{"status":"ok","message-type":"work","message-version":"1.0.0","message":{"indexed":{"date-parts":[[2026,1,28]],"date-time":"2026-01-28T10:38:25Z","timestamp":1769596705014,"version":"3.49.0"},"reference-count":154,"publisher":"Frontiers Media SA","license":[{"start":{"date-parts":[[2021,8,4]],"date-time":"2021-08-04T00:00:00Z","timestamp":1628035200000},"content-version":"vor","delay-in-days":0,"URL":"https:\/\/creativecommons.org\/licenses\/by\/4.0\/"}],"content-domain":{"domain":["frontiersin.org"],"crossmark-restriction":true},"short-container-title":["Front. Psychol."],"abstract":"<jats:p>Human walking is a socially embedded and shaped biological adaptation: it frees our hands, makes our minds mobile, and is deeply health promoting. Yet, today, physical inactivity is an unsolved, major public health problem. However, globally, tens of millions of people annually undertake ancient, significant and enduring traditions of physiologically and psychologically arduous walks (pilgrimages) of days-to-weeks extent. Pilgrim walking is a significant human activity requiring weighty commitments of time, action and belief, as well as community support. Paradoxically, human walking is most studied on treadmills, not \u2018<jats:italic>in the wild\u2019<\/jats:italic>, while mechanistically vital, treadmill studies of walking cannot, in principle, address why humans walk extraordinary distances together to demonstrate their adherence to a behaviourally demanding belief system.<\/jats:p><jats:p>Pilgrim walkers provide a rich \u2018living laboratory\u2019 bridging humanistic inquiries, to progressive theoretical and empirical investigations of human walking arising from a behaviourally demanding belief system. Pilgrims vary demographically and undertake arduous journeys on precisely mapped routes of tracked, titrated doses and durations on terrain of varying difficulty, allowing investigations from molecular to cultural levels of analysis. Using the reciprocal perspectives of \u2018<jats:italic>inside\u2192out<\/jats:italic>\u2019 (where processes within brain and body initiate, support and entrain movement) and \u2018<jats:italic>outside\u2192in<\/jats:italic>\u2019 (where processes in the world beyond brain and body drive activity<jats:italic>within<\/jats:italic>brain and body), we examine how pilgrim walking might shape personal, social and transcendental processes, revealing potential mechanisms supporting the body and brain in motion, to how pilgrim walking might offer policy solutions for physical inactivity.<\/jats:p>","DOI":"10.3389\/fpsyg.2021.654122","type":"journal-article","created":{"date-parts":[[2021,8,4]],"date-time":"2021-08-04T04:28:51Z","timestamp":1628051331000},"update-policy":"https:\/\/doi.org\/10.3389\/crossmark-policy","source":"Crossref","is-referenced-by-count":11,"title":["Biopsychosocial Functions of Human Walking and Adherence to Behaviourally Demanding Belief Systems: A Narrative Review"],"prefix":"10.3389","volume":"12","author":[{"given":"Shane","family":"O\u2019Mara","sequence":"first","affiliation":[],"role":[{"role":"author","vocabulary":"crossref"}]}],"member":"1965","published-online":{"date-parts":[[2021,8,4]]},"reference":[{"key":"ref1","doi-asserted-by":"publisher","first-page":"29","DOI":"10.7554\/eLife.19568","article-title":"New footprints from laetoli (Tanzania) provide evidence for marked body size variation in early hominins","volume":"5","author":"Addis","year":"2017","journal-title":"Nature"},{"key":"ref502","doi-asserted-by":"publisher","first-page":"527","DOI":"10.1037\/0033-295X.115.3.527","article-title":"What is the shape of developmental change?","volume":"115","author":"Adolph","year":"2008","journal-title":"Psychol. Rev."},{"key":"ref2","doi-asserted-by":"publisher","first-page":"2292","DOI":"10.1111\/j.1460-9568.2010.07251.x","article-title":"Hippocampal\u2013anterior thalamic pathways for memory: uncovering a network of direct and indirect actions","volume":"31","author":"Aggleton","year":"2010","journal-title":"Eur. J. Neurosci."},{"key":"ref3","doi-asserted-by":"publisher","first-page":"3743","DOI":"10.1016\/j.cub.2017.10.062","article-title":"Theta oscillations in the human medial temporal lobe during real-world ambulatory movement","volume":"27","author":"Aghajan","year":"2017","journal-title":"Curr. Biol."},{"key":"ref4","doi-asserted-by":"publisher","first-page":"336","DOI":"10.1038\/nature23018","article-title":"Large-scale physical activity data reveal worldwide activity inequality","volume":"547","author":"Althoff","year":"2017","journal-title":"Nature"},{"key":"ref5","doi-asserted-by":"publisher","first-page":"271","DOI":"10.1016\/j.tourman.2017.09.007","article-title":"A closer look at Santiago de Compostela\u2019s pilgrims through the lens of motivations","volume":"64","author":"Amaro","year":"2018","journal-title":"Tour. Manag."},{"key":"ref501","year":"2015"},{"key":"ref6","doi-asserted-by":"publisher","first-page":"236","DOI":"10.1016\/j.wem.2011.03.005","article-title":"Physiological adaptation of a mature adult walking the Alps","volume":"22","author":"Ardigo","year":"2011","journal-title":"Wilderness Environ. Med."},{"key":"ref7","doi-asserted-by":"publisher","first-page":"3712","DOI":"10.1002\/hbm.24627","article-title":"Neural processing of social interaction: coordinate-based meta-analytic evidence from human neuroimaging studies","volume":"40","author":"Arioli","year":"2019","journal-title":"Hum. Brain Mapp."},{"key":"ref8","doi-asserted-by":"publisher","first-page":"437","DOI":"10.1016\/j.tics.2012.06.010","article-title":"Top-down versus bottom-up attentional control: a failed theoretical dichotomy","volume":"16","author":"Awh","year":"2012","journal-title":"Trends Cogn. Sci."},{"key":"ref9","doi-asserted-by":"publisher","DOI":"10.1126\/science.aai9067","article-title":"On the origin of modern humans: Asian perspectives","volume":"358","author":"Bae","year":"2017","journal-title":"Science"},{"key":"ref10","doi-asserted-by":"publisher","first-page":"127","DOI":"10.3233\/BPL-160040","article-title":"The effects of acute exercise on mood, cognition, neurophysiology, and neurochemical pathways: a review","volume":"2","author":"Basso","year":"2017","journal-title":"Brain Plast"},{"key":"ref11","first-page":"279","article-title":"59 Inner Processes Serve Interpersonal Functions,","volume-title":"Scientists Making a Difference: One Hundred Eminent Behavioral and Brain Scientists Talk about their Most Important Contributions","author":"Baumeister","year":"2016"},{"key":"ref12","doi-asserted-by":"publisher","first-page":"497","DOI":"10.1037\/0033-2909.117.3.497","article-title":"The need to belong: desire for interpersonal attachments as a fundamental human motivation","volume":"117","author":"Baumeister","year":"1995","journal-title":"Psychol. Bull."},{"key":"ref13","doi-asserted-by":"publisher","first-page":"87","DOI":"10.1016\/j.tics.2015.10.004","article-title":"Creative cognition and brain network dynamics","volume":"20","author":"Beaty","year":"2016","journal-title":"Trends Cogn. Sci."},{"key":"ref14","doi-asserted-by":"publisher","DOI":"10.1136\/bmjopen-2012-000875","article-title":"The relationship between walking speed and changes in cardiovascular risk factors during a 12-day walking tour to Santiago de Compostela: a cohort study","volume":"2","author":"Bemelmans","year":"2012","journal-title":"BMJ Open"},{"key":"ref15","doi-asserted-by":"publisher","first-page":"621","DOI":"10.1016\/j.atherosclerosis.2010.06.012","article-title":"Vascular and metabolic effects of 12 days intensive walking to Santiago de Compostela","volume":"212","author":"Bemelmans","year":"2010","journal-title":"Atherosclerosis"},{"key":"ref16","doi-asserted-by":"publisher","DOI":"10.1136\/bmjopen-2016-013384","article-title":"Social prescribing: less rhetoric and more reality. A systematic review of the evidence","volume":"7","author":"Bickerdike","year":"2017","journal-title":"BMJ Open"},{"key":"ref17","doi-asserted-by":"publisher","first-page":"123","DOI":"10.7326\/M14-1651","article-title":"Sedentary time and its association with risk for disease incidence, mortality, and hospitalization in adults: a systematic review and meta-analysis","volume":"162","author":"Biswas","year":"2015","journal-title":"Ann. Intern. Med."},{"key":"ref18","doi-asserted-by":"publisher","first-page":"51","DOI":"10.3389\/fncom.2013.00051","article-title":"The neural origin of muscle synergies","volume":"7","author":"Bizzi","year":"2013","journal-title":"Front. Comput. Neurosci."},{"key":"ref19","doi-asserted-by":"publisher","first-page":"1143","DOI":"10.1002\/cphy.c110025","article-title":"Lack of exercise is a major cause of chronic diseases","volume":"2","author":"Booth","year":"2011","journal-title":"Compr. Physiol."},{"key":"ref20","doi-asserted-by":"publisher","first-page":"aax0903","DOI":"10.1126\/sciadv.aax0903","article-title":"Nature and mental health: an ecosystem service perspective","volume":"5","author":"Bratman","year":"2019","journal-title":"Sci. Adv."},{"key":"ref21","doi-asserted-by":"publisher","first-page":"8567","DOI":"10.1073\/pnas.1510459112","article-title":"Nature experience reduces rumination and subgenual prefrontal cortex activation","volume":"112","author":"Bratman","year":"2015","journal-title":"Proc. Natl. Acad. Sci."},{"key":"ref45","year":""},{"key":"ref22","doi-asserted-by":"publisher","first-page":"27","DOI":"10.1146\/annurev.psych.60.110707.163508","article-title":"The role of the hippocampus in prediction and imagination","volume":"61","author":"Buckner","year":"2010","journal-title":"Annu. Rev. Psychol."},{"key":"ref23","doi-asserted-by":"publisher","first-page":"769349","DOI":"10.1101\/769349","article-title":"Differential effects of opioid receptor modulators on motivational and stress-coping behaviors in the back-translational rat IFN-\u03b1 depression model","author":"Callaghan","year":"2019","journal-title":"bioRxiv"},{"key":"ref24","doi-asserted-by":"publisher","first-page":"377","DOI":"10.1016\/j.physbeh.2017.07.018","article-title":"Exercise prevents IFN-\u03b1-induced mood and cognitive dysfunction and increases BDNF expression in the rat","volume":"179","author":"Callaghan","year":"2017","journal-title":"Physiol. Behav."},{"key":"ref25","doi-asserted-by":"publisher","first-page":"89","DOI":"10.1016\/bs.pbr.2018.07.009","volume":"239","author":"Callaghan","year":"2018","journal-title":"Prog Brain Res."},{"key":"ref26","doi-asserted-by":"publisher","DOI":"10.1371\/journal.pone.0193506","article-title":"Motivational valence alters memory formation without altering exploration of a real-life spatial environment","volume":"13","author":"Chiew","year":"2018","journal-title":"PLoS One"},{"key":"ref27","doi-asserted-by":"publisher","first-page":"1235","DOI":"10.3390\/ijerph6031235","article-title":"How the mid-Victorians worked, ate and died","volume":"6","author":"Clayton","year":"2009","journal-title":"Int. J. Environ. Res. Public Health"},{"key":"ref28","doi-asserted-by":"publisher","first-page":"1112","DOI":"10.1016\/j.cell.2020.04.043","article-title":"Molecular choreography of acute exercise","volume":"181","author":"Contrepois","year":"2020","journal-title":"Cell"},{"key":"ref29","doi-asserted-by":"publisher","first-page":"170","DOI":"10.1080\/14427591.2013.764816","article-title":"A review of the literature exploring the occupation of contemporary pilgrimage","volume":"22","author":"Courtney","year":"2015","journal-title":"J. Occup. Sci."},{"key":"ref30","doi-asserted-by":"publisher","first-page":"528","DOI":"10.1016\/j.gaitpost.2013.01.027","article-title":"Treadmill versus overground and barefoot versus shod comparisons of triceps surae fascicle behaviour in human walking and running","volume":"38","author":"Cronin","year":"2013","journal-title":"Gait Posture"},{"key":"ref31","doi-asserted-by":"crossref","first-page":"209","DOI":"10.1007\/978-94-017-9088-8_14","article-title":"Toward a psychology of optimal experience","volume-title":"Flow and the Foundations of Positive Psychology","author":"Csikszentmihalyi","year":"2014"},{"key":"ref32","doi-asserted-by":"publisher","first-page":"344","DOI":"10.1046\/j.1460-9568.1999.00446.x","article-title":"Sustained activation of hippocampal pyramidal cells by \u2018space clamping\u2019 in a running wheel","volume":"11","author":"Czurk\u00f3","year":"1999","journal-title":"Eur. J. Neurosci."},{"key":"ref33","doi-asserted-by":"publisher","first-page":"312","DOI":"10.1016\/j.neuron.2013.09.007","article-title":"Goals and habits in the brain","volume":"80","author":"Dolan","year":"2013","journal-title":"Neuron"},{"key":"ref34","author":"Donoghue","year":"2016"},{"key":"ref35","first-page":"26","article-title":"When forest-based hunter-gatherers become sedentary: consequences for diet and health","volume":"57","author":"Dounias","year":"2006","journal-title":"UNASYLVA-FAO"},{"key":"ref36","doi-asserted-by":"publisher","first-page":"S294","DOI":"10.1177\/15648265070282S208","article-title":"From sago to rice, from forest to town: the consequences of sedentarization for the nutritional ecology of Punan former hunter-gatherers of Borneo","volume":"28","author":"Dounias","year":"2007","journal-title":"Food Nutr. Bull."},{"key":"ref37","doi-asserted-by":"crossref","first-page":"41","DOI":"10.1016\/B978-0-12-816660-4.00003-9","article-title":"The neurophysiological basis of optimism bias","volume-title":"Cognitive Biases in Health and Psychiatric Disorders.","author":"Dricu","year":"2020"},{"key":"ref38","doi-asserted-by":"publisher","first-page":"780","DOI":"10.1016\/j.tics.2018.06.004","article-title":"Mere membership","volume":"22","author":"Dunham","year":"2018","journal-title":"Trends Cogn. Sci."},{"key":"ref39","doi-asserted-by":"publisher","first-page":"107","DOI":"10.3167\/JYS.2010.110106","article-title":"Walking back to happiness? modern pilgrimage and the expression of suffering on Spain\u2019s Camino de Santiago","volume":"11","author":"Egan","year":"2010","journal-title":"Journeys"},{"key":"ref40","doi-asserted-by":"publisher","first-page":"999","DOI":"10.1016\/j.neubiorev.2010.08.002","article-title":"Threat detection: behavioral practices in animals and humans","volume":"35","author":"Eilam","year":"2011","journal-title":"Neurosci. Biobehav. Rev."},{"key":"ref41","doi-asserted-by":"publisher","first-page":"112","DOI":"10.1017\/S1355617708090152","article-title":"Walking and talking therapy: improving cognitive-motor dual-tasking in neurological illness","volume":"15","author":"Evans","year":"2009","journal-title":"J. Int. Neuropsychol. Soc."},{"key":"ref42","doi-asserted-by":"publisher","first-page":"1132","DOI":"10.1016\/j.jvs.2012.04.046","article-title":"Supervised walking therapy in patients with intermittent claudication","volume":"56","author":"Fakhry","year":"2012","journal-title":"J. Vasc. Surg."},{"key":"ref43","doi-asserted-by":"publisher","first-page":"28","DOI":"10.1093\/socrel\/sry019","article-title":"Atheists on the Santiago way: examining motivations to go on pilgrimage","volume":"80","author":"Farias","year":"2019","journal-title":"Sociol. Relig."},{"key":"ref44","doi-asserted-by":"publisher","first-page":"174","DOI":"10.1016\/j.conb.2016.09.013","article-title":"Cortical circuits and modules in movement generation: experiments and theories","volume":"41","author":"Flash","year":"2016","journal-title":"Curr. Opin. Neurobiol."},{"key":"ref46","doi-asserted-by":"publisher","first-page":"142","DOI":"10.1097\/NPT.0000000000000090","article-title":"Motor-cognitive dual-task training in neurologic disorders: a systematic review","volume":"39","author":"Fritz","year":"2015","journal-title":"J. Neurol. Phys. Ther."},{"key":"ref47","doi-asserted-by":"publisher","first-page":"1349","DOI":"10.1037\/pas0000434","article-title":"The psychological importance of collective assembly: development and validation of the tendency for effervescent assembly measure (TEAM)","volume":"29","author":"Gabriel","year":"2017","journal-title":"Psychol. Assess."},{"key":"ref48","doi-asserted-by":"publisher","first-page":"15","DOI":"10.1080\/14729679.2017.1321999","article-title":"A walk down memory lane: on the relationship between autobiographical memories and outdoor activities","volume":"18","author":"Gibson","year":"2018","journal-title":"J. Adventure Educ. Outdoor Learn."},{"key":"ref49","doi-asserted-by":"publisher","first-page":"617","DOI":"10.1037\/0022-3514.75.3.617","article-title":"Immune neglect: a source of durability bias in affective forecasting","volume":"75","author":"Gilbert","year":"1998","journal-title":"J. Pers. Soc. Psychol."},{"key":"ref50","volume-title":"Pilgrimage: From the Ganges to Graceland: An Encyclopedia.","author":"Gitlitz","year":"2002"},{"key":"ref51","doi-asserted-by":"crossref","first-page":"124","DOI":"10.1787\/2f7cd1bf-en","volume-title":"The Heavy Burden of Obesity: The Economics of Prevention.","author":"Goryakin","year":"2019"},{"key":"ref52","doi-asserted-by":"publisher","DOI":"10.1016\/j.physbeh.2011.06.005","article-title":"Aerobic exercise improves hippocampal function and increases BDNF in the serum of young adult males","volume":"104","author":"Griffin","year":"2011","journal-title":"Physiol. Behav."},{"key":"ref53","doi-asserted-by":"publisher","DOI":"10.21427\/D7242Z","article-title":"The importance of religious tourism and pilgrimage: reflecting on definitions, motives and data","volume":"5","author":"Griffin","year":"2017","journal-title":"Int. J. Relig. Tour. Pilgr."},{"key":"ref54","year":"2019"},{"key":"ref55","doi-asserted-by":"publisher","first-page":"1","DOI":"10.1016\/0022-5193(64)90038-4","article-title":"The genetical evolution of social behaviour. I","volume":"7","author":"Hamilton","year":"1964","journal-title":"J. Theor. Biol."},{"key":"ref56","doi-asserted-by":"publisher","DOI":"10.21427\/Q6DE-AV43","article-title":"The physiological effects of walking pilgrimage","volume":"7","author":"Harris","year":"2019","journal-title":"Int. J. Relig. Tour. Pilgr."},{"key":"ref57","doi-asserted-by":"publisher","first-page":"727","DOI":"10.1055\/s-0032-1331172","article-title":"Cardiovascular disease risk following a 758 km pilgrimage","volume":"34","author":"Harris","year":"2013","journal-title":"Int. J. Sports Med."},{"key":"ref58","doi-asserted-by":"publisher","first-page":"28","DOI":"10.1176\/appi.ajp.2017.16111223","article-title":"Exercise and the prevention of depression: results of the HUNT cohort study","volume":"175","author":"Harvey","year":"2018","journal-title":"Am. J. Psychiatry"},{"key":"ref59","doi-asserted-by":"publisher","first-page":"299","DOI":"10.1016\/j.tics.2007.05.001","article-title":"Deconstructing episodic memory with construction","volume":"11","author":"Hassabis","year":"2007","journal-title":"Trends Cogn. Sci."},{"key":"ref504","doi-asserted-by":"publisher","first-page":"32","DOI":"10.1016\/j.neulet.2019.02.009","article-title":"Effects of self-locomotion on the activity of place cells in the hippocampus of a freely behaving monkey","volume":"701","author":"Hazama","year":"2019","journal-title":"Neurosci. Lett."},{"key":"ref60","doi-asserted-by":"publisher","first-page":"456","DOI":"10.1126\/science.aap8369","article-title":"The earliest modern humans outside Africa","volume":"359","author":"Hershkovitz","year":"2018","journal-title":"Science"},{"key":"ref61","article-title":"The nineteenth century London labour market","volume-title":"London: Aspects of Change.","author":"Hobsbawn","year":"1964"},{"key":"ref62","volume-title":"Mothers and Others: The Evolutionary Origins of Mutual Understanding.","author":"Hrdy","year":"2009"},{"key":"ref63","doi-asserted-by":"crossref","first-page":"65","DOI":"10.4324\/9780203789445-4","article-title":"Comes the child before man: how cooperative breeding and prolonged postweaning dependence shaped human potential","volume-title":"Hunter-Gatherer Childhoods","author":"Hrdy","year":"2017"},{"key":"ref64","doi-asserted-by":"publisher","first-page":"309","DOI":"10.1111\/hsc.12839","article-title":"What approaches to social prescribing work, for whom, and in what circumstances? A realist review","volume":"28","author":"Husk","year":"2020","journal-title":"Health Soc. Care Community"},{"key":"ref65","doi-asserted-by":"publisher","first-page":"236","DOI":"10.1016\/j.cortex.2010.01.002","article-title":"Impaired capacity for autonoetic reliving during autobiographical event recall in mild Alzheimer\u2019s disease","volume":"47","author":"Irish","year":"2011","journal-title":"Cortex"},{"key":"ref66","doi-asserted-by":"publisher","first-page":"27","DOI":"10.3389\/fnbeh.2013.00027","volume":"7","author":"Irish","year":"2013","journal-title":"The pivotal role of semantic memory in remembering the past and imagining the future.Front. Behav. Neurosci."},{"key":"ref67","doi-asserted-by":"publisher","first-page":"1","DOI":"10.14336\/AD.2014.1210","article-title":"The critical need to promote research of aging and aging-related diseases to improve health and longevity of the elderly population","volume":"6","author":"Jin","year":"2015","journal-title":"Aging Dis."},{"key":"ref68","doi-asserted-by":"publisher","DOI":"10.21427\/V0CC-7135","article-title":"The processes, effects and therapeutics of pilgrimage walking the St. Olav way","volume":"8","author":"J\u00f8rgensen","year":"2020","journal-title":"Int. J. Relig. Tour. Pilgr."},{"key":"ref69","doi-asserted-by":"publisher","first-page":"3","DOI":"10.1177\/0013916587191001","article-title":"Aesthetics, affect, and cognition: environmental preference from an evolutionary perspective","volume":"19","author":"Kaplan","year":"1987","journal-title":"Environ. Behav."},{"key":"ref70","doi-asserted-by":"publisher","first-page":"1730","DOI":"10.1016\/S0140-6736(17)30752-3","article-title":"Coronary atherosclerosis in indigenous south American Tsimane: a cross-sectional cohort study","volume":"389","author":"Kaplan","year":"2017","journal-title":"Lancet"},{"key":"ref71","doi-asserted-by":"publisher","first-page":"593","DOI":"10.1007\/s10734-015-9926-2","article-title":"Taking your mind for a walk: a qualitative investigation of walking and thinking among nine Norwegian academics","volume":"71","author":"Kein\u00e4nen","year":"2016","journal-title":"High. Educ."},{"key":"ref505","doi-asserted-by":"publisher","first-page":"214","DOI":"10.1002\/dneu.22465","author":"Kenkel","year":"2017","journal-title":"Dev. Neurobiol."},{"key":"ref72","doi-asserted-by":"publisher","first-page":"297","DOI":"10.1080\/02699930302297","article-title":"Approaching awe, a moral, spiritual, and aesthetic emotion","volume":"17","author":"Keltner","year":"2003","journal-title":"Cogn. Emot."},{"key":"ref73","doi-asserted-by":"publisher","first-page":"231","DOI":"10.1038\/sj.jea.7500165","article-title":"The National Human Activity Pattern Survey (NHAPS): a resource for assessing exposure to environmental pollutants","volume":"11","author":"Klepeis","year":"2001","journal-title":"J. Expo. Sci. Environ. Epidemiol."},{"key":"ref74","doi-asserted-by":"publisher","first-page":"910","DOI":"10.1177\/1948550614538463","article-title":"Get up, stand up: the effects of a non-sedentary workspace on information elaboration and group performance","volume":"5","author":"Knight","year":"2014","journal-title":"Soc. Psychol. Personal. Sci."},{"key":"ref75","doi-asserted-by":"publisher","first-page":"1580","DOI":"10.3389\/fpsyg.2016.01580","article-title":"Sensorimotor-conceptual integration in free walking enhances divergent thinking for young and older adults","volume":"7","author":"Kuo","year":"2016","journal-title":"Front. Psychol."},{"key":"ref76","doi-asserted-by":"publisher","first-page":"1","DOI":"10.1016\/j.chb.2014.12.054","article-title":"The delayed effect of treadmill desk usage on recall and attention","volume":"46","author":"Labont\u00e9-LeMoyne","year":"2015","journal-title":"Comput. Human Behav."},{"key":"ref77","doi-asserted-by":"publisher","first-page":"723","DOI":"10.3389\/FPSYG.2015.00723","article-title":"Slow walking on a treadmill desk does not negatively affect executive abilities: an examination of cognitive control, conflict adaptation, response inhibition, and post-error slowing","volume":"6","author":"Larson","year":"","journal-title":"Front. Psychol."},{"key":"ref78","doi-asserted-by":"publisher","first-page":"e0121309","DOI":"10.1371\/JOURNAL.PONE.0121309","article-title":"Cognitive and typing outcomes measured simultaneously with slow treadmill walking or sitting: implications for treadmill desks","volume":"10","author":"Larson","year":"","journal-title":"PLoS One"},{"key":"ref79","doi-asserted-by":"publisher","first-page":"141","DOI":"10.1186\/s12877-016-0315-1","article-title":"Influence of combined physical and cognitive training on cognition: a systematic review","volume":"16","author":"Lauenroth","year":"2016","journal-title":"BMC Geriatr."},{"key":"ref80","doi-asserted-by":"publisher","first-page":"558","DOI":"10.1136\/bjsm.2006.032755","article-title":"The energy expenditure of using a \u201cwalk-and-work\u201d desk for office workers with obesity","volume":"41","author":"Levine","year":"2007","journal-title":"Br. J. Sports Med."},{"key":"ref81","doi-asserted-by":"publisher","first-page":"313","DOI":"10.1249\/JSR.0000000000000168","article-title":"Is exercise really medicine? An evolutionary perspective","volume":"14","author":"Lieberman","year":"2015","journal-title":"Curr. Sports Med. Rep."},{"key":"ref82","doi-asserted-by":"publisher","first-page":"68","DOI":"10.1016\/j.palaeo.2016.09.019","article-title":"Radioisotopic age, formation, and preservation of late Pleistocene human footprints at Engare Sero, Tanzania","volume":"463","author":"Liutkus-Pierce","year":"2016","journal-title":"Palaeogeogr. Palaeoclimatol. Palaeoecol."},{"key":"ref83","doi-asserted-by":"publisher","first-page":"413","DOI":"10.1016\/S0140-6736(20)30367-6","article-title":"Dementia prevention, intervention, and care: 2020 report of the lancet commission","volume":"396","author":"Livingston","year":"2020","journal-title":"Lancet"},{"key":"ref84","doi-asserted-by":"publisher","first-page":"2673","DOI":"10.1016\/S0140-6736(17)31363-6","article-title":"Dementia prevention, intervention, and care","volume":"390","author":"Livingston","year":"2017","journal-title":"Lancet"},{"key":"ref85","doi-asserted-by":"crossref","first-page":"72","DOI":"10.1079\/9781786390271.0072","article-title":"\u201cThe Camino de Santiago de Compostela: the most important historic pilgrimage way in Europe,\u201d","volume-title":"Religious Pilgrimage Routes and Trails: Sustainable Development and Management.","author":"Lois-Gonz\u00e1lez","year":"2018"},{"key":"ref86","doi-asserted-by":"publisher","DOI":"10.4137\/EBo.s33489","article-title":"Human dispersal out of Africa: a lasting debate","volume":"11","author":"L\u00f3pez","year":"2015","journal-title":"Evol. 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Soc. Psychol."},{"key":"ref92","volume-title":"The Farther Reaches of Human Nature.","author":"Maslow","year":"1971"},{"key":"ref93","doi-asserted-by":"publisher","first-page":"272","DOI":"10.1177\/1745691612464657","article-title":"The psychological self as actor, agent, and author","volume":"8","author":"McAdams","year":"2013","journal-title":"Perspect. Psychol. Sci."},{"key":"ref94","doi-asserted-by":"publisher","first-page":"1","DOI":"10.26613\/esic.3.1.110","article-title":"\u201cFirst we invented stories, then they changed us\u201d: the evolution of narrative identity","volume":"3","author":"McAdams","year":"2019","journal-title":"Evol. Stud. Imaginative Cult."},{"key":"ref95","doi-asserted-by":"publisher","first-page":"475","DOI":"10.1177\/0956797614568318","article-title":"Narrating the generative life","volume":"26","author":"McAdams","year":"2015","journal-title":"Psychol. Sci."},{"key":"ref96","doi-asserted-by":"publisher","first-page":"2745","DOI":"10.1523\/JNEUROSCI.1812-17.2018","article-title":"Mind-wandering in people with hippocampal damage","volume":"38","author":"McCormick","year":"2018","journal-title":"J. Neurosci."},{"key":"ref97","doi-asserted-by":"publisher","first-page":"775","DOI":"10.1037\/a0040270","article-title":"Walking facilitates positive affect (even when expecting the opposite)","volume":"16","author":"Miller","year":"2016","journal-title":"Emotion"},{"key":"ref98","doi-asserted-by":"publisher","first-page":"132","DOI":"10.1038\/s41559-017-0397-y","article-title":"Social polymorphism is favoured by the co-evolution of dispersal with social behaviour","volume":"2","author":"Mullon","year":"2018","journal-title":"Nat. Ecol. (4).ris Evol."},{"key":"ref99","volume-title":"Newgrange: Monument to Immortality.","author":"Murphy","year":"2012"},{"key":"ref100","doi-asserted-by":"publisher","first-page":"1468","DOI":"10.1097\/00005768-200209000-00011","article-title":"Accumulating brisk walking for fitness, cardiovascular risk, and psychological health","volume":"34","author":"Murphy","year":"2002","journal-title":"Med. Sci. Sports Exerc."},{"key":"ref101","doi-asserted-by":"publisher","first-page":"1101","DOI":"10.1177\/0956797611418527","article-title":"Underestimating nearby nature: affective forecasting errors obscure the happy path to sustainability","volume":"22","author":"Nisbet","year":"2011","journal-title":"Psychol. Sci."},{"key":"ref102","doi-asserted-by":"publisher","first-page":"71","DOI":"10.1016\/j.humov.2017.10.007","article-title":"Audiovisual integration increases the intentional step synchronization of side-by-side walkers","volume":"56","author":"Noy","year":"2017","journal-title":"Hum. Mov. 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Neurosci."},{"key":"ref107","doi-asserted-by":"publisher","first-page":"425","DOI":"10.1016\/j.tics.2020.03.001","article-title":"Unraveling the mysteries of motivation","volume":"24","author":"O\u2019Reilly","year":"2020","journal-title":"Trends Cogn. Sci."},{"key":"ref108","doi-asserted-by":"publisher","first-page":"147","DOI":"10.1016\/S0169-328X(98)00222-8","article-title":"Exercise-induced regulation of brain-derived neurotrophic factor (BDNF) transcripts in the rat hippocampus","volume":"61","author":"Oliff","year":"1998","journal-title":"Mol. Brain Res."},{"key":"ref109","doi-asserted-by":"publisher","first-page":"1142","DOI":"10.1037\/a0036577","article-title":"Give your ideas some legs: the positive effect of walking on creative thinking","volume":"40","author":"Oppezzo","year":"2014","journal-title":"J. Exp. Psychol. Learn. Mem. 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Sci."},{"key":"ref135","doi-asserted-by":"publisher","first-page":"e56211","DOI":"10.7554\/eLife.56211","article-title":"Dreaming with hippocampal damage","volume":"9","author":"Span\u00f2","year":"2020","journal-title":"elife"},{"key":"ref136","doi-asserted-by":"publisher","first-page":"533","DOI":"10.1016\/j.tins.2020.04.010","article-title":"Effects of exercise on brain and cognition across age groups and health states","volume":"43","author":"Stillman","year":"2020","journal-title":"Trends Neurosci."},{"key":"ref137","doi-asserted-by":"publisher","DOI":"10.1037\/emo0000876","article-title":"Big smile, small self: awe walks promote prosocial positive emotions in older adults","author":"Sturm","year":"2020","journal-title":"Emotion."},{"key":"ref138","doi-asserted-by":"publisher","first-page":"441","DOI":"10.1016\/S0926-6410(01)00080-5","article-title":"Cortical and subcortical vestibular response to caloric stimulation detected by functional magnetic resonance imaging","volume":"12","author":"Suzuki","year":"2001","journal-title":"Cogn. 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