{"status":"ok","message-type":"work","message-version":"1.0.0","message":{"indexed":{"date-parts":[[2026,8,31]],"date-time":"2026-08-31T06:15:31Z","timestamp":1788156931294,"version":"build-2803163510"},"reference-count":56,"publisher":"Frontiers Media SA","license":[{"start":{"date-parts":[[2026,8,31]],"date-time":"2026-08-31T00:00:00Z","timestamp":1788134400000},"content-version":"vor","delay-in-days":0,"URL":"https:\/\/creativecommons.org\/licenses\/by\/4.0\/"}],"content-domain":{"domain":["frontiersin.org"],"crossmark-restriction":true},"short-container-title":["Front. Plant Sci."],"abstract":"<jats:p>\n                    Peach gummosis, caused primarily by\n                    <jats:italic>Botryosphaeria dothidea sensu lato<\/jats:italic>\n                    , is a destructive canker disease with few environmentally friendly management options. Here we report an uncontrolled pre\/post field demonstration of a \u03b3-polyglutamic acid-based artificial biofilm \u2014 a biodegradable delivery platform whose agent-agnostic design remains untested \u2014 encapsulating\n                    <jats:italic>Bacillus amyloliquefaciens<\/jats:italic>\n                    KNU-28 across 143 trees in 18 commercial peach orchards in South Korea. Severity declined from a pre-treatment mean of 4.12 (April) to 2.32 by August (76.2% of 143 trees improved by \u22651 point) and to 1.78 by October in three regions (88.3% of 128 trees). In an ordinal cumulative-link mixed model, severity fell below the April baseline at every later visit; the average monthly odds ratio was 0.70 (95% CI 0.66\u20130.75); alternative model specifications agreed. Operator-independent image-based \u0394E quantification across 74 trees showed a concordant cohort-level decline (lesion-area declines of 19\u201330%; significant in one of four regions after FDR correction) but agreed only weakly with the ordinal score at tree level (Spearman \u03c1 = 0.16), and is complementary rather than confirmatory. We sequenced fungal ITS2 from 23 bark samples. Multivariate dispersion differed among bark groups (PERMDISP F\u00a0=\u00a020.7, p &amp;lt; 0.001), and healthy and gummosis-affected bark did not separate under compositionally aware metrics (Aitchison\/robust CLR, p &amp;gt; 0.34); the nominally significant Bray\u2013Curtis PERMANOVA (R\n                    <jats:sup>2<\/jats:sup>\n                    = 0.248, p = 0.001) therefore cannot be read as a centroid difference. Early- and later-treatment diseased communities did not differ (p = 0.542; all genera FDR q &amp;gt; 0.5). Gummosis-enrolled bark showed higher relative abundances of\n                    <jats:italic>Wickerhamomyces<\/jats:italic>\n                    ,\n                    <jats:italic>Cytospora<\/jats:italic>\n                    and\n                    <jats:italic>Botryosphaeria<\/jats:italic>\n                    (dominant ASVs assigned to\n                    <jats:italic>B. qingyuanensis<\/jats:italic>\n                    ) than healthy bark, though none survived FDR correction. Because the 19 diseased libraries came from only 10 trees, the genus\u2013severity correlations were replaced by a paired analysis, in which no genus differed nominally. Together, this demonstration establishes the operational feasibility of field-scale application and a parameter-transparent lesion-quantification method; the severity decline and mycobiota trends are accompanying observations. Because untreated controls, strain-level KNU-28 tracking, verification of delivery to bark and a powered microbiome cohort were absent, the data do not establish causal efficacy and warrant randomized, untreated-control trials.\n                  <\/jats:p>","DOI":"10.3389\/fpls.2026.1933566","type":"journal-article","created":{"date-parts":[[2026,8,31]],"date-time":"2026-08-31T05:39:13Z","timestamp":1788154753000},"update-policy":"https:\/\/doi.org\/10.3389\/crossmark-policy","source":"Crossref","is-referenced-by-count":0,"title":["A \u03b3-polyglutamic acid-based artificial biofilm for field delivery of Bacillus amyloliquefaciens KNU-28: an uncontrolled multi-orchard demonstration on peach gummosis with exploratory bark fungal community profiling"],"prefix":"10.3389","volume":"17","author":[{"given":"Min-Kyu","family":"Park","sequence":"first","affiliation":[{"name":"Department of Applied Biosciences, Kyungpook National University","place":["Daegu, Republic of Korea"]},{"name":"Kyungpook National University Next-Generation Sequencing (KNU NGS) Core Facility, Kyungpook National University","place":["Daegu, Republic of Korea"]},{"name":"Microbalance Inc.","place":["Daegu, Republic of Korea"]}],"role":[{"vocabulary":"crossref","role":"author"}]},{"given":"Yeong-Jun","family":"Park","sequence":"additional","affiliation":[{"name":"Department of Applied Biosciences, Kyungpook National University","place":["Daegu, Republic of Korea"]},{"name":"Kyungpook National University Next-Generation Sequencing (KNU NGS) Core Facility, Kyungpook National University","place":["Daegu, Republic of Korea"]},{"name":"Microbalance Inc.","place":["Daegu, Republic of Korea"]}],"role":[{"vocabulary":"crossref","role":"author"}]},{"given":"Tae-Ho","family":"Kim","sequence":"additional","affiliation":[{"name":"Department of Applied Biosciences, Kyungpook National University","place":["Daegu, Republic of Korea"]}],"role":[{"vocabulary":"crossref","role":"author"}]},{"given":"Yujin","family":"Hyun","sequence":"additional","affiliation":[{"name":"Department of Applied Biosciences, Kyungpook National University","place":["Daegu, Republic of Korea"]}],"role":[{"vocabulary":"crossref","role":"author"}]},{"given":"Sook-Min","family":"Kwon","sequence":"additional","affiliation":[{"name":"Department of Applied Biosciences, Kyungpook National University","place":["Daegu, Republic of Korea"]}],"role":[{"vocabulary":"crossref","role":"author"}]},{"given":"Muhyeok","family":"Kwon","sequence":"additional","affiliation":[{"name":"Department of Applied Biosciences, Kyungpook National University","place":["Daegu, Republic of Korea"]}],"role":[{"vocabulary":"crossref","role":"author"}]},{"given":"Tae-Kyung","family":"Hwang","sequence":"additional","affiliation":[{"name":"Gunwi Agricultural Technology Center","place":["Gunwi-gun, Daegu, Republic of Korea"]}],"role":[{"vocabulary":"crossref","role":"author"}]},{"given":"Jae-Ho","family":"Shin","sequence":"additional","affiliation":[{"name":"Department of Applied Biosciences, Kyungpook National University","place":["Daegu, Republic of Korea"]},{"name":"Kyungpook National University Next-Generation Sequencing (KNU NGS) Core Facility, Kyungpook National University","place":["Daegu, Republic of Korea"]},{"name":"Microbalance Inc.","place":["Daegu, Republic of Korea"]}],"role":[{"vocabulary":"crossref","role":"author"}]}],"member":"1965","published-online":{"date-parts":[[2026,8,31]]},"reference":[{"key":"B1","doi-asserted-by":"publisher","DOI":"10.15156\/BIO\/3301241","article-title":"UNITE QIIME release for fungi. Version 19.02.2025. UNITE community","author":"Abarenkov","year":"2025"},{"key":"B2","doi-asserted-by":"publisher","first-page":"32","DOI":"10.1111\/j.1442-9993.2001.01070.pp.x","article-title":"A new method for non-parametric multivariate analysis of variance","volume":"26","author":"Anderson","year":"2001","journal-title":"Austral Ecol."},{"key":"B3","doi-asserted-by":"publisher","first-page":"557","DOI":"10.1890\/12-2010.1","article-title":"PERMANOVA, ANOSIM, and the Mantel test in the face of heterogeneous dispersions: what null hypothesis are you testing","volume":"83","author":"Anderson","year":"2013","journal-title":"Ecol. Monogr."},{"key":"B4","doi-asserted-by":"publisher","first-page":"307","DOI":"10.1104\/pp.103.028712","article-title":"Biocontrol of Bacillus subtilis against infection of Arabidopsis roots by Pseudomonas syringae is facilitated by biofilm formation and surfactin production","volume":"134","author":"Bais","year":"2004","journal-title":"Plant Physiol."},{"key":"B5","doi-asserted-by":"publisher","first-page":"1089","DOI":"10.1128\/AEM.51.5.1089-1098.1986","article-title":"Alginate beads as synthetic inoculant carriers for slow release of bacteria that affect plant growth","volume":"51","author":"Bashan","year":"1986","journal-title":"Appl. Environ. Microbiol."},{"key":"B6","doi-asserted-by":"publisher","DOI":"10.1136\/bmj.308.6942.1499","article-title":"Regression towards the mean","volume":"308","author":"Bland","year":"1994","journal-title":"BMJ"},{"key":"B7","doi-asserted-by":"publisher","first-page":"127","DOI":"10.1016\/j.tree.2008.10.008","article-title":"Generalized linear mixed models: a practical guide for ecology and evolution","volume":"24","author":"Bolker","year":"2009","journal-title":"Trends Ecol. Evol."},{"key":"B8","doi-asserted-by":"publisher","first-page":"852","DOI":"10.1038\/s41587-019-0209-9","article-title":"Reproducible, interactive, scalable and extensible microbiome data science using QIIME 2","volume":"37","author":"Bolyen","year":"2019","journal-title":"Nat. Biotechnol."},{"key":"B9","doi-asserted-by":"publisher","first-page":"34915","DOI":"10.1021\/acsomega.6c02228","article-title":"Microbial-based agrochemicals: cell-in and cell-free formulations, application, and production via agricultural byproducts for a circular economy","volume":"11","author":"Buttr\u00f3s","year":"2026","journal-title":"ACS Omega"},{"key":"B10","doi-asserted-by":"publisher","first-page":"581","DOI":"10.1038\/nmeth.3869","article-title":"DADA2: high-resolution sample inference from Illumina amplicon data","volume":"13","author":"Callahan","year":"2016","journal-title":"Nat. Methods"},{"key":"B11","doi-asserted-by":"publisher","first-page":"1007","DOI":"10.1038\/nbt1325","article-title":"Comparative analysis of the complete genome sequence of the plant growth-promoting bacterium Bacillus amyloliquefaciens FZB42","volume":"25","author":"Chen","year":"2007","journal-title":"Nat. Biotechnol."},{"key":"B12","doi-asserted-by":"publisher","first-page":"669","DOI":"10.1016\/j.soilbio.2009.11.024","article-title":"Plant growth-promoting bacteria in the rhizo- and endosphere of plants: their role, colonization, mechanisms involved and prospects for utilization","volume":"42","author":"Compant","year":"2010","journal-title":"Soil Biol. Biochem."},{"key":"B13","doi-asserted-by":"publisher","first-page":"137","DOI":"10.1016\/j.postharvbio.2008.11.009","article-title":"Twenty years of postharvest biocontrol research: is it time for a new paradigm","volume":"52","author":"Droby","year":"2009","journal-title":"Postharvest Biol. Technol."},{"key":"B14","doi-asserted-by":"publisher","DOI":"10.3389\/fmicb.2017.00022","article-title":"Bacillus amyloliquefaciens, Bacillus velezensis, and Bacillus siamensis form an \u201coperational group Ba. amyloliquefaciens\u201d within the B. subtilis species complex","volume":"8","author":"Fan","year":"2017","journal-title":"Front. Microbiol."},{"key":"B15","doi-asserted-by":"publisher","first-page":"1684","DOI":"10.1038\/s41467-025-56988-3","article-title":"Rhizobacteria protective hydrogel to promote plant growth and adaption to acidic soil","volume":"16","author":"Feng","year":"2025","journal-title":"Nat. Commun."},{"key":"B16","doi-asserted-by":"publisher","first-page":"29138","DOI":"10.1007\/s11356-019-06023-y","article-title":"Dispersion, persistence, and stability of the biocontrol agent Penicillium frequentans strain 909 after stone fruit tree applications","volume":"26","author":"Guijarro","year":"2019","journal-title":"Environ. Sci. pollut. Res."},{"key":"B17","doi-asserted-by":"publisher","first-page":"411","DOI":"10.1146\/annurev.phyto.40.120401.130158","article-title":"Biological control of postharvest diseases of fruits","volume":"40","author":"Janisiewicz","year":"2002","journal-title":"Annu. Rev. Phytopathol."},{"key":"B18","doi-asserted-by":"publisher","DOI":"10.3390\/plants11212834","article-title":"Changes in microbial community structure in response to gummosis in peach tree bark","volume":"11","author":"Jo","year":"2022","journal-title":"Plants"},{"key":"B19","doi-asserted-by":"publisher","first-page":"211","DOI":"10.3109\/07388551.2010.513327","article-title":"Bio-encapsulation of microbial cells for targeted agricultural delivery","volume":"31","author":"John","year":"2011","journal-title":"Crit. Rev. Biotechnol."},{"key":"B20","doi-asserted-by":"publisher","first-page":"296","DOI":"10.4014\/jmb.2310.10005","article-title":"Biocontrol of peach gummosis by Bacillus velezensis KTA01 and its antifungal mechanism","volume":"34","author":"Kang","year":"2024","journal-title":"J. Microbiol. Biotechnol."},{"key":"B21","doi-asserted-by":"publisher","DOI":"10.1128\/mra.00734-22","article-title":"Complete genome sequence of Bacillus amyloliquefaciens KNU-28 isolated from peach leaves (Prunus persica [L.] Batsch)","volume":"11","author":"Kim","year":"2022","journal-title":"Microbiol. Resour. Announc"},{"key":"B22","doi-asserted-by":"publisher","DOI":"10.3389\/fpls.2019.00845","article-title":"Mode of action of microbial biological control agents against plant diseases: relevance beyond efficacy","volume":"10","author":"K\u00f6hl","year":"2019","journal-title":"Front. Plant Sci."},{"key":"B23","doi-asserted-by":"publisher","DOI":"10.3390\/gels11040302","article-title":"Encapsulation of Bacillus subtilis in chitosan gel beads for eco-friendly crop protection","volume":"11","author":"Krastev","year":"2025","journal-title":"Gels"},{"key":"B24","doi-asserted-by":"publisher","DOI":"10.3390\/jof9010057","article-title":"Antifungal activity and biocontrol potential of Simplicillium lamellicola JC-1 against multiple fungal pathogens of oilseed rape","volume":"9","author":"Li","year":"2023","journal-title":"J. Fungi"},{"key":"B25","doi-asserted-by":"publisher","DOI":"10.1371\/journal.pone.0162125","article-title":"Antifungal activity of isolated Bacillus amyloliquefaciens SYBC H47 for the biocontrol of peach gummosis","volume":"11","author":"Li","year":"2016","journal-title":"PloS One"},{"key":"B26","doi-asserted-by":"publisher","first-page":"190","DOI":"10.1016\/j.ijfoodmicro.2010.06.023","article-title":"Gamma-polyglutamic acid (\u03b3-PGA) produced by Bacillus amyloliquefaciens C06 promoting its colonization on fruit surface","volume":"142","author":"Liu","year":"2010","journal-title":"Int. J. Food Microbiol."},{"key":"B27","doi-asserted-by":"publisher","first-page":"391","DOI":"10.1007\/s00284-001-0011-y","article-title":"Secretion of \u03b2-1,3-glucanases by the yeast Pichia membranifaciens and its possible role in the biocontrol of Botrytis cinerea causing grey mold disease of the grapevine","volume":"44","author":"Masih","year":"2002","journal-title":"Curr. Microbiol."},{"key":"B28","doi-asserted-by":"publisher","first-page":"98","DOI":"10.1016\/j.biocontrol.2015.06.003","article-title":"Biological control in the microbiome era: challenges and opportunities","volume":"89","author":"Massart","year":"2015","journal-title":"Biol. Control"},{"key":"B29","doi-asserted-by":"publisher","DOI":"10.1371\/journal.pcbi.1003531","article-title":"Waste not, want not: why rarefying microbiome data is inadmissible","volume":"10","author":"McMurdie","year":"2014","journal-title":"PloS Comput. Biol."},{"key":"B30","doi-asserted-by":"publisher","first-page":"566","DOI":"10.1094\/Phyto-81-566","article-title":"Pathogenicity, distribution, sources of inoculum, and infection courts of Botryosphaeria dothidea on pistachio","volume":"81","author":"Michailides","year":"1991","journal-title":"Phytopathology"},{"key":"B31","doi-asserted-by":"publisher","first-page":"D259","DOI":"10.1093\/nar\/gky1022","article-title":"The UNITE database for molecular identification of fungi: handling dark taxa and parallel taxonomic classifications","volume":"47","author":"Nilsson","year":"2019","journal-title":"Nucleic Acids Res."},{"key":"B32","doi-asserted-by":"publisher","first-page":"199","DOI":"10.1007\/s10658-011-9919-7","article-title":"Biological control of postharvest diseases of fruit","volume":"133","author":"Nunes","year":"2012","journal-title":"Eur. J. Plant Pathol."},{"key":"B33","doi-asserted-by":"publisher","first-page":"1","DOI":"10.1099\/mic.0.081448-0","article-title":"Poly-\u03b3-glutamic acid: production, properties and applications","volume":"161","author":"Ogunleye","year":"2015","journal-title":"Microbiology"},{"key":"B34","doi-asserted-by":"publisher","first-page":"115","DOI":"10.1016\/j.tim.2007.12.009","article-title":"Bacillus lipopeptides: versatile weapons for plant disease biocontrol","volume":"16","author":"Ongena","year":"2008","journal-title":"Trends Microbiol."},{"key":"B35","doi-asserted-by":"publisher","first-page":"1084","DOI":"10.1111\/j.1462-2920.2006.01202.x","article-title":"Surfactin and fengycin lipopeptides of Bacillus subtilis as elicitors of induced systemic resistance in plants","volume":"9","author":"Ongena","year":"2007","journal-title":"Environ. Microbiol."},{"key":"B36","doi-asserted-by":"publisher","DOI":"10.1371\/journal.pone.0097629","article-title":"Comparison and validation of some ITS primer pairs useful for fungal metabarcoding studies","volume":"9","author":"Op De Beeck","year":"2014","journal-title":"PloS One"},{"key":"B37","doi-asserted-by":"publisher","first-page":"187","DOI":"10.1016\/j.copbio.2010.12.003","article-title":"Plant protection and growth stimulation by microorganisms: biotechnological applications of Bacilli in agriculture","volume":"22","author":"P\u00e9rez-Garc\u00eda","year":"2011","journal-title":"Curr. Opin. Biotechnol."},{"key":"B38","doi-asserted-by":"publisher","first-page":"789","DOI":"10.1038\/nrmicro3109","article-title":"Going back to the roots: the microbial ecology of the rhizosphere","volume":"11","author":"Philippot","year":"2013","journal-title":"Nat. Rev. Microbiol."},{"key":"B39","doi-asserted-by":"publisher","first-page":"51","DOI":"10.3114\/sim0021","article-title":"The Botryosphaeriaceae: genera and species known from culture","volume":"76","author":"Phillips","year":"2013","journal-title":"Stud. Mycol."},{"key":"B40","doi-asserted-by":"publisher","DOI":"10.3389\/fmicb.2018.03047","article-title":"Understand the potential role of Aureobasidium pullulans, a resident microorganism from grapevine, to prevent the infection caused by Diplodia seriata","volume":"9","author":"Pinto","year":"2018","journal-title":"Front. Microbiol."},{"key":"B41","doi-asserted-by":"publisher","DOI":"10.3390\/microorganisms7090322","article-title":"Organs, cultivars, soil, and fruit properties affect structure of endophytic mycobiota of Pinggu peach trees","volume":"7","author":"Ren","year":"2019","journal-title":"Microorganisms"},{"key":"B42","doi-asserted-by":"publisher","first-page":"430","DOI":"10.1094\/MPMI-20-4-0430","article-title":"The iturin and fengycin families of lipopeptides are key factors in antagonism of Bacillus subtilis toward Podosphaera fusca","volume":"20","author":"Romero","year":"2007","journal-title":"Mol. Plant-Microbe Interact."},{"key":"B43","doi-asserted-by":"publisher","first-page":"7338","DOI":"10.1038\/s41598-026-38691-5","article-title":"Biopolymer-based multilayer capsules for protection and controlled release of Pseudomonas fluorescens T17-4 and Bacillus velezensis VRU1","volume":"16","author":"Saberi Riseh","year":"2026","journal-title":"Sci. Rep."},{"key":"B44","doi-asserted-by":"publisher","DOI":"10.3390\/ijms222011165","article-title":"Encapsulation of plant biocontrol bacteria with alginate as a main polymer material","volume":"22","author":"Saberi Riseh","year":"2021","journal-title":"Int. J. Mol. Sci."},{"key":"B45","doi-asserted-by":"publisher","first-page":"6241","DOI":"10.1073\/pnas.1117018109","article-title":"Nuclear ribosomal internal transcribed spacer (ITS) region as a universal DNA barcode marker for Fungi","volume":"109","author":"Schoch","year":"2012","journal-title":"Proc. Natl. Acad. Sci. U.S.A."},{"key":"B46","doi-asserted-by":"publisher","DOI":"10.1186\/gb-2011-12-6-r60","article-title":"Metagenomic biomarker discovery and explanation","volume":"12","author":"Segata","year":"2011","journal-title":"Genome Biol."},{"key":"B47","doi-asserted-by":"publisher","first-page":"337","DOI":"10.5423\/PPJ.FT.04.2017.0087","article-title":"Development of a biofungicide using a mycoparasitic fungus Simplicillium lamellicola BCP and its control efficacy against gray mold diseases of tomato and ginseng","volume":"33","author":"Shin","year":"2017","journal-title":"Plant Pathol. J."},{"key":"B48","doi-asserted-by":"publisher","first-page":"90","DOI":"10.1016\/j.fbr.2007.06.002","article-title":"Botryosphaeriaceae as endophytes and latent pathogens of woody plants: diversity, ecology and impact","volume":"21","author":"Slippers","year":"2007","journal-title":"Fungal Biol. Rev."},{"key":"B49","doi-asserted-by":"publisher","first-page":"128","DOI":"10.1111\/j.1365-3059.2006.01486.x","article-title":"Taxonomy, phylogeny and identification of Botryosphaeriaceae associated with pome and stone fruit trees in South Africa and other regions of the world","volume":"56","author":"Slippers","year":"2007","journal-title":"Plant Pathol."},{"key":"B50","doi-asserted-by":"publisher","first-page":"2769","DOI":"10.1111\/mec.16460","article-title":"Best practices in metabarcoding of fungi: from experimental design to results","volume":"31","author":"Tedersoo","year":"2022","journal-title":"Mol. Ecol."},{"key":"B51","doi-asserted-by":"publisher","first-page":"1","DOI":"10.1146\/annurev-ecolsys-121415-032238","article-title":"The phyllosphere: microbial jungle at the plant\u2013climate interface","volume":"47","author":"Vacher","year":"2016","journal-title":"Annu. Rev. Ecol. Evol. Syst."},{"key":"B52","doi-asserted-by":"publisher","first-page":"157","DOI":"10.1038\/nrmicro2960","article-title":"Sticking together: building a biofilm the Bacillus subtilis way","volume":"11","author":"Vlamakis","year":"2013","journal-title":"Nat. Rev. Microbiol."},{"key":"B53","doi-asserted-by":"publisher","first-page":"1378","DOI":"10.1094\/PDIS-12-10-0893","article-title":"Identification and characterization of Botryosphaeria spp. causing gummosis of peach trees in Hubei Province, central China","volume":"95","author":"Wang","year":"2011","journal-title":"Plant Dis."},{"key":"B54","doi-asserted-by":"publisher","first-page":"1429","DOI":"10.1094\/Phyto-64-1429","article-title":"A gummosis disease of peach trees caused by Botryosphaeria dothidea","volume":"64","author":"Weaver","year":"1974","journal-title":"Phytopathology"},{"key":"B55","first-page":"315","article-title":"Amplification and direct sequencing of fungal ribosomal RNA genes for phylogenetics","volume-title":"PCR Protocols: A Guide to Methods and Applications","author":"White","year":"1990"},{"key":"B56","doi-asserted-by":"publisher","DOI":"10.3389\/fmicb.2016.01811","article-title":"Poly-\u03b3-glutamic acids contribute to biofilm formation and plant root colonization in selected environmental isolates of Bacillus subtilis","volume":"7","author":"Yu","year":"2016","journal-title":"Front. Microbiol."}],"container-title":["Frontiers in Plant Science"],"original-title":[],"link":[{"URL":"https:\/\/www.frontiersin.org\/articles\/10.3389\/fpls.2026.1933566\/full","content-type":"unspecified","content-version":"vor","intended-application":"similarity-checking"}],"deposited":{"date-parts":[[2026,8,31]],"date-time":"2026-08-31T05:39:16Z","timestamp":1788154756000},"score":1,"resource":{"primary":{"URL":"https:\/\/www.frontiersin.org\/articles\/10.3389\/fpls.2026.1933566\/full"}},"subtitle":[],"short-title":[],"issued":{"date-parts":[[2026,8,31]]},"references-count":56,"alternative-id":["10.3389\/fpls.2026.1933566"],"URL":"https:\/\/doi.org\/10.3389\/fpls.2026.1933566","relation":{},"ISSN":["1664-462X"],"issn-type":[{"value":"1664-462X","type":"electronic"}],"subject":[],"published":{"date-parts":[[2026,8,31]]},"article-number":"1933566"}}