{"status":"ok","message-type":"work","message-version":"1.0.0","message":{"indexed":{"date-parts":[[2026,7,11]],"date-time":"2026-07-11T10:38:43Z","timestamp":1783766323496,"version":"3.55.0"},"reference-count":76,"publisher":"MDPI AG","issue":"4","license":[{"start":{"date-parts":[[2023,3,31]],"date-time":"2023-03-31T00:00:00Z","timestamp":1680220800000},"content-version":"vor","delay-in-days":0,"URL":"https:\/\/creativecommons.org\/licenses\/by\/4.0\/"}],"funder":[{"name":"\u00daNKP-22-4 New National Excellence Program of the Ministry for Culture","award":["NKFIH FK 135 136, PB, GV; NKFIH KKP 144068, JP"],"award-info":[{"award-number":["NKFIH FK 135 136, PB, GV; NKFIH KKP 144068, JP"]}]},{"name":"\u00daNKP-22-4 New National Excellence Program of the Ministry for Culture","award":["BO-00106-21 (PB)"],"award-info":[{"award-number":["BO-00106-21 (PB)"]}]},{"name":"National Research Development and Innovation Office","award":["NKFIH FK 135 136, PB, GV; NKFIH KKP 144068, JP"],"award-info":[{"award-number":["NKFIH FK 135 136, PB, GV; NKFIH KKP 144068, JP"]}]},{"name":"National Research Development and Innovation Office","award":["BO-00106-21 (PB)"],"award-info":[{"award-number":["BO-00106-21 (PB)"]}]},{"name":"J\u00e1nos Bolyai Research Scholarship of the Hungarian Academy of Sciences","award":["NKFIH FK 135 136, PB, GV; NKFIH KKP 144068, JP"],"award-info":[{"award-number":["NKFIH FK 135 136, PB, GV; NKFIH KKP 144068, JP"]}]},{"name":"J\u00e1nos Bolyai Research Scholarship of the Hungarian Academy of Sciences","award":["BO-00106-21 (PB)"],"award-info":[{"award-number":["BO-00106-21 (PB)"]}]},{"name":"\u00daNKP-22-5 New National Excellence Program of the Ministry for Innovation and Technology from the source of the National Research, Development and Innovation Fund (PB)","award":["NKFIH FK 135 136, PB, GV; NKFIH KKP 144068, JP"],"award-info":[{"award-number":["NKFIH FK 135 136, PB, GV; NKFIH KKP 144068, JP"]}]},{"name":"\u00daNKP-22-5 New National Excellence Program of the Ministry for Innovation and Technology from the source of the National Research, Development and Innovation Fund (PB)","award":["BO-00106-21 (PB)"],"award-info":[{"award-number":["BO-00106-21 (PB)"]}]}],"content-domain":{"domain":[],"crossmark-restriction":false},"short-container-title":["Insects"],"abstract":"<jats:p>(1) Bioclimatic factors have a proven effect on species distributions in terrestrial, marine, or freshwater ecosystems. Because of anthropogenic effects, the changes in these variables are accelerated; thus, the knowledge of the impact has great importance from a conservation point of view. Two endemic dragonflies, the Balkan Goldenring (Cordulegaster heros) and the Two-Toothed Goldenring (C. bidentata), confined to the hilly and mountainous regions in Europe, are classified as \u201cNear Threatened\u201d according to the IUCN Red List. (2) Modeling the potential occurrence of both species under present and future climatic conditions provides a more accurate picture of the most suitable areas. The models were used to predict the responses of both species to 6 different climate scenarios for the year 2070. (3) We revealed which climatic and abiotic variables affect them the most and which areas are the most suitable for the species. We calculated how future climatic changes would affect the range of suitable areas for the two species. (4) According to our results, the suitable area for Cordulegaster bidentata and C. heros are strongly influenced by bioclimatic variables and showed an upward shift toward high elevations. The models predict a loss of suitable area in the case of C. bidentata and a large gain in the case of C. heros.<\/jats:p>","DOI":"10.3390\/insects14040348","type":"journal-article","created":{"date-parts":[[2023,3,31]],"date-time":"2023-03-31T08:27:27Z","timestamp":1680251247000},"page":"348","update-policy":"https:\/\/doi.org\/10.3390\/mdpi_crossmark_policy","source":"Crossref","is-referenced-by-count":10,"title":["Winners and Losers: Cordulegaster Species under the Pressure of Climate Change"],"prefix":"10.3390","volume":"14","author":[{"given":"Judit","family":"Fekete","sequence":"first","affiliation":[{"name":"Research Group of Limnology, Centre of Natural Science, University of Pannonia, Egyetem St. 10, H-8200 Veszpr\u00e9m, Hungary"},{"name":"Centre for Ecological Research, Institute of Aquatic Ecology, Department of Tisza Research, 18\/c Bem Sq., H-4026 Debrecen, Hungary"}],"role":[{"vocabulary":"crossref","role":"author"}]},{"ORCID":"https:\/\/orcid.org\/0000-0002-7958-1420","authenticated-orcid":false,"given":"Geert","family":"De Knijf","sequence":"additional","affiliation":[{"name":"Research Institute of Nature and Forest (INBO), Havenlaan 88 bus 73, 1000 Brussels, Belgium"}],"role":[{"vocabulary":"crossref","role":"author"}]},{"given":"Marco","family":"Dinis","sequence":"additional","affiliation":[{"name":"CIBIO\/InBIO, Centro de Investiga\u00e7\u00e3o em Biodiversidade e Recursos Gen\u00e9ticos da Universidade do Porto, Instituto de Ci\u00eancias Agr\u00e1rias de Vair\u00e3o, R. Padre Armando Quintas n\u25e6 7, 4485-661 Vair\u00e3o, Portugal"},{"name":"Departamento de Biologia, Faculdade de Ci\u00eancias, Universidade do Porto, 4099-002 Porto, Portugal"},{"name":"BIOPOLIS Program in Genomics, Biodiversity and Land Planning, CIBIO, Campus de Vair\u00e3o, 4485-661 Vair\u00e3o, Portugal"}],"role":[{"vocabulary":"crossref","role":"author"}]},{"ORCID":"https:\/\/orcid.org\/0000-0001-8285-2896","authenticated-orcid":false,"given":"Judit","family":"Padis\u00e1k","sequence":"additional","affiliation":[{"name":"Research Group of Limnology, Centre of Natural Science, University of Pannonia, Egyetem St. 10, H-8200 Veszpr\u00e9m, Hungary"},{"name":"ELKH-PE Limnoecology Research Group, Egyetem St. 10, H-8200 Veszpr\u00e9m, Hungary"}],"role":[{"vocabulary":"crossref","role":"author"}]},{"ORCID":"https:\/\/orcid.org\/0000-0002-1825-5744","authenticated-orcid":false,"given":"P\u00e1l","family":"Boda","sequence":"additional","affiliation":[{"name":"Centre for Ecological Research, Institute of Aquatic Ecology, Department of Tisza Research, 18\/c Bem Sq., H-4026 Debrecen, Hungary"}],"role":[{"vocabulary":"crossref","role":"author"}]},{"given":"Edv\u00e1rd","family":"Mizsei","sequence":"additional","affiliation":[{"name":"Kiskuns\u00e1g National Park Directorate, Liszt F. St. 19, H-6000 Kecskem\u00e9t, Hungary"},{"name":"Department of Ecology, University of Debrecen, Egyetem Sq. 1, H-4010 Debrecen, Hungary"}],"role":[{"vocabulary":"crossref","role":"author"}]},{"ORCID":"https:\/\/orcid.org\/0000-0001-5907-3472","authenticated-orcid":false,"given":"G\u00e1bor","family":"V\u00e1rb\u00edr\u00f3","sequence":"additional","affiliation":[{"name":"Centre for Ecological Research, Institute of Aquatic Ecology, Department of Tisza Research, 18\/c Bem Sq., H-4026 Debrecen, Hungary"}],"role":[{"vocabulary":"crossref","role":"author"}]}],"member":"1968","published-online":{"date-parts":[[2023,3,31]]},"reference":[{"key":"ref_1","unstructured":"Masson-Delmotte, V., Zhai, P., Pirani, A., Connors, S.L., P\u00e9an, C., Berger, S., Caud, N., Chen, Y., Goldfarb, L., and Gomis, M.I. (2021). Summary for Policymakers. Climate Change 2021: The Physical Science Basis. Contribution of Working Group I to the Sixth Assessment Report of the Intergovernmental Panel on Climate Change, Cambridge University Press."},{"key":"ref_2","doi-asserted-by":"crossref","first-page":"134","DOI":"10.1017\/S0376892902000097","article-title":"Threats to the running water ecosystems of the world","volume":"29","author":"Malmqvist","year":"2002","journal-title":"Environ. Conserv."},{"key":"ref_3","doi-asserted-by":"crossref","first-page":"1053","DOI":"10.1007\/s00267-009-9329-1","article-title":"Climate change and river ecosystems: Protection and adaptation options","volume":"44","author":"Palmer","year":"2009","journal-title":"Environ. Manag."},{"key":"ref_4","unstructured":"Corbet, P.S. (1999). Dragonflies: Behaviour and Ecology of Odonata, Harley Books."},{"key":"ref_5","doi-asserted-by":"crossref","first-page":"1546","DOI":"10.2307\/1939134","article-title":"Regulation of Freshwater Community Structure at Multiple Intensities of Dragonfly Predation","volume":"65","author":"Thorp","year":"1984","journal-title":"Ecology"},{"key":"ref_6","doi-asserted-by":"crossref","first-page":"857","DOI":"10.2307\/1941035","article-title":"Interactive effects of dragonfly predation in experimental pond communities","volume":"69","year":"1988","journal-title":"Ecology"},{"key":"ref_7","doi-asserted-by":"crossref","unstructured":"Worthen, W., Fravel, R., and Horne, C. (2021). Downstream Changes in Odonate (Insecta: Odonata) Communities along a Suburban to Urban Gradient: Untangling Natural and Anthropogenic Effects. Insects, 12.","DOI":"10.3390\/insects12030201"},{"key":"ref_8","unstructured":"Collins, N.M., and Thomas, J.A. (1991). Conservation of neotropical environments: Insects as indicators. The Conservation of Insects and Their Habitats, Academic Press."},{"key":"ref_9","doi-asserted-by":"crossref","first-page":"279","DOI":"10.1016\/S0006-3207(96)00032-8","article-title":"Dragonfly (Odonata) distribution patterns in urban and forest landscapes, and recommendations for riparian management","volume":"78","author":"Samways","year":"1996","journal-title":"Biol. Conserv."},{"key":"ref_10","doi-asserted-by":"crossref","first-page":"517","DOI":"10.1071\/MF9820517","article-title":"Effect of sewage effluent on dragonflies of Bulimba Creek, Brisbane","volume":"33","author":"Watson","year":"1982","journal-title":"Aust. J. Mar. Freshw. Res."},{"key":"ref_11","unstructured":"Kalkman, V.J., Boudot, J.-P., Bernard, R., Conze, K.-J., De Knijf, G., Dyatlova, E., Ferreira, S., Jovi\u0107, M., Ott, J., and Riservato, E. (2010). European Red List of Dragonflies, Publications Office of the European Union."},{"key":"ref_12","unstructured":"Boudot, J.P., and Kalkman, V.J. (2015). Atlas of the European Dragonflies and Damselflies, KNNV Publishing."},{"key":"ref_13","doi-asserted-by":"crossref","first-page":"155","DOI":"10.1080\/13887890.2020.1739150","article-title":"Flying activity and population dynamics of Cordulegaster heros Theischinger, 1979 (Insecta: Odonata: Cordulegastridae) in Slovakia","volume":"23","author":"Fric","year":"2020","journal-title":"Int. J. Odonatol."},{"key":"ref_14","doi-asserted-by":"crossref","first-page":"121","DOI":"10.1007\/s10750-015-2317-y","article-title":"Life history and multiscale habitat preferences of the red-listed Balkan Goldenring, Cordulegaster heros Theischinger, 1979 (Insecta, Odonata), in South-Hungarian headwaters: Does the species have mesohabitat-mediated microdistribution?","volume":"760","author":"Boda","year":"2015","journal-title":"Hydrobiologia"},{"key":"ref_15","first-page":"11","article-title":"Further data on Cordulegaster bidentata Selys, 1873 from the territory of B\u00fckk National Park","volume":"42","author":"Fekete","year":"2018","journal-title":"Folia Hist.-Nat. Musei Matra."},{"key":"ref_16","unstructured":"(2022). IUCN (2021): International Union for Conservation of Nature Annual Report, IUCN."},{"key":"ref_17","first-page":"395","article-title":"Larval habitats and longitudinal distribution patterns of Cordulegaster heros Theischinger and C. bidentata S\u00e9lys in an Austrian forest stream (Anisoptera: Cordulegastridae)","volume":"30","author":"Lang","year":"2001","journal-title":"Odonatologica"},{"key":"ref_18","doi-asserted-by":"crossref","first-page":"49","DOI":"10.1111\/bij.12190","article-title":"Molecular phylogeny of the Western Palaearctic Cordulegaster taxa (Odonata: Anisoptera: Cordulegastridae)","volume":"111","author":"Froufe","year":"2014","journal-title":"Biol. J. Linn. Soc."},{"key":"ref_19","first-page":"51","article-title":"Adatok az \u00c9szakborsodi-hegyvid\u00e9k Odonata faun\u00e1j\u00e1hoz","volume":"19","author":"Ambrus","year":"1994","journal-title":"Folia Hist.-Nat. Musei Matra."},{"key":"ref_20","first-page":"1","article-title":"Odonata from Montenegro, with notes on taxonomy, regional diversity and conservation","volume":"42","author":"Vanappelghem","year":"2013","journal-title":"Odonatologica"},{"key":"ref_21","unstructured":"Boudot, J.-P. IUCN Red List of Threatened Species (IUCN SSC Dragonfly Specialist Group) Cordulegaster heros: 2018."},{"key":"ref_22","first-page":"53","article-title":"New data on the distribution of Large Golden Ringed Dragonfly (Cordulegaster heros Theischinger, 1979) (Odonata) in Zselic hills","volume":"15","author":"Ferincz","year":"2009","journal-title":"Nat. Som."},{"key":"ref_23","first-page":"447","article-title":"Adatok a (Cordulegaster bidentata S\u00e9lys, 1843) \u00e9s a k\u00e9tcs\u00edkos hegyiszitak\u00f6t\u0151 (Cordulegaster heros Theischinger, 1979) elterjed\u00e9s\u00e9hez a Bakonyban","volume":"18","year":"2012","journal-title":"Term\u00e9szetv\u00e9delmi K\u00f6zlem\u00e9nyek"},{"key":"ref_24","first-page":"407","article-title":"Rheobionte Libellen (Odonata) in Fliessgew\u00e4ssern der L\u00fcneburger Heide (Niedersachsen)","volume":"3","author":"Bahlo","year":"1989","journal-title":"Braunschw. Nat. Schr."},{"key":"ref_25","doi-asserted-by":"crossref","first-page":"1144","DOI":"10.1086\/682688","article-title":"Modeling the distribution of odonates: A review","volume":"34","author":"Collins","year":"2015","journal-title":"Freshw. Sci."},{"key":"ref_26","doi-asserted-by":"crossref","first-page":"295","DOI":"10.1656\/1092-6194(2004)011[0295:SATSAS]2.0.CO;2","article-title":"Spatial and Temporal Segregation Among Six Species of Coexisting Ophiogomphus (Odonata: Gomphidae) in the Aroostook River, Maine","volume":"11","author":"Gibbs","year":"2004","journal-title":"Northeast. Nat."},{"key":"ref_27","doi-asserted-by":"crossref","first-page":"341","DOI":"10.1656\/045.017.0217","article-title":"Status and Ecology of a Rare Gomphid Dragonfly at the Northern Extent of its Range","volume":"17","author":"Corser","year":"2010","journal-title":"Northeast. Nat."},{"key":"ref_28","doi-asserted-by":"crossref","first-page":"252","DOI":"10.1111\/j.1752-4598.2010.00096.x","article-title":"New records and a niche model for the distribution of two Neotropical damselflies: Schistolobos boliviensis and Tuberculobasis inversa (Odonata: Coenagrionidae)","volume":"3","author":"Cortes","year":"2010","journal-title":"Insect Conserv. Divers."},{"key":"ref_29","doi-asserted-by":"crossref","first-page":"2009","DOI":"10.1111\/j.1365-2427.2011.02631.x","article-title":"Climate-change winners and losers: Stream macroinvertebrates of a submontane region in Central Europe","volume":"56","author":"Domisch","year":"2011","journal-title":"Freshw. Biol."},{"key":"ref_30","doi-asserted-by":"crossref","first-page":"752","DOI":"10.1111\/gcb.12107","article-title":"Modelling distribution in European stream macroinvertebrates under future climates","volume":"19","author":"Domisch","year":"2013","journal-title":"Glob. Chang. Biol."},{"key":"ref_31","doi-asserted-by":"crossref","first-page":"4239","DOI":"10.1007\/s10531-005-3577-z","article-title":"Application of predictive distribution modelling to invertebrates: Odonata in South Africa","volume":"15","author":"Finch","year":"2006","journal-title":"Biodivers. Conserv."},{"key":"ref_32","unstructured":"Howard, T.G. (2006). Salmon River Watershed Inventory and Landscape Analysis, New York Natural Heritage Program."},{"key":"ref_33","doi-asserted-by":"crossref","first-page":"1768","DOI":"10.1126\/science.1156831","article-title":"A Significant Upward Shift in Plant Species Optimum Elevation during the 20th Century","volume":"320","author":"Lenoir","year":"2008","journal-title":"Science"},{"key":"ref_34","doi-asserted-by":"crossref","first-page":"111","DOI":"10.1080\/13887890.2011.578565","article-title":"The status of two boreo-alpine species,Somatochlora alpestrisandS. arctica, in Romania and their vulnerability to the impact of climate change (Odonata: Corduliidae)","volume":"14","author":"Flenker","year":"2011","journal-title":"Int. J. Odonatol."},{"key":"ref_35","unstructured":"Levin, S. (2013). Latitudinal and Elevational Range Shifts under Contemporary Climate Change. Encyclopedia of Biodiversity, Elsevier. [2nd ed.]."},{"key":"ref_36","doi-asserted-by":"crossref","first-page":"677","DOI":"10.1146\/annurev.ecolsys.110308.120159","article-title":"Species Distribution Models: Ecological Explanation and Prediction across Space and Time","volume":"40","author":"Elith","year":"2009","journal-title":"Annu. Rev. Ecol. Evol. Syst."},{"key":"ref_37","doi-asserted-by":"crossref","unstructured":"Kaschner, K., Tittensor, D.P., Ready, J., Gerrodette, T., and Worm, B. (2011). Current and Future Patterns of Global Marine Mammal Biodiversity. PLoS ONE, 6.","DOI":"10.1371\/journal.pone.0019653"},{"key":"ref_38","doi-asserted-by":"crossref","first-page":"55","DOI":"10.1163\/15685381-00003031","article-title":"Species distribution modelling leads to the discovery of new populations of one of the least known European snakes, Vipera ursinii graeca, in Albania","volume":"37","author":"Mizsei","year":"2016","journal-title":"Amphib. -Reptil."},{"key":"ref_39","doi-asserted-by":"crossref","unstructured":"Srivastava, V., Lafond, V., and Griess, V.C. (2019). Species distribution models (SDM): Applications, benefits and challenges in invasive species management. CABI Rev., 1\u20133.","DOI":"10.1079\/PAVSNNR201914020"},{"key":"ref_40","doi-asserted-by":"crossref","first-page":"2699","DOI":"10.1111\/gcb.12525","article-title":"Seeing the woods for the trees\u2014When is microclimate important in species distribution models?","volume":"20","author":"Bennie","year":"2014","journal-title":"Glob. Chang. Biol."},{"key":"ref_41","doi-asserted-by":"crossref","first-page":"664","DOI":"10.1111\/ecog.02360","article-title":"NicheMapR\u2014An R package for biophysical modelling: The microclimate model","volume":"40","author":"Kearney","year":"2017","journal-title":"Ecography"},{"key":"ref_42","doi-asserted-by":"crossref","unstructured":"Peterson, A.T., Sober\u00f3n, J., Pearson, R.G., Anderson, R.P., Mart\u00ednez-Meyer, E., Nakamura, M., and Ara\u00fajo, M.B. (2011). Ecological niches and geographic distributions (MPB-49). Ecological Niches and Geographic Distributions (MPB-49), Princeton University Press.","DOI":"10.23943\/princeton\/9780691136868.001.0001"},{"key":"ref_43","doi-asserted-by":"crossref","first-page":"1424","DOI":"10.1111\/ele.12189","article-title":"Predicting species distributions for conservation decisions","volume":"16","author":"Guisan","year":"2013","journal-title":"Ecol. Lett."},{"key":"ref_44","doi-asserted-by":"crossref","first-page":"963","DOI":"10.1007\/s10531-021-02126-6","article-title":"High-resolution topographic variables accurately predict the distribution of rare plant species for conservation area selection in a narrow-endemism hotspot in New Caledonia","volume":"30","author":"Lannuzel","year":"2021","journal-title":"Biodivers. Conserv."},{"key":"ref_45","unstructured":"Core Writing Team, Pachauri, R.K., and Meyer, L.A. (2014). Climate Change 2014: Synthesis Report. Contribution of Working Groups I, II and III to the Fifth Assessment Report of the Intergovernmental Panel on Climate Change, IPCC."},{"key":"ref_46","doi-asserted-by":"crossref","first-page":"305","DOI":"10.1080\/14772000.2012.705357","article-title":"Sampling bias in geographic and environmental space and its effect on the predictive power of species distribution models","volume":"10","author":"Bystriakova","year":"2012","journal-title":"Syst. Biodivers."},{"key":"ref_47","doi-asserted-by":"crossref","first-page":"4302","DOI":"10.1002\/joc.5086","article-title":"WorldClim 2: New 1-km spatial resolution climate surfaces for global land areas","volume":"37","author":"Fick","year":"2017","journal-title":"Int. J. Climatol."},{"key":"ref_48","unstructured":"(2020, July 16). WorldClim, Maps, Graphs, Tables, and DATA of the Globla Climate. Available online: http:\/\/www.worldclim.org\/."},{"key":"ref_49","unstructured":"Hijmans, R.J. (2019, April 22). (2020) raster: Geographic Data Analysis and Modeling. R package version 2.8\u201319. Available online: https:\/\/cran.r-project.org\/package=raster."},{"key":"ref_50","unstructured":"Bivand, R., Rundel, C., Pebesma, E., Stuetz, R., Hufthammer, K.O., and Bivand, M.R. Package \u2018rgeos\u2019. The Comprehensive R Archive Network (CRAN)., 2017."},{"key":"ref_51","doi-asserted-by":"crossref","first-page":"321","DOI":"10.1111\/j.1472-4642.2010.00641.x","article-title":"Moving beyond static species distribution models in support of conservation biogeography: Moving beyond static species distribution models","volume":"16","author":"Franklin","year":"2010","journal-title":"Divers. Distrib."},{"key":"ref_52","doi-asserted-by":"crossref","first-page":"1704","DOI":"10.1111\/j.1365-2699.2006.01460.x","article-title":"Model-based uncertainty in species range prediction","volume":"33","author":"Pearson","year":"2006","journal-title":"J. Biogeogr."},{"key":"ref_53","doi-asserted-by":"crossref","first-page":"853","DOI":"10.1111\/2041-210X.12372","article-title":"fuzzySim: Applying fuzzy logic to binary similarity indices in ecology","volume":"6","author":"Barbosa","year":"2015","journal-title":"Methods Ecol. Evol."},{"key":"ref_54","doi-asserted-by":"crossref","first-page":"656","DOI":"10.1111\/2041-210X.13360","article-title":"GCM compare R: A web application to assess differences and assist in the selection of general circulation models for climate change research","volume":"11","author":"Fajardo","year":"2020","journal-title":"Methods Ecol. Evol."},{"key":"ref_55","doi-asserted-by":"crossref","first-page":"369","DOI":"10.1111\/j.1600-0587.2008.05742.x","article-title":"Package \u2018biomod2\u2032. Species distribution modeling within an ensemble forecasting framework","volume":"32","author":"Thuiller","year":"2016","journal-title":"Ecography"},{"key":"ref_56","doi-asserted-by":"crossref","first-page":"802","DOI":"10.1111\/j.1365-2656.2008.01390.x","article-title":"A working guide to boosted regression trees","volume":"77","author":"Elith","year":"2008","journal-title":"J. Anim. Ecol."},{"key":"ref_57","unstructured":"Ridgeway, G. (2018, April 12). Generalized Boosted Models: A Guide to the Gbm Package. Available online: http:\/\/gradientboostedmodels.googlecode.com\/git\/gbm\/inst\/doc\/gbm.pdf."},{"key":"ref_58","doi-asserted-by":"crossref","first-page":"5","DOI":"10.1023\/A:1010933404324","article-title":"Random forests","volume":"45","author":"Breiman","year":"2001","journal-title":"Mach. Learn."},{"key":"ref_59","doi-asserted-by":"crossref","first-page":"861","DOI":"10.1016\/j.patrec.2005.10.010","article-title":"An introduction to ROC analysis","volume":"27","author":"Fawcett","year":"2006","journal-title":"Pattern Recognit. Lett."},{"key":"ref_60","doi-asserted-by":"crossref","first-page":"385","DOI":"10.1111\/j.0906-7590.2005.03957.x","article-title":"Selecting thresholds of occurrence in the prediction of species distributions","volume":"28","author":"Liu","year":"2005","journal-title":"Ecography"},{"key":"ref_61","unstructured":"Lewin-Koh, J.N., and Bivand, R. (2022, December 02). \u201cPackage \u2018Maptools\u2019: Tools for Reading and Handling Spatial Objects,\u201d R Package Version 1.1-6. Available online: https:\/\/cran.r-project.org\/web\/packages\/maptools\/index.html."},{"key":"ref_62","doi-asserted-by":"crossref","unstructured":"Wickham, H. (2016). ggplot2: Elegant Graphics for Data Analysis, Springer-Verlag. Available online: https:\/\/ggplot2.tidyverse.org.","DOI":"10.1007\/978-3-319-24277-4_9"},{"key":"ref_63","unstructured":"Allan, J.D., Palmer, M., and Poff, N.L. (2005). Climate change and freshwater ecosystems. Climate Change and Biodiversity, Yale University Press."},{"key":"ref_64","doi-asserted-by":"crossref","first-page":"1557","DOI":"10.1111\/j.1365-2486.2005.001008.x","article-title":"Scenarios of freshwater fish extinctions from climate change and water withdrawal","volume":"11","author":"Xenopoulos","year":"2005","journal-title":"Glob. Chang. Biol."},{"key":"ref_65","doi-asserted-by":"crossref","first-page":"e113749","DOI":"10.1371\/journal.pone.0113749","article-title":"Improving the Use of Species Distribution Models in Conservation Planning and Management under Climate Change","volume":"9","author":"Porfirio","year":"2014","journal-title":"PLoS ONE"},{"key":"ref_66","unstructured":"Ambrus, A., Danyik, T., Kov\u00e1cs, T., and Olajos, P. (2018). Magyarorsz\u00e1g Szitak\u00f6t\u0151inek K\u00e9zik\u00f6nyve, Magyar Term\u00e9szettudom\u00e1nyi M\u00fazeum."},{"key":"ref_67","unstructured":"Hynes, H.B.N. (1970). The Ecology of Running Waters, Liverpool University Press."},{"key":"ref_68","doi-asserted-by":"crossref","first-page":"269","DOI":"10.1007\/s10750-017-3495-6","article-title":"Diversity and conservation of European dragonflies and damselflies (Odonata)","volume":"811","author":"Kalkman","year":"2018","journal-title":"Hydrobiologia"},{"key":"ref_69","unstructured":"Boudot, J.-P. IUCN Red List of Threatened Species (IUCN SSC Dragonfly Specialist Group) Cordulegaster bidentata: 2018."},{"key":"ref_70","doi-asserted-by":"crossref","first-page":"403","DOI":"10.1111\/geb.12004","article-title":"Recent range shifts of European dragonflies provide support for an inverse relationship between habitat predictability and dispersal","volume":"22","author":"Grewe","year":"2012","journal-title":"Glob. Ecol. Biogeogr."},{"key":"ref_71","doi-asserted-by":"crossref","first-page":"502","DOI":"10.1111\/j.1365-2486.2005.00904.x","article-title":"A northward shift of range margins in British Odonata","volume":"11","author":"Hickling","year":"2005","journal-title":"Glob. Chang. Biol."},{"key":"ref_72","doi-asserted-by":"crossref","first-page":"450","DOI":"10.1111\/j.1365-2486.2006.01116.x","article-title":"The distributions of a wide range of taxonomic groups are expanding polewards","volume":"12","author":"Hickling","year":"2006","journal-title":"Glob. Chang. Biol."},{"key":"ref_73","doi-asserted-by":"crossref","first-page":"1457","DOI":"10.1111\/een.12931","article-title":"What to do if streams go dry? Behaviour of Balkan Goldenring (Cordulegaster heros, Odonata) larvae in a simulated drought experiment in SW Hungary","volume":"45","author":"Pernecker","year":"2020","journal-title":"Ecol. Entomol."},{"key":"ref_74","first-page":"3","article-title":"Les Cordulegaster du Pal\u00e9arctique occidental: Identification et r\u00e9partition (Odonata, Anisoptera, Cordulegastridae)","volume":"17","author":"Boudot","year":"2001","journal-title":"Martinia"},{"key":"ref_75","doi-asserted-by":"crossref","unstructured":"Schneider, T., Vierstraete, A., M\u00fcller, O., van Pelt, G.J., Caspers, M., Ikemeyer, D., Snegovaya, N., and Dumont, H.J. (2021). Taxonomic revision of eastern part of Western Palaearctic Cordulegaster using molecular phylogeny and morphology, with the description of two new species (Odonata: Anisoptera: Cordulegastridae). 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