{"status":"ok","message-type":"work","message-version":"1.0.0","message":{"indexed":{"date-parts":[[2026,9,25]],"date-time":"2026-09-25T05:55:21Z","timestamp":1790315721171,"version":"4.1.0"},"reference-count":52,"publisher":"MDPI AG","issue":"8","license":[{"start":{"date-parts":[[2021,7,24]],"date-time":"2021-07-24T00:00:00Z","timestamp":1627084800000},"content-version":"vor","delay-in-days":0,"URL":"https:\/\/creativecommons.org\/licenses\/by\/4.0\/"}],"funder":[{"name":"MARS Petcare UK","award":["no number"],"award-info":[{"award-number":["no number"]}]},{"DOI":"10.13039\/100010732","name":"Marie-Claire Cronstedts Stiftelse","doi-asserted-by":"publisher","award":["no number"],"award-info":[{"award-number":["no number"]}],"id":[{"id":"10.13039\/100010732","id-type":"DOI","asserted-by":"publisher"}]}],"content-domain":{"domain":[],"crossmark-restriction":false},"short-container-title":["Animals"],"abstract":"<jats:p>Straw\u2019s low energy content means it is a roughage option for horses with low energy requirements. Previously, in a field study, straw was associated with an increased risk for gastric ulcers. This study evaluated the effect on gastric ulcers, metabolic profile and behaviour of replacing, in a forage-only ration, 50% of the daily allowance with wheat straw. Six equines were studied in a 2 \u00d7 21-day cross-over design. The control diet (CON: 100% grass forage) and the straw diet (S: 50% grass forage and 50% straw [DM basis]) were iso-energetic. Gastroscopy was performed prior to the study and on day 21 and blood samples were collected and behavioural observations were performed. Diet did not affect squamous or glandular gastric ulcer scores (p &gt; 0.05). Feed intake time was longer (p &lt; 0.05) plus energy intake and plasma insulin concentrations were lower on diet S compared to CON (p &lt; 0.0001). Plasma serotonin concentrations tended to be higher on diet S compared to CON (p = 0.05). The results suggest that good hygienic quality wheat straw can be included for up to 50% of the diet without causing gastric ulcers and that it can extend feeding time and promote a metabolic profile more suitable for overweight horses.<\/jats:p>","DOI":"10.3390\/ani11082197","type":"journal-article","created":{"date-parts":[[2021,7,25]],"date-time":"2021-07-25T22:06:21Z","timestamp":1627250781000},"page":"2197","update-policy":"https:\/\/doi.org\/10.3390\/mdpi_crossmark_policy","source":"Crossref","is-referenced-by-count":15,"title":["Straw as an Alternative to Grass Forage in Horses\u2014Effects on Post-Prandial Metabolic Profile, Energy Intake, Behaviour and Gastric Ulceration"],"prefix":"10.3390","volume":"11","author":[{"given":"Anna","family":"Jansson","sequence":"first","affiliation":[{"name":"Department of Anatomy, Physiology and Biochemistry, Swedish University of Agricultural Sciences, 75007 Uppsala, Sweden"}],"role":[{"vocabulary":"crossref","role":"author"}]},{"given":"Patricia","family":"Harris","sequence":"additional","affiliation":[{"name":"WALTHAM Petcare Science Institute Waltham-on-the Wolds, Melton Mowbray, Leics LE14 4RT, UK"}],"role":[{"vocabulary":"crossref","role":"author"}]},{"given":"Sara Larsdotter","family":"Davey","sequence":"additional","affiliation":[{"name":"University Animal Hospital, Swedish University of Agricultural Sciences, 75007 Uppsala, Sweden"}],"role":[{"vocabulary":"crossref","role":"author"}]},{"given":"Nanna","family":"Luthersson","sequence":"additional","affiliation":[{"name":"Hestedoktoren I\/S, Hvals\u00f8vej 298, 4360 Kr. Eskilstrup, Denmark"}],"role":[{"vocabulary":"crossref","role":"author"}]},{"given":"Sveinn","family":"Ragnarsson","sequence":"additional","affiliation":[{"name":"Department of Equine Science, H\u00f3lar University, IS-551 Sau\u00f0\u00e1rkr\u00f3kur, Iceland"}],"role":[{"vocabulary":"crossref","role":"author"}]},{"given":"Sara","family":"Ringmark","sequence":"additional","affiliation":[{"name":"Department of Anatomy, Physiology and Biochemistry, Swedish University of Agricultural Sciences, 75007 Uppsala, Sweden"}],"role":[{"vocabulary":"crossref","role":"author"}]}],"member":"1968","published-online":{"date-parts":[[2021,7,24]]},"reference":[{"key":"ref_1","first-page":"1","article-title":"Sm\u00e4ltbarhetsf\u00f6rs\u00f6k med h\u00e4star (Digestibility experiments on horses)","volume":"Meddelande nr 36","author":"Olsson","year":"1949","journal-title":"Lantbruksh\u00f6gskolan Husdjursf\u00f6rs\u00f6ksanstalten"},{"key":"ref_2","doi-asserted-by":"crossref","first-page":"347","DOI":"10.1017\/S0003356100023254","article-title":"Digestibility by ponies of oat straw treated with anhydrous ammonia","volume":"28","author":"Slagsvold","year":"1979","journal-title":"Anim. Sci."},{"key":"ref_3","doi-asserted-by":"crossref","first-page":"223","DOI":"10.1016\/S0737-0806(06)81450-4","article-title":"Digestibility of wheat straw or ammoniated wheat straw in equine diets","volume":"12","author":"Hansen","year":"1992","journal-title":"J. Equine Vet. Sci."},{"key":"ref_4","doi-asserted-by":"crossref","first-page":"335","DOI":"10.1111\/jvim.15423","article-title":"ECEIM consensus statement on equine metabolic syndrome","volume":"33","author":"Durham","year":"2019","journal-title":"J. Vet. Intern. Med."},{"key":"ref_5","doi-asserted-by":"crossref","first-page":"1","DOI":"10.12968\/ukve.2018.2.S2.3","article-title":"Equine obesity: Current perspectives","volume":"2","author":"Rendle","year":"2018","journal-title":"UK-Vet Equine"},{"key":"ref_6","doi-asserted-by":"crossref","first-page":"958","DOI":"10.1017\/S1751731116002469","article-title":"Review: Feeding conserved forage to horses: Recent advances and recommendations","volume":"11","author":"Harris","year":"2017","journal-title":"Animal"},{"key":"ref_7","first-page":"54","article-title":"Studies on the duration of feeding, masticatory frequency and mincing of feed in horses","volume":"82","author":"Meyer","year":"1975","journal-title":"DTW Deutsche Tierarztliche Wochenschrift"},{"key":"ref_8","doi-asserted-by":"crossref","first-page":"86","DOI":"10.1111\/j.2042-3306.1995.tb03041.x","article-title":"Management factors associated with stereotypic and redirected behaviour in the thoroughbred horse","volume":"27","author":"McGreevy","year":"1995","journal-title":"Equine Vet. J."},{"key":"ref_9","doi-asserted-by":"crossref","first-page":"475","DOI":"10.1017\/S1357729800009644","article-title":"Factors affecting behavioural disturbances in race-horses","volume":"66","author":"Redbo","year":"1998","journal-title":"Anim. Sci."},{"key":"ref_10","doi-asserted-by":"crossref","first-page":"297","DOI":"10.7120\/09627286.23.3.297","article-title":"Pre-feeding behaviour in UK leisure horses and associated feeding routine risk factors","volume":"23","author":"Hockenhull","year":"2014","journal-title":"Anim. Welf."},{"key":"ref_11","doi-asserted-by":"crossref","first-page":"625","DOI":"10.2746\/042516409X441929","article-title":"Risk factors associated with equine gastric ulceration syndrome (EGUS) in 201 horses in Denmark","volume":"41","author":"Luthersson","year":"2009","journal-title":"Equine Vet. J."},{"key":"ref_12","doi-asserted-by":"crossref","first-page":"125","DOI":"10.1016\/S0737-0806(01)70108-6","article-title":"Quality of different bedding materials and their influence on the compostability of horse manure","volume":"21","author":"Airaksinen","year":"2001","journal-title":"J. Equine Vet. Sci."},{"key":"ref_13","doi-asserted-by":"crossref","first-page":"432","DOI":"10.1111\/j.1439-0396.2007.00755.x","article-title":"Judgement of hygienic quality of roughage in horse stables in Switzerland","volume":"92","author":"Wichert","year":"2008","journal-title":"J. Anim. Physiol. Anim. Nutr."},{"key":"ref_14","doi-asserted-by":"crossref","first-page":"371","DOI":"10.1111\/j.2042-3306.1983.tb01826.x","article-title":"Relationship between condition score, physical measurements and body-fat percentage in mares","volume":"15","author":"Henneke","year":"1983","journal-title":"Equine Vet. J."},{"key":"ref_15","doi-asserted-by":"crossref","first-page":"784","DOI":"10.2460\/ajvr.2000.61.784","article-title":"Evaluation of diet as a cause of gastric ulcers in horses","volume":"61","author":"Nadeau","year":"2000","journal-title":"Am. J. Vet. Res."},{"key":"ref_16","doi-asserted-by":"crossref","first-page":"1288","DOI":"10.1111\/jvim.13578","article-title":"European College of Equine Internal Medicine Consensus Statement-Equine Gastric Ulcer Syndrome in Adult Horses","volume":"29","author":"Sykes","year":"2015","journal-title":"J. Vet. Intern. Med."},{"key":"ref_17","doi-asserted-by":"crossref","first-page":"795","DOI":"10.1111\/evj.12688","article-title":"Preliminary investigations into a novel, long-acting, injectable, intramuscular formulation of omeprazole in the horse","volume":"49","author":"Sykes","year":"2017","journal-title":"Equine Vet. J."},{"key":"ref_18","doi-asserted-by":"crossref","first-page":"169","DOI":"10.3920\/CEP13018","article-title":"A bibliometric review on nutrition of the exercising horse from 1970 to 2010","volume":"9","author":"Jansson","year":"2013","journal-title":"Comp. Exerc. Physiol."},{"key":"ref_19","unstructured":"NRC (2007). National Research Council Committee Nutrient Requirements of Horses, National Academies Press."},{"key":"ref_20","doi-asserted-by":"crossref","first-page":"746","DOI":"10.1017\/S1751731112002261","article-title":"Growth, training response and health in Standardbred yearlings fed a forage-only diet","volume":"7","author":"Ringmark","year":"2013","journal-title":"Animal"},{"key":"ref_21","doi-asserted-by":"crossref","first-page":"4869","DOI":"10.2527\/jas2017.1670","article-title":"Road transport and diet affect metabolic response to exercise in horses","volume":"95","author":"Connysson","year":"2017","journal-title":"J. Anim. Sci."},{"key":"ref_22","doi-asserted-by":"crossref","unstructured":"Connysson, M., Rhodin, M., and Jansson, A. (2019). Effects of Horse Housing System on Energy Balance during Post-Exercise Recovery. Animals, 9.","DOI":"10.3390\/ani9110976"},{"key":"ref_23","doi-asserted-by":"crossref","first-page":"83","DOI":"10.1079\/ECP200683","article-title":"Digestive and metabolic effects of altering feeding frequency in athletic horses","volume":"3","author":"Jansson","year":"2006","journal-title":"Equine Comp. Exerc. Physiol."},{"key":"ref_24","doi-asserted-by":"crossref","first-page":"103","DOI":"10.1111\/evj.12169","article-title":"Insulin dysregulation","volume":"46","author":"Frank","year":"2014","journal-title":"Equine Vet. J."},{"key":"ref_25","doi-asserted-by":"crossref","first-page":"e60","DOI":"10.1136\/vr.105793","article-title":"Inducing weight loss in native ponies: Is straw a viable alternative to hay?","volume":"187","author":"Dosi","year":"2020","journal-title":"Vet. Rec."},{"key":"ref_26","doi-asserted-by":"crossref","first-page":"171","DOI":"10.2746\/042516408X342975","article-title":"Prediction of incipient pasture-associated laminitis from hyperinsulinaemia, hyperleptinaemia and generalised and localised obesity in a cohort of ponies","volume":"41","author":"Carter","year":"2009","journal-title":"Equine Vet. J."},{"key":"ref_27","doi-asserted-by":"crossref","first-page":"986","DOI":"10.1016\/j.jevs.2014.05.006","article-title":"The Effect of Hay Net Design on Rate of Forage Consumption When Feeding Adult Horses","volume":"34","author":"Glunk","year":"2014","journal-title":"J. Equine Vet. Sci."},{"key":"ref_28","doi-asserted-by":"crossref","unstructured":"Saastamoinen, M., Fradinho, M.J., Santos, A.S., and Miraglia, N. (2012). Forage feeding systems. Forages and Grazing in Horse Nutrition, Wageningen Academic Publishers.","DOI":"10.3920\/978-90-8686-755-4"},{"key":"ref_29","doi-asserted-by":"crossref","first-page":"571","DOI":"10.2460\/ajvr.1990.51.04.571","article-title":"Effect of meal feeding on plasma volume and urinary electrolyte clearance in ponies","volume":"51","author":"Clarke","year":"1990","journal-title":"Am. J. Vet. Res."},{"key":"ref_30","doi-asserted-by":"crossref","first-page":"1610","DOI":"10.1152\/jappl.1999.86.5.1610","article-title":"Effects of feeding frequency and voluntary salt intake on fluid and electrolyte regulation in athletic horses","volume":"86","author":"Jansson","year":"1999","journal-title":"J. Appl. Physiol."},{"key":"ref_31","unstructured":"Jansson, A., Lindberg, J., Rundgren, M., M\u00fcller, C., Connysson, M., Kjellberg, L., and Lundberg, M. (2011). Utfodringsrekommendationer f\u00f6r h\u00e4st, Swedish University of Agricultural Sciences."},{"key":"ref_32","doi-asserted-by":"crossref","first-page":"117","DOI":"10.1152\/jappl.2002.92.1.117","article-title":"Glycemic index of a meal fed before exercise alters substrate use and glucose flux in exercising horses","volume":"92","author":"Hinchcliff","year":"2002","journal-title":"J. Appl. Physiol."},{"key":"ref_33","doi-asserted-by":"crossref","first-page":"311","DOI":"10.1016\/j.tvjl.2005.08.011","article-title":"Digestibility of a complete ration in horses fed once or three times a day and correlation with key blood parameters","volume":"173","author":"Buyse","year":"2007","journal-title":"Vet. J."},{"key":"ref_34","doi-asserted-by":"crossref","first-page":"25","DOI":"10.1186\/1754-6834-3-25","article-title":"Cellulosic ethanol: Interactions between cultivar and enzyme loading in wheat straw processing","volume":"3","author":"Lindedam","year":"2010","journal-title":"Biotechnol. Biofuels"},{"key":"ref_35","doi-asserted-by":"crossref","first-page":"1831","DOI":"10.1007\/s12155-015-9631-0","article-title":"Analysis of Wheat Straw Biodiversity for Use as a Feedstock for Biofuel Production","volume":"8","author":"Tishler","year":"2015","journal-title":"BioEnergy Res."},{"key":"ref_36","doi-asserted-by":"crossref","first-page":"133","DOI":"10.1136\/vr.c502","article-title":"Effect of diet on plasma tryptophan and serotonin in trained mares and geldings","volume":"166","author":"Alberghina","year":"2010","journal-title":"Vet. Rec."},{"key":"ref_37","doi-asserted-by":"crossref","first-page":"56","DOI":"10.1016\/j.biochi.2018.06.008","article-title":"Serotonin in the gut: Blessing or a curse","volume":"161","author":"Banskota","year":"2019","journal-title":"Biochimie"},{"key":"ref_38","doi-asserted-by":"crossref","first-page":"352","DOI":"10.1016\/j.cmet.2007.09.012","article-title":"Serotonin and the Orchestration of Energy Balance","volume":"6","author":"Tecott","year":"2007","journal-title":"Cell Metab."},{"key":"ref_39","doi-asserted-by":"crossref","first-page":"37","DOI":"10.1016\/0166-4328(96)00066-6","article-title":"Serotonergic control of the organization of feeding and satiety","volume":"73","author":"Simansky","year":"1996","journal-title":"Behav. Brain Res."},{"key":"ref_40","doi-asserted-by":"crossref","first-page":"88","DOI":"10.1016\/j.physbeh.2015.07.017","article-title":"Effects of chewing on appetite, food intake and gut hormones: A systematic review and meta-analysis","volume":"151","author":"Hetherington","year":"2015","journal-title":"Physiol. Behav."},{"key":"ref_41","doi-asserted-by":"crossref","first-page":"3","DOI":"10.1111\/j.1365-2036.2004.02180.x","article-title":"Review article: Serotonin receptors and transporters\u2014Roles in normal and abnormal gastrointestinal motility","volume":"20","author":"Gershon","year":"2004","journal-title":"Aliment. Pharmacol. Ther."},{"key":"ref_42","doi-asserted-by":"crossref","first-page":"588","DOI":"10.1016\/j.cmet.2012.09.014","article-title":"Gut-derived serotonin is a multifunctional determinant to fasting adaptation","volume":"16","author":"Sumara","year":"2012","journal-title":"Cell Metab."},{"key":"ref_43","doi-asserted-by":"crossref","first-page":"246","DOI":"10.1111\/evj.12990","article-title":"Assessing gastrointestinal motility in healthy horses comparing auscultation, ultrasonography and an acoustic gastrointestinal surveillance biosensor: A randomised, blinded, controlled crossover proof of principle study","volume":"51","author":"VanderBroek","year":"2019","journal-title":"Equine Vet. J."},{"key":"ref_44","doi-asserted-by":"crossref","first-page":"1419","DOI":"10.2460\/javma.2001.219.1419","article-title":"Feeding practices associated with colic in horses","volume":"219","author":"Hudson","year":"2001","journal-title":"J. Am. Vet. Med. Assoc."},{"key":"ref_45","doi-asserted-by":"crossref","first-page":"648","DOI":"10.1111\/j.2042-3306.2006.tb05620.x","article-title":"Effects on exercise response, fluid and acid-base balance of protein intake from forage-only diets in Standardbred horses","volume":"38","author":"Connysson","year":"2006","journal-title":"Equine Vet. J."},{"key":"ref_46","doi-asserted-by":"crossref","first-page":"341","DOI":"10.1111\/j.2042-3306.2010.00283.x","article-title":"Effects of crude protein intake from forage-only diets on muscle amino acids and glycogen levels in horses in training","volume":"42","author":"Connysson","year":"2010","journal-title":"Equine Vet. J."},{"key":"ref_47","doi-asserted-by":"crossref","first-page":"133","DOI":"10.1017\/S1478061509342334","article-title":"Effects on fluid balance, digestion and exercise response in Standardbred horses fed silage, haylage and hay","volume":"5","author":"Muhonen","year":"2008","journal-title":"Comp. Exerc. Physiol."},{"key":"ref_48","doi-asserted-by":"crossref","first-page":"2291","DOI":"10.2527\/jas.2008-1461","article-title":"Effects on the equine colon ecosystem of grass silage and haylage diets after an abrupt change from hay","volume":"87","author":"Muhonen","year":"2009","journal-title":"J. Anim. Sci."},{"key":"ref_49","doi-asserted-by":"crossref","first-page":"273","DOI":"10.1111\/j.1439-0396.2010.01049.x","article-title":"Comparison of grass haylage digestibility and metabolic plasma profile in Icelandic and Standardbred horses","volume":"95","author":"Ragnarsson","year":"2011","journal-title":"J. Anim. Physiol. Anim. Nutr."},{"key":"ref_50","doi-asserted-by":"crossref","first-page":"209","DOI":"10.3920\/CEP13014","article-title":"Insulin response to feeding forage with varying crude protein and amino acid content in horses at rest and after exercise","volume":"9","author":"Ringmark","year":"2013","journal-title":"Comp. Exerc. Physiol."},{"key":"ref_51","doi-asserted-by":"crossref","first-page":"1","DOI":"10.1017\/S1751731117000593","article-title":"Effects of training distance on feed intake, growth, body condition and muscle glycogen content in young Standardbred horses fed a forage-only diet","volume":"11","author":"Ringmark","year":"2017","journal-title":"Animal"},{"key":"ref_52","doi-asserted-by":"crossref","unstructured":"Muhonen, S., Connysson, M., Lindberg, J.E., Julliand, V., Bertilsson, J., and Jansson, A. (2008). Effects of crude protein intake from grass silage-only diets on the equine colon ecosystem after an abrupt feed change. J. Anim. 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