{"status":"ok","message-type":"work","message-version":"1.0.0","message":{"indexed":{"date-parts":[[2026,5,29]],"date-time":"2026-05-29T21:53:39Z","timestamp":1780091619858,"version":"3.54.0"},"reference-count":65,"publisher":"MDPI AG","issue":"20","license":[{"start":{"date-parts":[[2025,10,14]],"date-time":"2025-10-14T00:00:00Z","timestamp":1760400000000},"content-version":"vor","delay-in-days":0,"URL":"https:\/\/creativecommons.org\/licenses\/by\/4.0\/"}],"funder":[{"DOI":"10.13039\/501100005056","name":"the National Institute of Fisheries Science","doi-asserted-by":"publisher","award":["R2025009"],"award-info":[{"award-number":["R2025009"]}],"id":[{"id":"10.13039\/501100005056","id-type":"DOI","asserted-by":"publisher"}]}],"content-domain":{"domain":[],"crossmark-restriction":false},"short-container-title":["Animals"],"abstract":"<jats:p>Large-bodied sharks are key apex predators in marine ecosystems; however, ecological data from Korean waters are limited. From February to November 2024, 44 individuals from six species were collected. Of these, 24 individuals were analyzed for ecological characteristics: shortfin mako (Isurus oxyrinchus, n = 6), salmon shark (Lamna ditropis, n = 11), blue shark (Prionace glauca, n = 6), and great white shark (Carcharodon carcharias, n = 1). Age and growth parameters were estimated from vertebral band counts using the von Bertalanffy growth model. Diet was assessed via stomach content and DNA metabarcoding, and trophic relationships were examined using stable isotopes. The monthly occurrence peaked in July, with shortfin mako, salmon shark, and blue shark being the most frequently observed species. Estimated ages ranged 8\u201316, 4\u201313, and 1\u201311 years, respectively. The diets were predominantly fish-based, with species-specific prey preferences. Stable isotope data revealed trophic differentiation, suggesting niche partitioning among species. The eastern coastal waters of Korea appear to serve as foraging grounds and potential nursery habitats for large shark species. This is the first comprehensive ecological baseline for these species in Korean waters, which supports future assessments, conservation, and management strategies.<\/jats:p>","DOI":"10.3390\/ani15202974","type":"journal-article","created":{"date-parts":[[2025,10,14]],"date-time":"2025-10-14T13:41:29Z","timestamp":1760449289000},"page":"2974","update-policy":"https:\/\/doi.org\/10.3390\/mdpi_crossmark_policy","source":"Crossref","is-referenced-by-count":1,"title":["Ecological Characteristics of Large-Bodied Sharks in the East Sea of Korea"],"prefix":"10.3390","volume":"15","author":[{"given":"Gi Chang","family":"Seong","sequence":"first","affiliation":[{"name":"East Sea Fisheries Research Institute, National Institute of Fisheries Science, Gangneung 25435, Republic of Korea"}],"role":[{"vocabulary":"crossref","role":"author"}]},{"given":"Jeong-Ik","family":"Baek","sequence":"additional","affiliation":[{"name":"East Sea Fisheries Research Institute, National Institute of Fisheries Science, Gangneung 25435, Republic of Korea"}],"role":[{"vocabulary":"crossref","role":"author"}]},{"given":"Jong-Ku","family":"Gal","sequence":"additional","affiliation":[{"name":"East Sea Fisheries Research Institute, National Institute of Fisheries Science, Gangneung 25435, Republic of Korea"}],"role":[{"vocabulary":"crossref","role":"author"}]},{"given":"Sun-Kil","family":"Lee","sequence":"additional","affiliation":[{"name":"East Sea Fisheries Research Institute, National Institute of Fisheries Science, Gangneung 25435, Republic of Korea"}],"role":[{"vocabulary":"crossref","role":"author"}]},{"given":"Jeong-Min","family":"Shim","sequence":"additional","affiliation":[{"name":"East Sea Fisheries Research Institute, National Institute of Fisheries Science, Gangneung 25435, Republic of Korea"}],"role":[{"vocabulary":"crossref","role":"author"}]},{"given":"Maeng-Jin","family":"Kim","sequence":"additional","affiliation":[{"name":"East Sea Fisheries Research Institute, National Institute of Fisheries Science, Gangneung 25435, Republic of Korea"}],"role":[{"vocabulary":"crossref","role":"author"}]}],"member":"1968","published-online":{"date-parts":[[2025,10,14]]},"reference":[{"key":"ref_1","first-page":"611","article-title":"Importance of sharks in ocean ecosystem","volume":"8","author":"Motivarash","year":"2020","journal-title":"J. Entomol. Zool. Stud."},{"key":"ref_2","doi-asserted-by":"crossref","unstructured":"T\u00fcrko\u011flu, M., \u00d6nal, U., and Ismen, A. (2018). Overview on Mediterranean shark\u2019s fisheries: Impact on the biodiversity. Marine Ecology-Biotic and Abiotic Interactions, InTech.","DOI":"10.5772\/intechopen.69018"},{"key":"ref_3","doi-asserted-by":"crossref","first-page":"39","DOI":"10.60084\/ljes.v1i2.54","article-title":"Effect of life history on Alopias pelagicus overexploitation vulnerability: A literature review","volume":"1","author":"Hewindati","year":"2023","journal-title":"Leuser J. Environ. Stud."},{"key":"ref_4","doi-asserted-by":"crossref","first-page":"R565","DOI":"10.1016\/j.cub.2017.04.038","article-title":"Challenges and priorities in shark and ray conservation","volume":"27","author":"Dulvy","year":"2017","journal-title":"Curr. Biol."},{"key":"ref_5","doi-asserted-by":"crossref","first-page":"e00590","DOI":"10.7554\/eLife.00590","article-title":"Extinction risk and conservation of the world\u2019s sharks and rays","volume":"3","author":"Dulvy","year":"2014","journal-title":"eLife"},{"key":"ref_6","doi-asserted-by":"crossref","first-page":"4773","DOI":"10.1016\/j.cub.2021.08.062","article-title":"Overfishing drives over one-third of all sharks and rays toward a global extinction crisis","volume":"31","author":"Dulvy","year":"2021","journal-title":"Curr. Biol."},{"key":"ref_7","doi-asserted-by":"crossref","first-page":"111","DOI":"10.1007\/s00442-021-05075-7","article-title":"Forecasting intraspecific changes in distribution of a wide-ranging marine predator under climate change","volume":"198","author":"Niella","year":"2022","journal-title":"Oecologia"},{"key":"ref_8","unstructured":"Clarke, S., Yokawa, K., Matsunaga, H., and Nakano, H. (2011). Analysis of North Pacific Shark Data from Japanese Commercial Longline and Research\/Training Vessel Records, Western and Central Pacific Fisheries Commission."},{"key":"ref_9","doi-asserted-by":"crossref","first-page":"259","DOI":"10.1111\/fog.12149","article-title":"Distribution, body length, and abundance of blue shark and shortfin mako offshore of northeastern Japan, as determined from observed pelagic longline data, 2000\u20132014","volume":"25","author":"Ohshimo","year":"2016","journal-title":"Fish. Oceanogr."},{"key":"ref_10","first-page":"21","article-title":"CPUE standardization and catch estimate of blue shark by Taiwanese large-scale tuna longline fishery in the North Pacific Ocean","volume":"23","author":"Tsai","year":"2015","journal-title":"J. Mar. Sci. Technol."},{"key":"ref_11","doi-asserted-by":"crossref","first-page":"518","DOI":"10.1071\/MF11086","article-title":"The importance of research and public opinion to conservation management of sharks and rays: A synthesis","volume":"62","author":"Simpfendorfer","year":"2011","journal-title":"Mar. Freshw. Res."},{"key":"ref_12","doi-asserted-by":"crossref","first-page":"16055","DOI":"10.1002\/ece3.8277","article-title":"Defining distribution and habitat use of west--central Florida\u2019s coastal sharks through a research and education program","volume":"11","author":"Mullins","year":"2021","journal-title":"Ecol. Evol."},{"key":"ref_13","doi-asserted-by":"crossref","first-page":"270","DOI":"10.2112\/JCR-SI116-055.1","article-title":"Comparative analysis of changes in fish catches by species and sea surface temperature change in coastal waters of the Republic of Korea","volume":"116","author":"Park","year":"2024","journal-title":"J. Coast. Res."},{"key":"ref_14","doi-asserted-by":"crossref","first-page":"298","DOI":"10.47853\/FAS.2022.e27","article-title":"Preliminary study on spatio-temporal variations of five giant and 17 large fish species around the Korean peninsula from 2011 to 2016","volume":"25","author":"Kim","year":"2022","journal-title":"Fish. Aquat. Sci."},{"key":"ref_15","first-page":"254","article-title":"Feeding habits of 6 shark species in the Southern Sea of Korea","volume":"43","author":"Huh","year":"2010","journal-title":"Korean J. Fish. Aquat. Sci."},{"key":"ref_16","first-page":"44","article-title":"Distribution of the white shark, Carcharodon carcharias and other sharks around the Korean waters","volume":"21","author":"Choi","year":"2009","journal-title":"Korean J. Ichthyol."},{"key":"ref_17","first-page":"595","article-title":"First reliable record of Echinorhinus cookei (Chondrichthyes: Elasmobranchii) collected from Busan, Korea","volume":"51","author":"Lee","year":"2018","journal-title":"Korean J. Fish. Aquat. Sci."},{"key":"ref_18","first-page":"965","article-title":"Two unrecorded species of sharks in Korean waters","volume":"53","author":"Choi","year":"2020","journal-title":"Korean J. Fish. Aquat. Sci."},{"key":"ref_19","doi-asserted-by":"crossref","first-page":"84","DOI":"10.35399\/ISK.36.1.10","article-title":"Taxonomic system of sharks (Chondrichthyes: Elasmobranchii) in Korean waters","volume":"36","author":"Kim","year":"2024","journal-title":"Korean J. Ichthyol."},{"key":"ref_20","unstructured":"Choi, Y., Kim, J.-G., Im, Y., and Kim, H. (2022). Pictorial Guidebook to Sharks of Korean Coastal Waters, National Institute of Fisheries Science."},{"key":"ref_21","doi-asserted-by":"crossref","first-page":"7","DOI":"10.1186\/s40462-018-0125-5","article-title":"Summer at the beach: Spatio-temporal patterns of white shark occurrence along the inshore areas of False Bay, South Africa","volume":"6","author":"Kock","year":"2018","journal-title":"Mov. Ecol."},{"key":"ref_22","doi-asserted-by":"crossref","first-page":"1529","DOI":"10.1139\/cjfas-2020-0036","article-title":"Using temporally explicit habitat suitability models to infer the migratory pattern of a large mobile shark","volume":"77","author":"Irigoyen","year":"2020","journal-title":"Can. J. Fish. Aquat. Sci."},{"key":"ref_23","doi-asserted-by":"crossref","first-page":"10","DOI":"10.1577\/1548-8446(2002)027<0010:ECIFM>2.0.CO;2","article-title":"Ecological considerations in fisheries management: When does it matter?","volume":"27","author":"Link","year":"2002","journal-title":"Fisheries"},{"key":"ref_24","doi-asserted-by":"crossref","unstructured":"Flinn, S.A., and Midway, S.R. (2021). Trends in growth modeling in fisheries science. Fishes, 6.","DOI":"10.3390\/fishes6010001"},{"key":"ref_25","first-page":"627","article-title":"Age and growth of the blue shark (Prionace glauca) in the North Atlantic Ocean","volume":"101","author":"Skomal","year":"2003","journal-title":"Fish. Bull.-Natl. Ocean. Atmos. Adm."},{"key":"ref_26","doi-asserted-by":"crossref","first-page":"143","DOI":"10.1071\/MF05077","article-title":"Age, growth, maturity, longevity and natural mortality of the shortfin mako shark (Isurus oxyrinchus) in New Zealand waters","volume":"57","author":"Bishop","year":"2006","journal-title":"Mar. Freshw. Res."},{"key":"ref_27","first-page":"278","article-title":"Growth and maturity of salmon sharks (Lamna ditropis) in the eastern and western North Pacific, and comments on back-calculation methods","volume":"104","author":"Goldman","year":"2006","journal-title":"Fish. Bull."},{"key":"ref_28","unstructured":"Goldman, K.J., Cailliet, G.M., Andrews, A.H., and Natanson, L.J. (2012). Assessing the age and growth of chondrichthyan fishes. Biol. Sharks Their Relat., 13."},{"key":"ref_29","doi-asserted-by":"crossref","unstructured":"Huxley, J.S. (1932). Problems of Relative Growth, Dover Publishing.","DOI":"10.5962\/bhl.title.6427"},{"key":"ref_30","first-page":"181","article-title":"A quantitative theory of organic growth (inquiries on growth laws. II)","volume":"10","year":"1938","journal-title":"Hum. Biol."},{"key":"ref_31","doi-asserted-by":"crossref","first-page":"141","DOI":"10.2307\/1538217","article-title":"A new graphic method of describing the growth of animals","volume":"90","author":"Walford","year":"1946","journal-title":"Biol. Bull."},{"key":"ref_32","unstructured":"R Core Team (2024). R: A Language and Environment for Statistical Computing, R Foundation for Statistical Computing. Available online: https:\/\/www.R-project.org\/."},{"key":"ref_33","unstructured":"Kim, I.S., Choi, Y., Lee, C.L., Lee, Y.J., Kim, B.J., and Kim, J.H. (2005). Illustrated Book of Korean Fishes, Kyo-Hak Publ."},{"key":"ref_34","unstructured":"Kim, J.K., Ryu, J.H., Kwun, H.J., Ji, H.S., Park, J.H., Myoung, S.H., Song, Y.S., Lee, S.J., Yu, H.J., and Bae, S.E. (2019). Distribution Map of Sea Fishes in the Korean Peninsula, Mapledesign Publ."},{"key":"ref_35","first-page":"1","article-title":"Food habits of albacore, bluefin tuna, and bonito in California waters","volume":"152","author":"Pinkas","year":"1971","journal-title":"Fish. Bull."},{"key":"ref_36","doi-asserted-by":"crossref","first-page":"6310","DOI":"10.1002\/ece3.6362","article-title":"In silico and empirical evaluation of twelve metabarcoding primer sets for insectivorous diet analyses","volume":"10","author":"Tournayre","year":"2020","journal-title":"Ecol. Evol."},{"key":"ref_37","doi-asserted-by":"crossref","unstructured":"Tabassum, N., Lee, J.-H., Lee, S.-R., Kim, J.-U., Park, H., Kim, H.-W., and Kim, J.-H. (2022). Molecular diet analysis of Ad\u00e9lie penguins (Pygoscelis adeliae) in the Ross Sea using fecal DNA. Biology, 11.","DOI":"10.3390\/biology11020182"},{"key":"ref_38","doi-asserted-by":"crossref","first-page":"581","DOI":"10.1038\/nmeth.3869","article-title":"DADA2: High-resolution sample inference from Illumina amplicon data","volume":"13","author":"Callahan","year":"2016","journal-title":"Nat. Methods"},{"key":"ref_39","doi-asserted-by":"crossref","first-page":"408","DOI":"10.2307\/1935376","article-title":"Nonsynchronous spatial overlap of lizards in patchy habitats","volume":"51","author":"Schoener","year":"1970","journal-title":"Ecology"},{"key":"ref_40","doi-asserted-by":"crossref","first-page":"595","DOI":"10.1111\/j.1365-2656.2011.01806.x","article-title":"Comparing isotopic niche widths among and within communities: SIBER\u2013Stable Isotope Bayesian Ellipses in R","volume":"80","author":"Jackson","year":"2011","journal-title":"J. Anim. Ecol."},{"key":"ref_41","doi-asserted-by":"crossref","first-page":"42","DOI":"10.1890\/0012-9658(2007)88[42:CSIRPF]2.0.CO;2","article-title":"Can stable isotope ratios provide for community-wide measures of trophic structure?","volume":"88","author":"Layman","year":"2007","journal-title":"Ecology"},{"key":"ref_42","doi-asserted-by":"crossref","unstructured":"Santos, C.C., Domingo, A., Carlson, J., Natanson, L.J., Travassos, P., Mac\u00edas, D., Cort\u00e9s, E., Miller, P., Hazin, F., and Mas, F. (2021). Movements, habitat use, and diving behavior of shortfin mako in the Atlantic Ocean. Front. Mar. Sci., 8.","DOI":"10.3389\/fmars.2021.686343"},{"key":"ref_43","doi-asserted-by":"crossref","first-page":"2117","DOI":"10.1890\/14-2113.1","article-title":"Evaluating the landscape of fear between apex predatory sharks and mobile sea turtles across a large dynamic seascape","volume":"96","author":"Hammerschlag","year":"2015","journal-title":"Ecology"},{"key":"ref_44","doi-asserted-by":"crossref","first-page":"1569","DOI":"10.1111\/ecog.04463","article-title":"Behavioral response of a mobile marine predator to environmental variables differs across ecoregions","volume":"42","author":"Byrne","year":"2019","journal-title":"Ecography"},{"key":"ref_45","doi-asserted-by":"crossref","first-page":"4381","DOI":"10.1038\/s41467-022-32035-3","article-title":"Small-scale fisheries catch more threatened elasmobranchs inside partially protected areas than in unprotected areas","volume":"13","author":"Milisenda","year":"2022","journal-title":"Nat. Commun."},{"key":"ref_46","doi-asserted-by":"crossref","first-page":"432","DOI":"10.1111\/j.1365-2656.2011.01914.x","article-title":"L\u00e9vy flight and Brownian search patterns of a free--ranging predator reflect different prey field characteristics","volume":"81","author":"Sims","year":"2012","journal-title":"J. Anim. Ecol."},{"key":"ref_47","doi-asserted-by":"crossref","unstructured":"Nasby-Lucas, N., Dewar, H., Sosa-Nishizaki, O., Wilson, C., Hyde, J.R., Vetter, R.D., Wraith, J., Block, B.A., Kinney, M.J., and Sippel, T. (2019). Movements of electronically tagged shortfin mako sharks (Isurus oxyrinchus) in the eastern North Pacific Ocean. Anim. Biotelemetry, 7.","DOI":"10.1186\/s40317-019-0174-6"},{"key":"ref_48","doi-asserted-by":"crossref","unstructured":"Smoothey, A.F., Lee, K.A., and Peddemors, V.M. (2019). Long-term patterns of abundance, residency and movements of bull sharks (Carcharhinus leucas) in Sydney Harbour, Australia. Sci. Rep., 9.","DOI":"10.1038\/s41598-019-54365-x"},{"key":"ref_49","doi-asserted-by":"crossref","unstructured":"Balchin, G.P., Schuller, A., Di Stefano, I., Robertson, M., Pollard, K., and Hughes, W.O.H. (2025). Seasonality, long-term trends and co-occurrence of sharks in a top predator assemblage. PLoS ONE, 20.","DOI":"10.1371\/journal.pone.0318011"},{"key":"ref_50","doi-asserted-by":"crossref","first-page":"402","DOI":"10.1002\/mcf2.10029","article-title":"Size, age, and spatial\u2013temporal distribution of shortfin mako in the Mexican Pacific Ocean","volume":"10","author":"Tavares","year":"2018","journal-title":"Mar. Coast. Fish."},{"key":"ref_51","doi-asserted-by":"crossref","first-page":"14","DOI":"10.1016\/j.fishres.2005.05.004","article-title":"Age and growth of the shortfin mako shark, Isurus oxyrinchus, from the western coast of Baja California Sur, Mexico","volume":"76","year":"2005","journal-title":"Fish. Res."},{"key":"ref_52","first-page":"269","article-title":"Age, growth and maturity of blue shark (Prionace glauca) in the northwest Atlantic Ocean","volume":"80","author":"Carlson","year":"2023","journal-title":"Collect. Vol. Sci. Pap. ICCAT"},{"key":"ref_53","doi-asserted-by":"crossref","first-page":"394","DOI":"10.1071\/MF08125","article-title":"Age and growth of the shortfin mako (Isurus oxyrinchus) in the south-eastern Pacific off Chile","volume":"60","author":"Cerna","year":"2009","journal-title":"Mar. Freshw. Res."},{"key":"ref_54","doi-asserted-by":"crossref","first-page":"1346","DOI":"10.1071\/MF17098","article-title":"Age and growth estimates of the blue shark Prionace glauca in the central South Pacific Ocean","volume":"69","author":"Joung","year":"2018","journal-title":"Mar. Freshw. Res."},{"key":"ref_55","doi-asserted-by":"crossref","first-page":"20","DOI":"10.1071\/MF10123","article-title":"Size at maturity and reproductive traits of shortfin mako, Isurus oxyrinchus, in the western and central North Pacific","volume":"62","author":"Semba","year":"2011","journal-title":"Mar. Freshw. Res."},{"key":"ref_56","first-page":"101","article-title":"Reproductive biology of the blue shark Prionace glauca (Chondrichthyes: Carcharhinidae) off Baja California Sur, M\u00e9xico","volume":"16","year":"2010","journal-title":"aqua"},{"key":"ref_57","doi-asserted-by":"crossref","first-page":"18","DOI":"10.1016\/j.fishres.2014.03.003","article-title":"Sexual development, size at maturity, size at maternity and fecundity of the blue shark Prionace glauca (Linnaeus, 1758) in the Southwest Atlantic","volume":"160","author":"Cardoso","year":"2014","journal-title":"Fish. Res."},{"key":"ref_58","doi-asserted-by":"crossref","first-page":"245","DOI":"10.3354\/meps268245","article-title":"Foraging ecology of the early life stages of four sympatric shark species","volume":"268","author":"Bethea","year":"2004","journal-title":"Mar. Ecol. Prog. Ser."},{"key":"ref_59","doi-asserted-by":"crossref","first-page":"3719","DOI":"10.1111\/mec.12785","article-title":"An holistic ecological analysis of the diet of Cory\u2019s shearwaters using prey morphological characters and DNA barcoding","volume":"23","author":"Alonso","year":"2014","journal-title":"Mol. Ecol."},{"key":"ref_60","doi-asserted-by":"crossref","first-page":"2665","DOI":"10.1139\/f94-266","article-title":"Modeling predator\u2013prey dynamics in a fluctuating environment","volume":"51","author":"Collie","year":"1994","journal-title":"Can. J. Fish. Aquat. Sci."},{"key":"ref_61","doi-asserted-by":"crossref","unstructured":"Arn\u00e9s-Urgell\u00e9s, C., Salinas-de-Le\u00f3n, P., Rastoin-Laplane, E., Vaca-Pita, L., Su\u00e1rez-Moncada, J., and P\u00e1ez-Rosas, D. (2021). The effects of climatic variability on the feeding ecology of the scalloped hammerhead shark (Sphyrna lewini) in the tropical Eastern Pacific. Front. Mar. Sci., 8.","DOI":"10.3389\/fmars.2021.625748"},{"key":"ref_62","doi-asserted-by":"crossref","first-page":"299","DOI":"10.1038\/s41559-017-0432-z","article-title":"A global perspective on the trophic geography of sharks","volume":"2","author":"Bird","year":"2018","journal-title":"Nat. Ecol. Evol."},{"key":"ref_63","doi-asserted-by":"crossref","first-page":"26","DOI":"10.1016\/j.fishres.2008.12.002","article-title":"An ecosystem-based fisheries assessment approach for Korean fisheries","volume":"100","author":"Zhang","year":"2009","journal-title":"Fish. Res."},{"key":"ref_64","doi-asserted-by":"crossref","unstructured":"Adams, K.R., Gibbs, L., Knott, N.A., Broad, A., Hing, M., Taylor, M.D., and Davis, A.R. (2020). Coexisting with sharks: A novel, socially acceptable and non-lethal shark mitigation approach. Sci. Rep., 10.","DOI":"10.1038\/s41598-020-74270-y"},{"key":"ref_65","doi-asserted-by":"crossref","first-page":"e9885","DOI":"10.1002\/ece3.9885","article-title":"The rise of animal biotelemetry and genetics research data integration","volume":"13","author":"Banks","year":"2023","journal-title":"Ecol. Evol."}],"container-title":["Animals"],"original-title":[],"language":"en","link":[{"URL":"https:\/\/www.mdpi.com\/2076-2615\/15\/20\/2974\/pdf","content-type":"unspecified","content-version":"vor","intended-application":"similarity-checking"}],"deposited":{"date-parts":[[2025,10,15]],"date-time":"2025-10-15T06:50:58Z","timestamp":1760511058000},"score":1,"resource":{"primary":{"URL":"https:\/\/www.mdpi.com\/2076-2615\/15\/20\/2974"}},"subtitle":[],"short-title":[],"issued":{"date-parts":[[2025,10,14]]},"references-count":65,"journal-issue":{"issue":"20","published-online":{"date-parts":[[2025,10]]}},"alternative-id":["ani15202974"],"URL":"https:\/\/doi.org\/10.3390\/ani15202974","relation":{},"ISSN":["2076-2615"],"issn-type":[{"value":"2076-2615","type":"electronic"}],"subject":[],"published":{"date-parts":[[2025,10,14]]}}}