{"status":"ok","message-type":"work","message-version":"1.0.0","message":{"indexed":{"date-parts":[[2026,8,27]],"date-time":"2026-08-27T05:18:11Z","timestamp":1787807891620,"version":"build-2784847793"},"reference-count":51,"publisher":"MDPI AG","issue":"9","license":[{"start":{"date-parts":[[2026,8,25]],"date-time":"2026-08-25T00:00:00Z","timestamp":1787616000000},"content-version":"vor","delay-in-days":0,"URL":"https:\/\/creativecommons.org\/licenses\/by\/4.0\/"}],"funder":[{"DOI":"10.13039\/501100004608","name":"Natural Science Foundation of Jiangsu Province","doi-asserted-by":"crossref","award":["BK20220743"],"award-info":[{"award-number":["BK20220743"]}],"id":[{"id":"10.13039\/501100004608","id-type":"DOI","asserted-by":"crossref"}]},{"name":"Open Research Fund of the State Key Laboratory of Vegetable Biobreeding","award":["SKLVB202602"],"award-info":[{"award-number":["SKLVB202602"]}]},{"name":"Jiangsu Provincial Key R&D Programme-Modern Agriculture","award":["BE2022339"],"award-info":[{"award-number":["BE2022339"]}]},{"DOI":"10.13039\/501100001809","name":"National Natural Science Foundation of China","doi-asserted-by":"publisher","award":["32202489"],"award-info":[{"award-number":["32202489"]}],"id":[{"id":"10.13039\/501100001809","id-type":"DOI","asserted-by":"publisher"}]},{"award":["32202489"],"award-info":[{"award-number":["32202489"]}],"id":[{"id":"https:\/\/ror.org\/01h0zpd94","id-type":"ROR","asserted-by":"publisher"}]}],"content-domain":{"domain":[],"crossmark-restriction":false},"short-container-title":["Antioxidants"],"abstract":"<jats:p>Drought is one of the major environmental constraints limiting tomato growth and productivity. Identifying regulators that enhance drought tolerance without compromising plant growth is therefore important for tomato production. Homeodomain-leucine zipper (HD-Zip) transcription factors (TFs) play essential roles in plant development and abiotic stress responses; however, the functions of most HD-Zip II members in tomato remain poorly understood. Here, we identified SlHZ07, a drought-responsive HD-Zip II TF, through transcriptome analysis and characterized its biological function in tomato. SlHZ07 was rapidly induced by drought stress and localized predominantly to the nucleus. Overexpression of SlHZ07 significantly enhanced drought tolerance, whereas RNAi-mediated suppression increased drought sensitivity. Physiological analyses showed that SlHZ07 overexpression reduced reactive oxygen species (ROS) accumulation, enhanced antioxidant enzyme activities, upregulated expression of ROS-scavenging genes, and alleviated membrane damage under drought stress. Hormone analyses revealed that SlHZ07 positively regulated jasmonic acid (JA) accumulation and the expression of JA biosynthetic genes, including OPR2, OPR3, JAR1, and AOC, but did not alter endogenous abscisic acid (ABA) levels under well-watered conditions. Furthermore, SlHZ07 promoted vegetative growth by increasing endogenous gibberellin (GA) levels and upregulating the expression of the GA biosynthetic genes GA20ox2 and GA20ox4. Together, our findings identify SlHZ07 as a previously uncharacterized positive regulator that coordinates plant growth and drought adaptation by integrating GA biosynthesis, JA homeostasis, and ROS detoxification. These results expand our understanding of HD-Zip II TFs and provide a promising genetic target for improving drought tolerance in tomato.<\/jats:p>","DOI":"10.3390\/antiox15091062","type":"journal-article","created":{"date-parts":[[2026,8,25]],"date-time":"2026-08-25T12:57:17Z","timestamp":1787662637000},"page":"1062","update-policy":"https:\/\/doi.org\/10.3390\/mdpi_crossmark_policy","source":"Crossref","is-referenced-by-count":0,"title":["The HD-Zip II Transcription Factor SlHZ07 Promotes Growth and Drought Tolerance in Tomato"],"prefix":"10.3390","volume":"15","author":[{"given":"Shuchao","family":"Dong","sequence":"first","affiliation":[{"name":"Jiangsu Key Laboratory for Genetic Improvement and Efficient Utilization of Horticultural Crops, Institute of Vegetable Crops, Jiangsu Academy of Agricultural Sciences, Nanjing 210014, China"},{"name":"State Key Laboratory of Vegetable Biobreeding, Institute of Vegetables and Flowers, Chinese Academy of Agricultural Sciences, Beijing 100081, China"}],"role":[{"vocabulary":"crossref","role":"author"}]},{"given":"Jiaxin","family":"Li","sequence":"additional","affiliation":[{"name":"Jiangsu Key Laboratory for Genetic Improvement and Efficient Utilization of Horticultural Crops, Institute of Vegetable Crops, Jiangsu Academy of Agricultural Sciences, Nanjing 210014, China"},{"name":"College of Horticulture and Plant Protection, Yangzhou University, Yangzhou 225100, China"}],"role":[{"vocabulary":"crossref","role":"author"}]},{"given":"Jingwen","family":"Zhang","sequence":"additional","affiliation":[{"name":"Jiangsu Key Laboratory for Genetic Improvement and Efficient Utilization of Horticultural Crops, Institute of Vegetable Crops, Jiangsu Academy of Agricultural Sciences, Nanjing 210014, China"},{"name":"College of Horticulture, Nanjing Agricultural University, Nanjing 210095, China"}],"role":[{"vocabulary":"crossref","role":"author"}]},{"ORCID":"https:\/\/orcid.org\/0000-0003-2853-5800","authenticated-orcid":false,"given":"Liuxia","family":"Song","sequence":"additional","affiliation":[{"name":"Jiangsu Key Laboratory for Genetic Improvement and Efficient Utilization of Horticultural Crops, Institute of Vegetable Crops, Jiangsu Academy of Agricultural Sciences, Nanjing 210014, China"}],"role":[{"vocabulary":"crossref","role":"author"}]},{"given":"Yinlei","family":"Wang","sequence":"additional","affiliation":[{"name":"Jiangsu Key Laboratory for Genetic Improvement and Efficient Utilization of Horticultural Crops, Institute of Vegetable Crops, Jiangsu Academy of Agricultural Sciences, Nanjing 210014, China"}],"role":[{"vocabulary":"crossref","role":"author"}]},{"given":"Liping","family":"Zhao","sequence":"additional","affiliation":[{"name":"Jiangsu Key Laboratory for Genetic Improvement and Efficient Utilization of Horticultural Crops, Institute of Vegetable Crops, Jiangsu Academy of Agricultural Sciences, Nanjing 210014, China"}],"role":[{"vocabulary":"crossref","role":"author"}]},{"given":"Jie","family":"Chen","sequence":"additional","affiliation":[{"name":"Jiangsu Key Laboratory for Genetic Improvement and Efficient Utilization of Horticultural Crops, Institute of Vegetable Crops, Jiangsu Academy of Agricultural Sciences, Nanjing 210014, China"}],"role":[{"vocabulary":"crossref","role":"author"}]},{"given":"Yariv","family":"Brotman","sequence":"additional","affiliation":[{"name":"School of Plant Sciences and Food Security, Institute for Cereal Crops Research, Tel Aviv University, Tel Aviv 6997801, Israel"}],"role":[{"vocabulary":"crossref","role":"author"}]},{"given":"Junming","family":"Li","sequence":"additional","affiliation":[{"name":"State Key Laboratory of Vegetable Biobreeding, Institute of Vegetables and Flowers, Chinese Academy of Agricultural Sciences, Beijing 100081, China"}],"role":[{"vocabulary":"crossref","role":"author"}]},{"given":"Tongmin","family":"Zhao","sequence":"additional","affiliation":[{"name":"Jiangsu Key Laboratory for Genetic Improvement and Efficient Utilization of Horticultural Crops, Institute of Vegetable Crops, Jiangsu Academy of Agricultural Sciences, Nanjing 210014, China"},{"name":"College of Horticulture and Plant Protection, Yangzhou University, Yangzhou 225100, China"},{"name":"College of Horticulture, Nanjing Agricultural University, Nanjing 210095, China"}],"role":[{"vocabulary":"crossref","role":"author"}]}],"member":"1968","published-online":{"date-parts":[[2026,8,25]]},"reference":[{"key":"ref_1","doi-asserted-by":"crossref","unstructured":"Li, Y., Henke, M., Zhang, D., Wang, C., and Wei, M. 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