{"status":"ok","message-type":"work","message-version":"1.0.0","message":{"indexed":{"date-parts":[[2026,2,24]],"date-time":"2026-02-24T13:21:57Z","timestamp":1771939317618,"version":"3.50.1"},"reference-count":13,"publisher":"MDPI AG","issue":"5","license":[{"start":{"date-parts":[[2026,2,24]],"date-time":"2026-02-24T00:00:00Z","timestamp":1771891200000},"content-version":"vor","delay-in-days":0,"URL":"https:\/\/creativecommons.org\/licenses\/by\/4.0\/"}],"content-domain":{"domain":[],"crossmark-restriction":false},"short-container-title":["Animals"],"abstract":"<jats:p>The long hair phenotype of the dog is ascribed to variants in the fibroblast growth factor 5 (FGF5) gene. Currently, there are five variant alleles known, Lh1 through Lh5, with any combination of these alleles resulting in a long-haired phenotype in dogs. Recently, genotyping of several long-haired dogs for the known Lh1\u2013Lh5 variants yielded unexpected results as these dogs contained no or only one variant in the FGF5 gene known to result in the long-hair phenotype. Analysis of the coding region for the FGF5 gene identified four novel variants associated with the long-haired phenotype. One dog of mixed ancestry had an insertion of G very similar to and near to the long-hair variant Lh4, NC_006614.4: g.37352832insCC. A group of 24 Tibetan Mastiffs showed the Lh1 variant NC_006614.4:g.37372096C&gt;A in 11 dogs only, and no other known Lh variant. Within the coding frame of the FGF5 gene, three unknown variants were discovered that were predicted to have a phenotypic effect. These new variants were inherited in a Mendelian fashion and, in combination with one of the known FGF5 alleles, likely predict the long-hair phenotype. Lastly, eight of the 24 Tibetan Mastiff dogs carried three FGF5 variants.<\/jats:p>","DOI":"10.3390\/ani16050699","type":"journal-article","created":{"date-parts":[[2026,2,24]],"date-time":"2026-02-24T12:19:23Z","timestamp":1771935563000},"page":"699","update-policy":"https:\/\/doi.org\/10.3390\/mdpi_crossmark_policy","source":"Crossref","is-referenced-by-count":0,"title":["Discovery of Four New FGF5 Variants Causing Long Hair in the Dog"],"prefix":"10.3390","volume":"16","author":[{"ORCID":"https:\/\/orcid.org\/0009-0001-3977-2089","authenticated-orcid":false,"given":"Robin E.","family":"Everts","sequence":"first","affiliation":[{"name":"AlphaDogDNA by Etalon Inc., Menlo Park, CA 94025, USA"}],"role":[{"role":"author","vocabulary":"crossref"}]},{"ORCID":"https:\/\/orcid.org\/0009-0006-7203-8720","authenticated-orcid":false,"given":"Tim","family":"Roane","sequence":"additional","affiliation":[{"name":"AlphaDogDNA by Etalon Inc., Menlo Park, CA 94025, USA"}],"role":[{"role":"author","vocabulary":"crossref"}]},{"ORCID":"https:\/\/orcid.org\/0009-0007-1639-1615","authenticated-orcid":false,"given":"Rachael","family":"Caron","sequence":"additional","affiliation":[{"name":"AlphaDogDNA by Etalon Inc., Menlo Park, CA 94025, USA"}],"role":[{"role":"author","vocabulary":"crossref"}]},{"given":"Cameron","family":"Kunstadt","sequence":"additional","affiliation":[{"name":"AlphaDogDNA by Etalon Inc., Menlo Park, CA 94025, USA"}],"role":[{"role":"author","vocabulary":"crossref"}]},{"ORCID":"https:\/\/orcid.org\/0000-0002-8810-2014","authenticated-orcid":false,"given":"Gabriel","family":"Foster","sequence":"additional","affiliation":[{"name":"AlphaDogDNA by Etalon Inc., Menlo Park, CA 94025, USA"}],"role":[{"role":"author","vocabulary":"crossref"}]},{"given":"Christa","family":"Lafayette","sequence":"additional","affiliation":[{"name":"AlphaDogDNA by Etalon Inc., Menlo Park, CA 94025, USA"}],"role":[{"role":"author","vocabulary":"crossref"}]}],"member":"1968","published-online":{"date-parts":[[2026,2,24]]},"reference":[{"key":"ref_1","doi-asserted-by":"crossref","first-page":"3","DOI":"10.1111\/age.13154","article-title":"Canine Coat Pigmentation Genetics: A Review","volume":"53","author":"Brancalion","year":"2022","journal-title":"Anim. 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