2026,8,31]],"date-time":"2026-08-31T00:00:00Z","timestamp":1788134400000},"content-version":"vor","delay-in-days":0,"URL":"https:\/\/creativecommons.org\/licenses\/by\/4.0\/"}],"content-domain":{"domain":["frontiersin.org"],"crossmark-restriction":true},"short-container-title":["Front. Plant Sci."],"abstract":"<jats:p>Microgreens are considered a good source of nutritional and functional compounds, offering humans significant health benefits. Biostimulants are recognized as a sustainable tool to elicit growth and development of vegetables. The aim of this research was to evaluate the effects of a plant protein hydrolysate (PH) and a seaweed extract (SWE) \u2013 supplied via seed priming or root application \u2013 on pea and arugula microgreens performance. Moreover, a partial budget analysis was conducted to assess the economic implication of biostimulants application. Hydropriming and PH-priming treatments enhanced pea microgreens yield compared to the root application treatment\u2019s counterpart by 53.2% and 33.3%, respectively. Nitrates were increased by SWE application and reduced by PH supply, in both species, compared with the not biostimulated plants, particularly when biostimulants were applied via root application [SWE,\u00a0+\u00a026.6% (pea); PH, -19.9% (pea), SWE,\u00a0+\u00a06.5% (arugula); PH, -24.3% (arugula)]. The combination of biostimulants and priming enhanced ascorbic acid compared to the control \u00d7 root application (negative control) in pea microgreens by 20.9%, whereas, in arugula, either the biostimulants or priming alone significantly improved the aforesaid parameter. Biostimulants boosted total polyphenol content in pea microgreens, supplied via priming (SWE,\u00a0+\u00a021.5%; PH, 22.8%, compared to the hydropriming) or root application (SWE,\u00a0+\u00a015.7%; PH, 14.6%, compared with the negative control). Total polyphenols in arugula microgreens increased when PH and SWE were supplied via priming (+14.3%, for both biostimulants, compared to the hydropriming). Biostimulants significantly increased the antioxidant capacity (DPPH), although the method of application did not significantly affect DPPH. These findings confirmed the efficacy of biostimulants in modulating microgreens qualitative features. Furthermore, the partial budget analysis underlined that the use of both biostimulants, independently of the type of application, is economically suitable for pea, but not always convenient for arugula. The application of PH or SWE, especially via root application, was a suitable strategy to increase the nutritive and functional values of pea and arugula microgreens, contributing to their potential as valuable novel foods for human consumption.<\/jats:p>","DOI":"10.3389\/fpls.2026.1913861","type":"journal-article","created":{"date-parts":[[2026,8,31]],"date-time":"2026-08-31T04:28:07Z","timestamp":1788150487000},"update-policy":"https:\/\/doi.org\/10.3389\/crossmark-policy","source":"Crossref","is-referenced-by-count":0,"title":["Combining non-microbial biostimulants and different application methods affects yield, antioxidant activity, nutritional and functional value of pea and arugula microgreens"],"prefix":"10.3389","volume":"17","author":[{"given":"Beppe Benedetto","family":"Consentino","sequence":"first","affiliation":[{"name":"Department of Agricultural, Food and Forest Sciences, University of Palermo","place":["Palermo, Italy"]}],"role":[{"vocabulary":"crossref","role":"author"}]},{"given":"Pietro","family":"Bellitto","sequence":"additional","affiliation":[{"name":"Department of Agricultural, Food and Forest Sciences, University of Palermo","place":["Palermo, Italy"]}],"role":[{"vocabulary":"crossref","role":"author"}]},{"given":"Fabiana","family":"Mancuso","sequence":"additional","affiliation":[{"name":"Department of Agricultural, Food and Forest Sciences, University of Palermo","place":["Palermo, Italy"]}],"role":[{"vocabulary":"crossref","role":"author"}]},{"given":"Francesco","family":"Salamone","sequence":"additional","affiliation":[{"name":"Department of Agricultural, Food and Forest Sciences, University of Palermo","place":["Palermo, Italy"]}],"role":[{"vocabulary":"crossref","role":"author"}]},{"given":"Theodora","family":"Ntanasi","sequence":"additional","affiliation":[{"name":"Laboratory of Vegetable Production, Department of Crop Science, Agricultural University of Athens","place":["Athens, Greece"]}],"role":[{"vocabulary":"crossref","role":"author"}]},{"given":"Georgia","family":"Ntatsi","sequence":"additional","affiliation":[{"name":"Laboratory of Vegetable Production, Department of Crop Science, Agricultural University of Athens","place":["Athens, Greece"]}],"role":[{"vocabulary":"crossref","role":"author"}]},{"given":"Simona","family":"Napoli","sequence":"additional","affiliation":[{"name":"Department of Agricultural, Food and Forest Sciences, University of Palermo","place":["Palermo, Italy"]}],"role":[{"vocabulary":"crossref","role":"author"}]},{"given":"Filippo","family":"Sgroi","sequence":"additional","affiliation":[{"name":"Department of Economics, Business and Statistics, University of Palermo","place":["Palermo, Italy"]}],"role":[{"vocabulary":"crossref","role":"author"}]},{"given":"Salvatore","family":"La Bella","sequence":"additional","affiliation":[{"name":"Department of Agricultural, Food and Forest Sciences, University of Palermo","place":["Palermo, Italy"]},{"name":"Research Consortium for the Development of Innovative Agro-Environmental Systems (Corissia)","place":["Palermo, Italy"]}],"role":[{"vocabulary":"crossref","role":"author"}]},{"given":"Leo","family":"Sabatino","sequence":"additional","affiliation":[{"name":"Department of Agricultural, Food and Forest Sciences, University of Palermo","place":["Palermo, Italy"]}],"role":[{"vocabulary":"crossref","role":"author"}]}],"member":"1965","published-online":{"date-parts":[[2026,8,31]]},"reference":[{"key":"B1","doi-asserted-by":"publisher","DOI":"10.3390\/horticulturae6020028","article-title":"Effect of seaweed extract on productivity and quality attributes of four onion cultivars","volume":"6","author":"Abbas","year":"2020","journal-title":"Horticulturae"},{"key":"B2","doi-asserted-by":"publisher","DOI":"10.3390\/plants10030531","article-title":"Biostimulant properties of seaweed extracts in plants: implications towards sustainable crop production","volume":"10","author":"Ali","year":"2021","journal-title":"Plants"},{"key":"B3","doi-asserted-by":"publisher","first-page":"116","DOI":"10.15835\/nbha48111806","article-title":"Seed priming as a promising technique to improve growth, chlorophyll, photosynthesis and nutrient contents in cucumber seedlings","volume":"48","author":"Anwar","year":"2020","journal-title":"Not. Bot. Horti Agrobot Cluj Napoca"},{"key":"B4","first-page":"501","article-title":"The effect of seed priming on germination and seedling growth of watermelon (Citrullus lanatus)","volume":"4","author":"Armin","year":"2010","journal-title":"Adv. Environ. Biol."},{"key":"B5","doi-asserted-by":"publisher","DOI":"10.3390\/agronomy12081943","article-title":"Improvement of photosynthetic pigment characteristics, mineral content, and antioxidant activity of lettuce (Lactuca sativa L.) by arbuscular mycorrhizal fungus and seaweed extract foliar application","volume":"12","author":"Asadi","year":"2022","journal-title":"Agronomy"},{"key":"B6","doi-asserted-by":"publisher","first-page":"6213","DOI":"10.1038\/s41598-025-85860-z","article-title":"Nutritional quality profiles of six microgreens","volume":"15","author":"Balik","year":"2025","journal-title":"Sci. Rep."},{"key":"B7","doi-asserted-by":"publisher","DOI":"10.3390\/molecules28020867","article-title":"Microgreens\u2014a comprehensive review of bioactive molecules and health benefits","volume":"28","author":"Bhaswant","year":"2023","journal-title":"Molecules"},{"key":"B8","doi-asserted-by":"publisher","first-page":"96","DOI":"10.47115\/bsagriculture.1595301","article-title":"Determination of quality characteristics in mature parsley (Petroselinum hortense) plants, parsley microgreens, and primed parsley microgreens","volume":"8","author":"\u00c7ak\u0131r","year":"2025","journal-title":"BSJ Agric."},{"key":"B9","doi-asserted-by":"publisher","DOI":"10.3390\/nu12020334","article-title":"Defining a healthy diet: Evidence for the role of contemporary dietary patterns in health and disease","volume":"12","author":"Cena","year":"2020","journal-title":"Nutrients"},{"key":"B10","doi-asserted-by":"publisher","first-page":"2374","DOI":"10.1021\/jf970055t","article-title":"Antioxidant activities of caffeic acid and its related hydroxycinnamic acid compounds","volume":"45","author":"Chen","year":"1997","journal-title":"J. Agric. Food. Chem."},{"key":"B11","doi-asserted-by":"publisher","DOI":"10.26353\/j.itahort\/2023.3.3747","article-title":"Hydropriming and plant-based protein hydrolysate biostimulants impact the primary and secondary metabolites of brassicaceous microgreens differentially","volume":"30","author":"Ciriello","year":"2023","journal-title":"Italus Hortus"},{"key":"B12","doi-asserted-by":"publisher","first-page":"5921","DOI":"10.1002\/jsfa.13416","article-title":"Plant-derived biostimulant as priming agents enhanced antioxidant and nutritive properties in brassicaceous microgreens","volume":"104","author":"Ciriello","year":"2024","journal-title":"J. Sci. Food Agric."},{"key":"B13","doi-asserted-by":"publisher","DOI":"10.3390\/horticulturae8050409","article-title":"Biostimulatory action of a plant-derived protein hydrolysate on morphological traits, photosynthetic parameters, and mineral composition of two basil cultivars grown hydroponically under variable electrical conductivity","volume":"8","author":"Ciriello","year":"2022","journal-title":"Horticulturae"},{"key":"B14","doi-asserted-by":"publisher","DOI":"10.3389\/fpls.2017.02202","article-title":"Biostimulant action of protein hydrolysates: unraveling their effects on plant physiology and microbiome","volume":"8","author":"Colla","year":"2017","journal-title":"Front. Plant Sci."},{"key":"B15","doi-asserted-by":"publisher","first-page":"28","DOI":"10.1016\/j.scienta.2015.08.037","article-title":"Protein hydrolysates as biostimulants in horticulture","volume":"196","author":"Colla","year":"2015","journal-title":"Sci. Hortic."},{"key":"B16","doi-asserted-by":"publisher","first-page":"27","DOI":"10.17660\/ActaHortic.2016.1148.3","article-title":"Protein hydrolysate-based biostimulants: origin, biological activity and application methods","volume":"1148","author":"Colla","year":"2015","journal-title":"Acta Hortic."},{"key":"B17","doi-asserted-by":"publisher","DOI":"10.3390\/horticulturae7120549","article-title":"Seaweed extract improves Lagenaria siceraria young shoot production, mineral profile and functional quality","volume":"7","author":"Consentino","year":"2021","journal-title":"Horticulturae"},{"key":"B18","doi-asserted-by":"publisher","DOI":"10.3390\/horticulturae8080722","article-title":"Grafting eggplant onto underutilized solanum species and biostimulatory action of Azospirillum brasilense modulate growth, yield, NUE and nutritional and functional traits","volume":"8","author":"Consentino","year":"2022","journal-title":"Horticulturae"},{"key":"B19","first-page":"7702","article-title":"Studies concerning the extraction of chlorophyll and total carotenoids from vegetables","volume":"17","author":"Costache","year":"2012","journal-title":"Rom. Biotechnol. Lett."},{"key":"B20","doi-asserted-by":"publisher","DOI":"10.3390\/foods8070250","article-title":"Evaluation of the bioaccessibility of antioxidant bioactive compounds and minerals of four genotypes of Brassicaceae microgreens","volume":"8","author":"de la Fuente","year":"2019","journal-title":"Foods"},{"key":"B21","doi-asserted-by":"publisher","first-page":"981","DOI":"10.1021\/acsfoodscitech.3c00040","article-title":"Brassicaceae microgreens: phytochemical compositions, influences of growing practices, postharvest technology, health, and food applications","volume":"3","author":"Dereje","year":"2023","journal-title":"ACS Food Sci. Technol."},{"key":"B22","doi-asserted-by":"publisher","DOI":"10.3390\/horticulturae9050592","article-title":"Synergistic effect of a plant-derived protein hydrolysate and arbuscular mycorrhizal fungi on eggplant grown in open fields: a two-year study","volume":"9","author":"Di Miceli","year":"2023","journal-title":"Horticulturae"},{"key":"B23","doi-asserted-by":"publisher","DOI":"10.3390\/su16177436","article-title":"The eliciting effect of aqueous extracts from Ascophyllum nodosum algae on the cultivation of arugula (Eruca sativa Mill.) microgreens","volume":"16","author":"Dryga\u015b","year":"2024","journal-title":"Sustainability"},{"key":"B24","doi-asserted-by":"publisher","DOI":"10.1016\/j.cofs.2022.100863","article-title":"Advances and emerging trends in cultivation substrates for growing sprouts and microgreens toward safe and sustainable agriculture","volume":"46","author":"Du","year":"2022","journal-title":"Curr. Opin. Food Sci."},{"key":"B25","doi-asserted-by":"publisher","DOI":"10.3390\/horticulturae7090279","article-title":"Protein hydrolysate combined with hydroponics divergently modifies growth and shuffles pigments and free amino acids of carrot and dill microgreens","volume":"7","author":"El-Nakhel","year":"2021","journal-title":"Horticulturae"},{"key":"B26","doi-asserted-by":"publisher","first-page":"406","DOI":"10.1039\/an9588300406","article-title":"The colorimetric determination of phosphorus","volume":"83","author":"Fogg","year":"1958","journal-title":"Analyst"},{"key":"B27","doi-asserted-by":"publisher","DOI":"10.3390\/data6050050","article-title":"Dataset on the effects of anti-insect nets of different porosity on mineral and organic acids profile of Cucurbita pepo L. Fruits and leaves","volume":"6","author":"Formisano","year":"2021","journal-title":"Data"},{"key":"B28","doi-asserted-by":"publisher","DOI":"10.3390\/plants9070922","article-title":"Stand-alone and combinatorial effects of plant-based biostimulants on the production and leaf quality of perennial wall rocket","volume":"9","author":"Giordano","year":"2020","journal-title":"Plants"},{"key":"B29","doi-asserted-by":"crossref","first-page":"1","DOI":"10.1016\/B978-0-12-823048-0.00018-6","article-title":"Chapter 1 - Categories of various plant biostimulants \u2013 mode of application and shelf-life","volume-title":"Biostimulants for Crops From Seed Germination to Plant Development","author":"Gupta","year":"2021"},{"key":"B30","doi-asserted-by":"publisher","first-page":"1381","DOI":"10.1007\/s11738-012-1186-5","article-title":"Seed priming for abiotic stress tolerance: an overview","volume":"35","author":"Jisha","year":"2013","journal-title":"Acta Physiol. Plant"},{"key":"B31","doi-asserted-by":"publisher","DOI":"10.3390\/agronomy13010009","article-title":"Influence of microalgae Planktochlorella nurekis clones on seed germination","volume":"13","author":"Karbarz","year":"2023","journal-title":"Agronomy"},{"key":"B32","doi-asserted-by":"publisher","first-page":"5089","DOI":"10.1111\/ijfs.16607","article-title":"Nutritional quality and bioactive compounds of arugula (Eruca sativa L.) sprouts and microgreens","volume":"58","author":"Komeroski","year":"2023","journal-title":"Int. J. Food Sci. Technol."},{"key":"B33","doi-asserted-by":"publisher","first-page":"599","DOI":"10.1007\/s10811-019-01938-y","article-title":"Science behind biostimulant action of seaweed extract on growth and crop yield: insights into transcriptional changes in roots of maize treated with Kappaphycus alvarezii seaweed extract under soil moisture stressed conditions","volume":"32","author":"Kumar","year":"2020","journal-title":"J. Appl. Phycol."},{"key":"B34","doi-asserted-by":"publisher","first-page":"107","DOI":"10.1016\/j.foodchem.2018.10.098","article-title":"Functional quality in novel food sources: Genotypic variation in the nutritive and phytochemical composition of thirteen microgreens species","volume":"277","author":"Kyriacou","year":"2019","journal-title":"Food Chem."},{"key":"B35","doi-asserted-by":"publisher","DOI":"10.3390\/foods10061333","article-title":"Preharvest nutrient deprivation reconfigures nitrate, mineral, and phytochemical content of microgreens","volume":"10","author":"Kyriacou","year":"2021","journal-title":"Foods"},{"key":"B36","doi-asserted-by":"publisher","first-page":"103","DOI":"10.1016\/j.tifs.2016.09.005","article-title":"Micro-scale vegetable production and the rise of microgreens","volume":"57","author":"Kyriacou","year":"2016","journal-title":"Trends Food Sci. Technol."},{"key":"B37","doi-asserted-by":"publisher","DOI":"10.3390\/plants10061139","article-title":"Impact of Ecklonia maxima seaweed extract and Mo foliar treatments on biofortification, spinach yield, quality and NUE","volume":"10","author":"La Bella","year":"2021","journal-title":"Plants"},{"key":"B38","doi-asserted-by":"publisher","DOI":"10.3390\/horticulturae9111217","article-title":"Shoot yield and mineral nutrient concentrations of five microgreens in the Brassicaceae family affected by fertigation rate","volume":"9","author":"Li","year":"2023","journal-title":"Horticulturae"},{"key":"B39","doi-asserted-by":"publisher","DOI":"10.3390\/horticulturae7060129","article-title":"Shoot production and mineral nutrients of five microgreens as affected by hydroponic substrate type and post-emergent fertilization","volume":"7","author":"Li","year":"2021","journal-title":"Horticulturae"},{"key":"B40","doi-asserted-by":"publisher","DOI":"10.3390\/ijms24119714","article-title":"Biostimulant properties of protein hydrolysates: recent advances and future challenges","volume":"24","author":"Mal\u00e9cange","year":"2023","journal-title":"IJMS"},{"key":"B41","doi-asserted-by":"publisher","first-page":"43","DOI":"10.1016\/j.ecoenv.2017.08.017","article-title":"Influence of biostimulants-seed-priming on Ceratotheca triloba germination and seedling growth under low temperatures, low osmotic potential and salinity stress","volume":"147","author":"Masondo","year":"2018","journal-title":"Ecotoxicol. Environ. Saf."},{"key":"B42","doi-asserted-by":"publisher","first-page":"571","DOI":"10.1016\/j.foodchem.2004.10.006","article-title":"Determination of the total phenolic, flavonoid and proline contents in Burkina Fasan honey, as well as their radical scavenging activity","volume":"91","author":"Meda","year":"2005","journal-title":"Food Chem."},{"key":"B43","first-page":"012113","article-title":"Determination of sodium, potassium, magnesium, and calcium minerals level in fresh and boiled broccoli and cauliflower by atomic absorption spectrometry","author":"Nerdy","year":"2018"},{"key":"B44","doi-asserted-by":"publisher","first-page":"495","DOI":"10.13128\/ahsc8127","article-title":"Effects of foliar application of glycine and glutamine amino acids on growth and quality of sweet basil","volume":"33","author":"Noroozlo","year":"2019","journal-title":"Adv. Hortic. Sci."},{"key":"B45","doi-asserted-by":"publisher","first-page":"251","DOI":"10.1016\/j.mayocpiqo.2019.05.001","article-title":"A clinician\u2019s guide to healthy eating for cardiovascular disease prevention","volume":"3","author":"Pallazola","year":"2019","journal-title":"Mayo Clinic Proceedings: Innovations Qual. Outcomes"},{"key":"B46","doi-asserted-by":"publisher","DOI":"10.3390\/agriculture12091329","article-title":"Mugwort (Artemisia vulgaris L.) aqueous extract: hormesis and biostimulant activity for seed germination and seedling growth in vegetable crops","volume":"12","author":"Pannacci","year":"2022","journal-title":"Agriculture"},{"key":"B47","doi-asserted-by":"publisher","DOI":"10.3390\/horticulturae10101041","article-title":"Protein hydrolysates - production, effects on plant metabolism, and use in agriculture","volume":"10","author":"Paskovi\u0107","year":"2024","journal-title":"Horticulturae"},{"key":"B48","doi-asserted-by":"publisher","DOI":"10.3390\/horticulturae11010039","article-title":"Bacillus thuringiensis and Trichoderma asperellum as biostimulants in hydroponic tendril pea (Pisum sativum) microgreens","volume":"11","author":"P\u00e9rez-Leal","year":"2025","journal-title":"Horticulturae"},{"key":"B49","first-page":"2239","article-title":"Microgreens: A super food for nutritional security in the 21st century-a review","volume":"40","author":"Rai","year":"2022","journal-title":"Environ. Ecol."},{"key":"B50","doi-asserted-by":"publisher","first-page":"311","DOI":"10.1007\/s10722-022-01506-3","article-title":"Microgreens: A next generation nutraceutical for multiple disease management and health promotion","volume":"70","author":"Rizvi","year":"2023","journal-title":"Genet. Resour. Crop Evol."},{"key":"B51","doi-asserted-by":"publisher","first-page":"359","DOI":"10.1023\/A:1010781500705","article-title":"Regulation of source\/sink relations by cytokinins","volume":"32","author":"Roitsch","year":"2000","journal-title":"Plant Growth Regul."},{"key":"B52","doi-asserted-by":"publisher","DOI":"10.3389\/fpls.2020.00040","article-title":"Editorial: biostimulants in agriculture","volume":"11","author":"Rouphael","year":"2020","journal-title":"Front. Plant Sci."},{"key":"B53","doi-asserted-by":"publisher","DOI":"10.3390\/agronomy11061023","article-title":"Protein hydrolysates and Mo-biofortification interactively modulate plant performance and quality of \u2018Canasta\u2019 lettuce grown in a protected environment","volume":"11","author":"Sabatino","year":"2021","journal-title":"Agronomy"},{"key":"B54","doi-asserted-by":"crossref","DOI":"10.1201\/9781003346173-13","article-title":"Seedling and seed priming in regulating secondary metabolite level for stress tolerance","volume-title":"Biology and Biotechnology of Environmental Stress Tolerance in Plants","author":"Salami","year":"2023"},{"key":"B55","doi-asserted-by":"publisher","first-page":"11800","DOI":"10.1021\/jf802362g","article-title":"Effects of an Alfalfa protein hydrolysate on the gene expression and activity of enzymes of the tricarboxylic acid (TCA) cycle and nitrogen metabolism in Zea mays L","volume":"56","author":"Schiavon","year":"2008","journal-title":"J. Agric. Food. Chem."},{"key":"B56","doi-asserted-by":"publisher","first-page":"1765","DOI":"10.1002\/jsfa.5968","article-title":"Effects of brassinosteroid analogues on total phenols, antioxidant activity, sugars, organic acids and yield of field grown endive (Cichorium endivia L.)","volume":"93","author":"Serna","year":"2013","journal-title":"J. Sci. Food Agric."},{"key":"B57","doi-asserted-by":"publisher","DOI":"10.3390\/agronomy7030061","article-title":"Response of chlorophyll, carotenoid and spad-502 measurement to salinity and nutrient stress in wheat (Triticum aestivum L.)","volume":"7","author":"Shah","year":"2017","journal-title":"Agronomy"},{"key":"B58","doi-asserted-by":"publisher","first-page":"805","DOI":"10.1080\/01904167.2012.663436","article-title":"Seed priming improves early seedling growth and nutrient uptake in mungbean","volume":"35","author":"Shah","year":"2012","journal-title":"J. Plant Nutr."},{"key":"B59","first-page":"257","article-title":"Effect of foliar spraying with amino acids and seaweed extract on growth chemical constitutes, yield and its quality of celeriac plant","volume":"58","author":"Shehata","year":"2011","journal-title":"Eur. J. Sci. Res."},{"key":"B60","doi-asserted-by":"publisher","DOI":"10.3390\/agronomy11071297","article-title":"Plant biostimulants: A categorical review, their implications for row crop production, and relation to soil health indicators","volume":"11","author":"Sible","year":"2021","journal-title":"Agronomy"},{"key":"B61","doi-asserted-by":"publisher","DOI":"10.3390\/plants13020210","article-title":"Amino acids biostimulants and protein hydrolysates in agricultural sciences","volume":"13","author":"Sun","year":"2024","journal-title":"Plants"},{"key":"B62","doi-asserted-by":"publisher","DOI":"10.13057\/nusbiosci\/n020204","article-title":"Effect of seaweed extracts on growth and yield of rice plants","volume":"8","author":"Sunarpi","year":"2011","journal-title":"Asian J. Trop. Bio\/Technol."},{"key":"B63","doi-asserted-by":"publisher","DOI":"10.3390\/agriculture13020439","article-title":"Seed priming treatments to improve heat stress tolerance of garden pea (Pisum sativum L.)","volume":"13","author":"Tamind\u017ei\u0107","year":"2023","journal-title":"Agriculture"},{"key":"B64","doi-asserted-by":"publisher","DOI":"10.1016\/j.foodres.2020.110039","article-title":"Metabolomic insight into the profile, in vitro bioaccessibility and bioactive properties of polyphenols and glucosinolates from four Brassicaceae microgreens","volume":"140","author":"Tomas","year":"2021","journal-title":"Food Res. Int."},{"key":"B65","doi-asserted-by":"publisher","DOI":"10.3390\/agronomy13010145","article-title":"Effect of application of biostimulants on the biomass, nitrate, pigments, and antioxidants content in radish and turnip microgreens","volume":"13","author":"Toscano","year":"2023","journal-title":"Agronomy"},{"key":"B66","doi-asserted-by":"publisher","first-page":"47","DOI":"10.1007\/s10709-020-00093-4","article-title":"Epigenetic memory and priming in plants","volume":"148","author":"Turgut-Kara","year":"2020","journal-title":"Genetica"},{"key":"B67","doi-asserted-by":"publisher","DOI":"10.3390\/nu15041022","article-title":"The role of consumption of molybdenum biofortified crops in bone homeostasis and healthy aging","volume":"15","author":"Vasto","year":"2023","journal-title":"Nutrients"},{"key":"B68","doi-asserted-by":"publisher","DOI":"10.3390\/horticulturae10050459","article-title":"Joint action of Trichoderma atroviride and a vegetal derived-protein hydrolysate improves performances of woodland strawberry in Italy","volume":"10","author":"Vultaggio","year":"2024","journal-title":"Horticulturae"},{"key":"B69","doi-asserted-by":"publisher","first-page":"1441","DOI":"10.21608\/jpp.2014.64669","article-title":"Effect of seaweed extract and amino acids on growth and productivity and some biocostituents of common bean (Phaseolus vulgaris) plants","volume":"5","author":"Zewail","year":"2014","journal-title":"J. Plant Prod."},{"key":"B70","doi-asserted-by":"publisher","first-page":"203","DOI":"10.1016\/j.tifs.2020.02.031","article-title":"A review on the effects of light-emitting diode (LED) light on the nutrients of sprouts and microgreens","volume":"99","author":"Zhang","year":"2020","journal-title":"Trends Food Sci. Technol."},{"key":"B71","doi-asserted-by":"publisher","DOI":"10.3389\/fpls.2022.1072782","article-title":"The differential modulation of secondary metabolism induced by a protein hydrolysate and a seaweed extract in tomato plants under salinity","volume":"13","author":"Zhang","year":"2023","journal-title":"Front. Plant Sci."},{"key":"B72","doi-asserted-by":"publisher","DOI":"10.1371\/journal.pone.0257236","article-title":"Effects of seed priming treatments on the germination and development of two rapeseed (Brassica napus L.) varieties under the co-influence of low temperature and drought","volume":"16","author":"Zhu","year":"2021","journal-title":"PloS One"}],"container-title":["Frontiers in Plant Science"],"original-title":[],"link":[{"URL":"https:\/\/www.frontiersin.org\/articles\/10.3389\/fpls.2026.1913861\/full","content-type":"unspecified","content-version":"vor","intended-application":"similarity-checking"}],"deposited":{"date-parts":[[2026,8,31]],"date-time":"2026-08-31T04:28:10Z","timestamp":1788150490000},"score":1,"resource":{"primary":{"URL":"https:\/\/www.frontiersin.org\/articles\/10.3389\/fpls.2026.1913861\/full"}},"subtitle":[],"short-title":[],"issued":{"date-parts":[[2026,8,31]]},"references-count":72,"alternative-id":["10.3389\/fpls.2026.1913861"],"URL":"https:\/\/doi.org\/10.3389\/fpls.2026.1913861","relation":{},"ISSN":["1664-462X"],"issn-type":[{"value":"1664-462X","type":"electronic"}],"subject":[],"published":{"date-parts":[[2026,8,31]]},"article-number":"1913861"}}