{"status":"ok","message-type":"work","message-version":"1.0.0","message":{"indexed":{"date-parts":[[2026,9,22]],"date-time":"2026-09-22T19:00:08Z","timestamp":1790103608033,"version":"4.0.1"},"reference-count":92,"publisher":"MDPI AG","issue":"9","license":[{"start":{"date-parts":[[2026,9,21]],"date-time":"2026-09-21T00:00:00Z","timestamp":1789948800000},"content-version":"vor","delay-in-days":0,"URL":"https:\/\/creativecommons.org\/licenses\/by\/4.0\/"}],"funder":[{"DOI":"10.13039\/501100019370","name":"FCT\u2014Portuguese Foundation for Science and Technology","doi-asserted-by":"publisher","award":["UID\/04033\/2025"],"award-info":[{"award-number":["UID\/04033\/2025"]}],"id":[{"id":"10.13039\/501100019370","id-type":"DOI","asserted-by":"publisher"}]},{"DOI":"10.13039\/501100019370","name":"FCT\u2014Portuguese Foundation for Science and Technology","doi-asserted-by":"publisher","award":["LA\/P\/0126\/2020"],"award-info":[{"award-number":["LA\/P\/0126\/2020"]}],"id":[{"id":"10.13039\/501100019370","id-type":"DOI","asserted-by":"publisher"}]},{"DOI":"10.13039\/501100019370","name":"FCT\u2014Portuguese Foundation for Science and Technology","doi-asserted-by":"publisher","award":["UIDB\/04621\/2025"],"award-info":[{"award-number":["UIDB\/04621\/2025"]}],"id":[{"id":"10.13039\/501100019370","id-type":"DOI","asserted-by":"publisher"}]}],"content-domain":{"domain":[],"crossmark-restriction":false},"short-container-title":["Insects"],"abstract":"<jats:p>Interrow cover crops have been increasingly adopted in Mediterranean vineyards to improve soil conservation and other ecosystem services. However, their long-term effects on vegetation dynamics and soil surface arthropod communities remain poorly understood. A study was conducted over three sampling years (2017, 2018 and 2023) in the Douro Demarcated Region (northeastern Portugal) to evaluate, one, two, and seven years after establishment, differences associated with two sown cover crop mixtures, namely a sown legume mixture (LEG) and a sown mixture of legumes and grasses (LEGRA) and spontaneous vegetation (SPONT) in vegetation dynamics and the activity density, richness, diversity, and stability of soil surface arthropod communities. Vegetation was assessed through floristic surveys, and arthropods were sampled using pitfall traps. Vegetation composition and soil surface arthropods community structure varied among sampling years, indicating that temporal variation also contributed to differences in both communities. LEG consistently maintained the highest vegetation cover, whereas LEGRA showed the lowest plant richness. Despite species turnover, LEG and LEGRA retained their functional identity, while SPONT exhibited greater temporal variation in floristic composition. Cover crop treatments were associated with differences in soil surface arthropod activity density, diversity, and community stability. LEGRA showed the highest activity density, whereas higher diversity and evenness were observed in LEG. SPONT presented intermediate values, differing significantly from LEGRA only in Shannon Wiener diversity index. All treatments maintained a relatively balanced trophic structure, suggesting a potential for natural regulation of herbivore population. Vegetation cover and plant richness were positively associated with soil surface arthropod diversity, highlighting the importance of vegetation characteristics for that communities in Mediterranean vineyards.<\/jats:p>","DOI":"10.3390\/insects17090975","type":"journal-article","created":{"date-parts":[[2026,9,22]],"date-time":"2026-09-22T17:56:41Z","timestamp":1790099801000},"page":"975","update-policy":"https:\/\/doi.org\/10.3390\/mdpi_crossmark_policy","source":"Crossref","is-referenced-by-count":0,"title":["Vegetation and Soil Surface Arthropods in Mediterranean Vineyards with Different Interrow Cover Crop Combinations"],"prefix":"10.3390","volume":"17","author":[{"given":"Jo\u00e3o","family":"Roque","sequence":"first","affiliation":[{"name":"Centre for the Research and Technology of Agro-Environmental and Biological Sciences (CITAB), University of Tr\u00e1s-os-Montes e Alto Douro (UTAD), 5000-801 Vila Real, Portugal"}],"role":[{"vocabulary":"crossref","role":"author"},{"role":"validation","vocabulary":"credit"},{"role":"writing-original-draft","vocabulary":"credit"}]},{"given":"Maria","family":"Ferreira","sequence":"additional","affiliation":[{"name":"Agronomy Department, University of Tr\u00e1s-os-Montes e Alto Douro (UTAD), 5000-801 Vila Real, Portugal"}],"role":[{"vocabulary":"crossref","role":"author"},{"role":"writing-original-draft","vocabulary":"credit"},{"role":"investigation","vocabulary":"credit"}]},{"ORCID":"https:\/\/orcid.org\/0000-0001-9236-9785","authenticated-orcid":false,"given":"Cristina","family":"Carlos","sequence":"additional","affiliation":[{"name":"Centre for the Research and Technology of Agro-Environmental and Biological Sciences (CITAB), University of Tr\u00e1s-os-Montes e Alto Douro (UTAD), 5000-801 Vila Real, Portugal"},{"name":"Agronomy Department, University of Tr\u00e1s-os-Montes e Alto Douro (UTAD), 5000-801 Vila Real, Portugal"}],"role":[{"vocabulary":"crossref","role":"author"},{"role":"conceptualization","vocabulary":"credit"},{"role":"methodology","vocabulary":"credit"},{"role":"writing-review-editing","vocabulary":"credit"},{"role":"funding-acquisition","vocabulary":"credit"}]},{"ORCID":"https:\/\/orcid.org\/0000-0002-9065-4336","authenticated-orcid":false,"given":"Irene","family":"Oliveira","sequence":"additional","affiliation":[{"name":"Centre for the Research and Technology of Agro-Environmental and Biological Sciences (CITAB), University of Tr\u00e1s-os-Montes e Alto Douro (UTAD), 5000-801 Vila Real, Portugal"},{"name":"Center for Computational and Stochastic Mathematics (CEMAT), University of Lisbon, 1049-001 Lisbon, Portugal"},{"name":"Department of Mathematics, University of Tr\u00e1s-os-Montes e Alto Douro (UTAD), 5000-801 Vila Real, Portugal"}],"role":[{"vocabulary":"crossref","role":"author"},{"role":"formal-analysis","vocabulary":"credit"},{"role":"visualization","vocabulary":"credit"}]},{"given":"Ant\u00f3nio","family":"Cresp\u00ed","sequence":"additional","affiliation":[{"name":"Centre for the Research and Technology of Agro-Environmental and Biological Sciences (CITAB), University of Tr\u00e1s-os-Montes e Alto Douro (UTAD), 5000-801 Vila Real, Portugal"}],"role":[{"vocabulary":"crossref","role":"author"},{"role":"methodology","vocabulary":"credit"},{"role":"investigation","vocabulary":"credit"},{"role":"writing-review-editing","vocabulary":"credit"}]},{"ORCID":"https:\/\/orcid.org\/0000-0001-7770-4477","authenticated-orcid":false,"given":"F\u00e1tima","family":"Gon\u00e7alves","sequence":"additional","affiliation":[{"name":"Centre for the Research and Technology of Agro-Environmental and Biological Sciences (CITAB), University of Tr\u00e1s-os-Montes e Alto Douro (UTAD), 5000-801 Vila Real, Portugal"},{"name":"Agronomy Department, University of Tr\u00e1s-os-Montes e Alto Douro (UTAD), 5000-801 Vila Real, Portugal"}],"role":[{"vocabulary":"crossref","role":"author"},{"role":"conceptualization","vocabulary":"credit"},{"role":"methodology","vocabulary":"credit"},{"role":"validation","vocabulary":"credit"},{"role":"investigation","vocabulary":"credit"},{"role":"data-curation","vocabulary":"credit"},{"role":"writing-review-editing","vocabulary":"credit"},{"role":"funding-acquisition","vocabulary":"credit"},{"role":"supervision","vocabulary":"credit"}]}],"member":"1968","published-online":{"date-parts":[[2026,9,21]]},"reference":[{"key":"ref_1","doi-asserted-by":"crossref","first-page":"685","DOI":"10.1098\/rstb.2007.2178","article-title":"Soil health in agricultural systems","volume":"363","author":"Kibblewhite","year":"2008","journal-title":"Philos. Trans. R. Soc. B Biol. Sci."},{"key":"ref_2","doi-asserted-by":"crossref","unstructured":"Lal, R. (2004). Carbon Sequestration in Dryland Agriculture. Challenges and Strategies of Dryland Agriculture, CSSA Special Publications.","DOI":"10.2135\/cssaspecpub32.c20"},{"key":"ref_3","unstructured":"OIV (2026). State of the World Wine Sector in 2025, International Organisation of Vine and Wine."},{"key":"ref_4","doi-asserted-by":"crossref","first-page":"128029","DOI":"10.1016\/j.jenvman.2025.128029","article-title":"Extensive vineyard management and semi-natural habitats increase biodiversity and ecosystem services: Insights from a global meta-analysis","volume":"395","author":"Winter","year":"2025","journal-title":"J. Environ. Manag."},{"key":"ref_5","doi-asserted-by":"crossref","first-page":"2484","DOI":"10.1111\/1365-2664.13124","article-title":"Effects of vegetation management intensity on biodiversity and ecosystem services in vineyards: A meta-analysis","volume":"55","author":"Winter","year":"2018","journal-title":"J. Appl. Ecol."},{"key":"ref_6","doi-asserted-by":"crossref","first-page":"1","DOI":"10.1016\/j.catena.2016.02.010","article-title":"Soil water erosion on Mediterranean vineyards: A review","volume":"141","author":"Prosdocimi","year":"2016","journal-title":"CATENA"},{"key":"ref_7","doi-asserted-by":"crossref","first-page":"110","DOI":"10.1016\/j.scitotenv.2014.03.069","article-title":"Soil losses in rural watersheds with environmental land use conflicts","volume":"485\u2013486","author":"Pacheco","year":"2014","journal-title":"Sci. Total Environ."},{"key":"ref_8","doi-asserted-by":"crossref","first-page":"29","DOI":"10.20870\/oeno-one.2023.57.2.7136","article-title":"Cover crops and no-tillage show negligible effects on grapevine physiology in mediterranean vineyard agroecosystems","volume":"57","author":"Zumkeller","year":"2023","journal-title":"OENO One"},{"key":"ref_9","doi-asserted-by":"crossref","unstructured":"Martins, S., Brito, C., Silva, E., Gon\u00e7alves, A., Arrobas, M., Pereira, E., Rodrigues, M.\u00c2., Nunes, F.M., and Correia, C.M. (2023). Synergy between Zeolites and Leguminous Cover Crops Improved Olive Tree Performance and Soil Properties in a Rainfed Olive Orchard. Agronomy, 13.","DOI":"10.3390\/agronomy13112674"},{"key":"ref_10","doi-asserted-by":"crossref","unstructured":"Brito, C., Martins, S., Saraiva, F., Roque, J., Arrobas, M., Rodrigues, M.\u00c2., and Correia, C. (2025). Biomass and nutrient release dynamics of decomposing rolling down legume cover crops in a steep-sloped Mediterranean vineyard. Front. Agron., 7.","DOI":"10.3389\/fagro.2025.1604142"},{"key":"ref_11","doi-asserted-by":"crossref","unstructured":"Panagos, P., Ballabio, C., Poesen, J., Lugato, E., Scarpa, S., Montanarella, L., and Borrelli, P. (2020). A Soil Erosion Indicator for Supporting Agricultural, Environmental and Climate Policies in the European Union. Remote Sens., 12.","DOI":"10.3390\/rs12091365"},{"key":"ref_12","doi-asserted-by":"crossref","unstructured":"Tiwari, H., Canavera, G., Pelusi, F., and Poni, S. (2025). Shifting from Tillage to Cover Cropping in Warm Climate Viticulture: Seeking the Optimal Balance. Agronomy, 15.","DOI":"10.3390\/agronomy15102245"},{"key":"ref_13","doi-asserted-by":"crossref","unstructured":"Zalai, M., Bujt\u00e1s, O., S\u00e1rospataki, M., and Dorner, Z. (2025). Grassy and Herbaceous Interrow Cover Crops in European Vineyards: A Review of Their Short-Term Effects on Water Management and Regulating Ecosystem Services. Land, 14.","DOI":"10.3390\/land14081526"},{"key":"ref_14","doi-asserted-by":"crossref","first-page":"1","DOI":"10.1890\/ES12-00236.1","article-title":"Spatiotemporal variation of event related N2O and CH4 emissions during fertigation in a California almond orchard","volume":"4","author":"Alsina","year":"2013","journal-title":"Ecosphere"},{"key":"ref_15","doi-asserted-by":"crossref","first-page":"108972","DOI":"10.1016\/j.agee.2024.108972","article-title":"A native plant species cover crop positively impacted vineyard water dynamics, soil health, and vine vigor","volume":"367","author":"Fernando","year":"2024","journal-title":"Agric. Ecosyst. Environ."},{"key":"ref_16","doi-asserted-by":"crossref","first-page":"109384","DOI":"10.1016\/j.agee.2024.109384","article-title":"Exploring the influence of cover crops with native plant species on soil and berry microbiota in a Moscatel Galego vineyard: Implications for sustainable viticulture","volume":"380","author":"Teixeira","year":"2025","journal-title":"Agric. Ecosyst. Environ."},{"key":"ref_17","doi-asserted-by":"crossref","unstructured":"Wall, D.H., and Bardgett, R.D. (2012). Soil Ecology and Ecosystem Services, Oxford University Press.","DOI":"10.1093\/acprof:oso\/9780199575923.001.0001"},{"key":"ref_18","doi-asserted-by":"crossref","first-page":"233","DOI":"10.1016\/j.agee.2006.12.013","article-title":"Soil biodiversity for agricultural sustainability","volume":"121","author":"Brussaard","year":"2007","journal-title":"Agric. Ecosyst. Environ."},{"key":"ref_19","doi-asserted-by":"crossref","first-page":"287","DOI":"10.1111\/j.1466-8238.2009.00517.x","article-title":"Macroecological patterns in soil communities","volume":"19","year":"2010","journal-title":"Glob. Ecol. Biogeogr."},{"key":"ref_20","doi-asserted-by":"crossref","first-page":"629","DOI":"10.3390\/agriculture3040629","article-title":"Role of Arthropods in Maintaining Soil Fertility","volume":"3","author":"Culliney","year":"2013","journal-title":"Agriculture"},{"key":"ref_21","doi-asserted-by":"crossref","first-page":"19","DOI":"10.1016\/S0167-8809(99)00028-6","article-title":"The ecological role of biodiversity in agroecosystems","volume":"74","author":"Altieri","year":"1999","journal-title":"Agric. Ecosyst. Environ."},{"key":"ref_22","doi-asserted-by":"crossref","first-page":"106863","DOI":"10.1016\/j.agee.2020.106863","article-title":"Do soil management practices affect the activity density, diversity, and stability of soil arthropods in vineyards?","volume":"294","author":"Nunes","year":"2020","journal-title":"Agric. Ecosyst. Environ."},{"key":"ref_23","unstructured":"Majer, J. (1987). The conservation and study of invertebrates in remnants of native vegetation. Nature Conservation: The Role of Remnants of Native Vegetation, Surrey Beatty And Sons Pty Limited."},{"key":"ref_24","doi-asserted-by":"crossref","first-page":"106571","DOI":"10.1016\/j.agee.2019.106571","article-title":"Ground cover management in a Mediterranean vineyard: Impact on insect abundance and diversity","volume":"283","year":"2019","journal-title":"Agric. Ecosyst. Environ."},{"key":"ref_25","doi-asserted-by":"crossref","first-page":"1","DOI":"10.1016\/j.agee.2018.11.004","article-title":"Organic cropping practice decreases pest abundance and positively influences predator-prey interactions","volume":"272","author":"Jacobsen","year":"2019","journal-title":"Agric. Ecosyst. Environ."},{"key":"ref_26","doi-asserted-by":"crossref","first-page":"70","DOI":"10.1016\/j.agee.2018.12.004","article-title":"Species and functional diversity of arthropod assemblages (Araneae, Carabidae, Heteroptera and Orthoptera) in grazed and mown salt grasslands","volume":"273","author":"Torma","year":"2019","journal-title":"Agric. Ecosyst. Environ."},{"key":"ref_27","doi-asserted-by":"crossref","first-page":"625","DOI":"10.1017\/S0007485317001158","article-title":"Organic vs. organic\u2014Soil arthropods as bioindicators of ecological sustainability in greenhouse system experiment under Mediterranean conditions","volume":"108","author":"Madzaric","year":"2017","journal-title":"Bull. Entomol. Res."},{"key":"ref_28","doi-asserted-by":"crossref","first-page":"158","DOI":"10.1016\/j.soilbio.2011.11.014","article-title":"A soil fauna index for assessing soil quality","volume":"47","author":"Yan","year":"2012","journal-title":"Soil Biol. Biochem."},{"key":"ref_29","doi-asserted-by":"crossref","first-page":"3680","DOI":"10.1038\/s41598-022-07529-1","article-title":"Vegetation increases abundances of ground and canopy arthropods in Mediterranean vineyards","volume":"12","author":"Blaise","year":"2022","journal-title":"Sci. Rep."},{"key":"ref_30","doi-asserted-by":"crossref","first-page":"759","DOI":"10.1017\/S0007485316000493","article-title":"Habitat management of organic vineyard in Northern Italy: The role of cover plants management on arthropod functional biodiversity","volume":"106","author":"Burgio","year":"2016","journal-title":"Bull. Entomol. Res."},{"key":"ref_31","doi-asserted-by":"crossref","first-page":"970","DOI":"10.1603\/EN09144","article-title":"Effects of native grass cover crops on beneficial and pest invertebrates in Australian vineyards","volume":"39","author":"Danne","year":"2010","journal-title":"Environ. Entomol."},{"key":"ref_32","doi-asserted-by":"crossref","first-page":"426","DOI":"10.1111\/ajgw.12290","article-title":"Cover crop differentially affects arthropods, but not diseases, occurring on grape leaves in vineyards","volume":"23","author":"Vogelweith","year":"2017","journal-title":"Aust. J. Grape Wine Res."},{"key":"ref_33","doi-asserted-by":"crossref","first-page":"388","DOI":"10.1111\/1365-2664.14833","article-title":"Weed management modifies functional properties of both weeds and microbial nitrogen-cycling communities in Mediterranean vineyards","volume":"62","author":"Bopp","year":"2025","journal-title":"J. Appl. Ecol."},{"key":"ref_34","doi-asserted-by":"crossref","first-page":"102706","DOI":"10.1016\/j.agsy.2019.102706","article-title":"Vegetation management intensity and landscape diversity alter plant species richness, functional traits and community composition across European vineyards","volume":"177","author":"Hall","year":"2020","journal-title":"Agric. Syst."},{"key":"ref_35","doi-asserted-by":"crossref","first-page":"296","DOI":"10.1111\/ajgw.12278","article-title":"Daily prediction of seasonal grapevine production in the Douro wine region based on favourable meteorological conditions","volume":"23","author":"Fraga","year":"2017","journal-title":"Aust. J. Grape Wine Res."},{"key":"ref_36","doi-asserted-by":"crossref","first-page":"574","DOI":"10.1080\/09583157.2014.881981","article-title":"Flowers for better pest control? Ground cover plants enhance apple orchard spiders (Araneae), but not necessarily their impact on pests","volume":"24","author":"Keresztes","year":"2014","journal-title":"Biocontrol Sci. Technol."},{"key":"ref_37","doi-asserted-by":"crossref","first-page":"499","DOI":"10.1017\/S0007485399000656","article-title":"The within-field spatial and temporal distribution of arthropods in winter wheat","volume":"89","author":"Holland","year":"2007","journal-title":"Bull. Entomol. Res."},{"key":"ref_38","doi-asserted-by":"crossref","first-page":"559","DOI":"10.1146\/annurev-ento-011613-161952","article-title":"Movement of entomophagous arthropods in agricultural landscapes: Links to pest suppression","volume":"59","author":"Schellhorn","year":"2014","journal-title":"Annu. Rev. Entomol."},{"key":"ref_39","doi-asserted-by":"crossref","first-page":"51","DOI":"10.1016\/j.agee.2005.08.033","article-title":"Carabid movement and vegetation density: Implications for interpreting pitfall trap data from split-field trials","volume":"113","author":"Thomas","year":"2006","journal-title":"Agric. Ecosyst. Environ."},{"key":"ref_40","first-page":"387","article-title":"Phytostructural characterization of several vegetation types in northern Portugal. II. The structural expressivity and the resistance of the vegetation","volume":"139","author":"Bernardos","year":"2005","journal-title":"Plant Biosyst.\u2014Int. J. Deal. All Asp. Plant Biol."},{"key":"ref_41","unstructured":"Southwood, T., and Henderson, P. (2000). Ecological Methods, Blackwell Science. [3rd ed.]."},{"key":"ref_42","doi-asserted-by":"crossref","first-page":"3953","DOI":"10.1002\/ece3.2176","article-title":"A review of extensive variation in the design of pitfall traps and a proposal for a standard pitfall trap design for monitoring ground-active arthropod biodiversity","volume":"6","author":"Brown","year":"2016","journal-title":"Ecol. Evol."},{"key":"ref_43","doi-asserted-by":"crossref","first-page":"2201","DOI":"10.1007\/s10531-010-9832-y","article-title":"Rapid biodiversity assessment of arthropods for monitoring average local species richness and related ecosystem services","volume":"19","author":"Obrist","year":"2010","journal-title":"Biodivers. Conserv."},{"key":"ref_44","doi-asserted-by":"crossref","first-page":"562","DOI":"10.1046\/j.1523-1739.1993.07030562.x","article-title":"A Possible Method for the Rapid Assessment of Biodiversity","volume":"7","author":"Oliver","year":"1993","journal-title":"Conserv. Biol."},{"key":"ref_45","unstructured":"Aguiar, C., and Serrano, A. (2013). Cat\u00e1logo e Atlas dos Cole\u00f3pteros Carab\u00eddeos (Coleoptera: Carabidae) de Portugal Continental, Sociedade Portuguesa de Entomologia."},{"key":"ref_46","unstructured":"Booth, R.G., Cox, M.L., and Madge, R.B. (1990). Coleoptera. IIE Guides to Insects of Importance to Man, CAB International."},{"key":"ref_47","first-page":"49","article-title":"A guide to ants of Continental Portugal (Hymenoptera: Formicidae)","volume":"5","author":"Collingwood","year":"1998","journal-title":"Bol. DA Soc. Port. DE Entomol."},{"key":"ref_48","unstructured":"G\u00f3mez, K., and Espadaler, X. (2026, July 31). Hormigas Ib\u00e9ricas. Available online: https:\/\/hormigas.mirmiberica.org\/."},{"key":"ref_49","doi-asserted-by":"crossref","first-page":"458","DOI":"10.1007\/s00442-004-1740-y","article-title":"The response of carabids to landscape simplification differs between trophic groups","volume":"142","author":"Purtauf","year":"2005","journal-title":"Oecologia"},{"key":"ref_50","unstructured":"Triplehorn, C.A., Johnson, N.F., and Borror, D.J. (2005). Borror and DeLong\u2019s Introduction to the Study of Insects, Thompson Brooks\/Cole."},{"key":"ref_51","doi-asserted-by":"crossref","first-page":"226","DOI":"10.1111\/j.1365-2311.2010.01175.x","article-title":"Trophic level modulates carabid beetle responses to habitat and landscape structure: A pan-European study","volume":"35","author":"Vanbergen","year":"2010","journal-title":"Ecol. Entomol."},{"key":"ref_52","unstructured":"McCune, B., and Grace, J. (2002). Analysis of Ecological Communities, MJM Software Design."},{"key":"ref_53","unstructured":"Magurran, A.E. (2004). Measuring Biological Diversity, Blackwell Publishing."},{"key":"ref_54","doi-asserted-by":"crossref","unstructured":"Gotelli, N., and Chao, A. (2013). Measuring and Estimating Species Richness, Species Diversity, and Biotic Similarity from Sampling Data, Academic press.","DOI":"10.1016\/B978-0-12-384719-5.00424-X"},{"key":"ref_55","doi-asserted-by":"crossref","first-page":"363","DOI":"10.1111\/j.2006.0030-1299.14714.x","article-title":"Entropy and diversity","volume":"113","author":"Jost","year":"2006","journal-title":"Oikos"},{"key":"ref_56","doi-asserted-by":"crossref","first-page":"175","DOI":"10.1016\/j.ecoleng.2014.05.010","article-title":"Ecological engineering of ground cover vegetation enhances the diversity and stability of peach orchard canopy arthropod communities","volume":"70","author":"Wan","year":"2014","journal-title":"Ecol. Eng."},{"key":"ref_57","unstructured":"Team, R.C. (2025). R: A Language and Environment for Statistical Computing, R Core Team."},{"key":"ref_58","first-page":"345","article-title":"Species assemblages and indicator species:the need for a flexible asymmetrical approach","volume":"67","author":"Legendre","year":"1997","journal-title":"Ecol. Monogr."},{"key":"ref_59","doi-asserted-by":"crossref","first-page":"11","DOI":"10.5334\/jors.452","article-title":"Ecoindicators: An R Package for the Identification of Indicator Taxonomic Units","volume":"11","author":"Duhour","year":"2023","journal-title":"J. Open Res. Softw."},{"key":"ref_60","unstructured":"Juggins, S. (Rioja: Analysis of Quaternary Science Data, 2024). Rioja: Analysis of Quaternary Science Data, R package version 1.0-7."},{"key":"ref_61","unstructured":"Oksanen, J., Simpson, G., Blanchet, F., Kindt, R., Legendre, P., Minchin, P., O\u2019Hara, R., Solymos, P., Stevens, M., and Szoecs, E. (vegan: Community Ecology Package, 2026). vegan: Community Ecology Package, R package version 2.7-5."},{"key":"ref_62","doi-asserted-by":"crossref","unstructured":"Wickham, H. (2009). ggplot2: Elegant Graphics for Data Analysis, Springer.","DOI":"10.1007\/978-0-387-98141-3"},{"key":"ref_63","doi-asserted-by":"crossref","first-page":"1","DOI":"10.18637\/jss.v067.i01","article-title":"Fitting Linear Mixed-Effects Models Using lme4","volume":"67","author":"Bates","year":"2015","journal-title":"J. Stat. Softw."},{"key":"ref_64","doi-asserted-by":"crossref","first-page":"1686","DOI":"10.21105\/joss.01686","article-title":"Welcome to the Tidyverse","volume":"4","author":"Wickham","year":"2019","journal-title":"J. Open Source Softw."},{"key":"ref_65","unstructured":"van den Brand, T. (ggh4x: Hacks for \u2018ggplot2\u2019, 2025). ggh4x: Hacks for \u2018ggplot2\u2019, R package version 0.3.1."},{"key":"ref_66","doi-asserted-by":"crossref","first-page":"295","DOI":"10.20870\/oeno-one.2021.55.1.3599","article-title":"Cover crops in viticulture. A systematic review (1): Implications on soil characteristics and biodiversity in vineyard","volume":"55","author":"Abad","year":"2021","journal-title":"OENO One"},{"key":"ref_67","doi-asserted-by":"crossref","first-page":"158","DOI":"10.1016\/j.agee.2017.09.030","article-title":"Management of service crops for the provision of ecosystem services in vineyards: A review","volume":"251","author":"Garcia","year":"2018","journal-title":"Agric. Ecosyst. Environ."},{"key":"ref_68","doi-asserted-by":"crossref","first-page":"108740","DOI":"10.1016\/j.agee.2023.108740","article-title":"Effect of spontaneous vegetation on beneficial arthropods in Mediterranean vineyards","volume":"359","author":"Rocher","year":"2024","journal-title":"Agric. Ecosyst. Environ."},{"key":"ref_69","doi-asserted-by":"crossref","first-page":"452","DOI":"10.1016\/j.fcr.2016.08.005","article-title":"Competition between cocksfoot (Dactylis glomerata L.) and companion species: Evidence for allelopathy","volume":"196","author":"Li","year":"2016","journal-title":"Field Crops Res."},{"key":"ref_70","unstructured":"Espejo-P\u00e9rez, A., Rodr\u00edguez-Lizana, A., Gir\u00e1ldez, J., and Ord\u00f3\u00f1ez, R. (2005, January 9\u201311). Efecto de la cubierta vegetal en la p\u00e9rdida de suelo en olivar ecol\u00f3gico. Proceedings of the Congreso Internacional Sobre Agricultura de Conservaci\u00f3n, Espa\u00f1a, Spain."},{"key":"ref_71","doi-asserted-by":"crossref","first-page":"2449","DOI":"10.2134\/agronj15.0086","article-title":"Cover Crops and Ecosystem Services: Insights from Studies in Temperate Soils","volume":"107","author":"Shaver","year":"2015","journal-title":"Agron. J."},{"key":"ref_72","doi-asserted-by":"crossref","first-page":"65","DOI":"10.1046\/j.1365-2664.2003.00762.x","article-title":"Plant traits as predictors of performance in ecological restoration","volume":"40","author":"Pywell","year":"2003","journal-title":"J. Appl. Ecol."},{"key":"ref_73","doi-asserted-by":"crossref","unstructured":"Smith, R.G., Atwood, L.W., and Warren, N.D. (2014). Increased Productivity of a Cover Crop Mixture Is Not Associated with Enhanced Agroecosystem Services. PLoS ONE, 9.","DOI":"10.1371\/journal.pone.0097351"},{"key":"ref_74","doi-asserted-by":"crossref","first-page":"17445","DOI":"10.1038\/s41598-017-17601-w","article-title":"Soil biota in vineyards are more influenced by plants and soil quality than by tillage intensity or the surrounding landscape","volume":"7","author":"Buchholz","year":"2017","journal-title":"Sci. Rep."},{"key":"ref_75","doi-asserted-by":"crossref","first-page":"1653","DOI":"10.2478\/s11756-019-00272-5","article-title":"Cover crop farming system affects macroarthropods community diversity in Regosol of Caatinga, Brazil","volume":"74","author":"Santos","year":"2019","journal-title":"Biologia"},{"key":"ref_76","doi-asserted-by":"crossref","unstructured":"Heimpel, G., and Jervis, M. (2005). Does floral nectar improve biological control by parasitoids?. Plant-Provided Food for Carnivorous Insects: A Protective Mutualism and Its Applications, Cambridge University Press.","DOI":"10.1017\/CBO9780511542220.010"},{"key":"ref_77","doi-asserted-by":"crossref","first-page":"91","DOI":"10.1016\/j.apsoil.2014.10.008","article-title":"Contribution of legume tree residues and macrofauna to the improvement of abiotic soil properties in the eastern Amazon","volume":"86","author":"Moura","year":"2015","journal-title":"Appl. Soil Ecol."},{"key":"ref_78","doi-asserted-by":"crossref","first-page":"140","DOI":"10.1016\/j.still.2007.09.006","article-title":"Impact of a no-till with mulch soil management strategy on soil macrofauna communities in a cotton cropping system","volume":"97","author":"Bikay","year":"2007","journal-title":"Soil Tillage Res."},{"key":"ref_79","first-page":"473","article-title":"Conservation agriculture and soil fauna: Only benefits or also potential threats? A review","volume":"2","author":"Conti","year":"2015","journal-title":"ECronicon Agric."},{"key":"ref_80","doi-asserted-by":"crossref","first-page":"33","DOI":"10.1016\/j.still.2010.08.010","article-title":"Tillage and soil ecology: Partners for sustainable agriculture","volume":"111","author":"Anger","year":"2010","journal-title":"Soil Tillage Res."},{"key":"ref_81","doi-asserted-by":"crossref","first-page":"109842","DOI":"10.1016\/j.agee.2025.109842","article-title":"Unravelling the effects of vineyard inter-row vegetation on soil biodiversity in South Africa","volume":"393","author":"Mamabolo","year":"2025","journal-title":"Agric. Ecosyst. Environ."},{"key":"ref_82","doi-asserted-by":"crossref","unstructured":"Gon\u00e7alves, F., Carlos, C., Crespo, L., Zina, V., Oliveira, A., Salva\u00e7\u00e3o, J., Pereira, J.A., and Torres, L. (2021). Soil Arthropods in the Douro Demarcated Region Vineyards: General Characteristics and Ecosystem Services Provided. Sustainability, 13.","DOI":"10.3390\/su13147837"},{"key":"ref_83","doi-asserted-by":"crossref","first-page":"45","DOI":"10.1890\/13-0133.1","article-title":"Rarefaction and extrapolation with Hill numbers: A framework for sampling and estimation in species diversity studies","volume":"84","author":"Chao","year":"2014","journal-title":"Ecol. Monogr."},{"key":"ref_84","unstructured":"ADVID (2017). Boletim Ano Vit\u00edcola 2017. Balanc\u0327o Final do Ano Vit\u00edcola, ADVID\u2014Associac\u0327\u00e3o para o Desenvolvimento de Viticultura Duriense."},{"key":"ref_85","unstructured":"ADVID (2018). Boletim Ano Vit\u00edcola 2018. Balanc\u0327o Final do Ano Vit\u00edcola, ADVID\u2014Associac\u0327\u00e3o para o Desenvolvimento de Viticultura Duriense."},{"key":"ref_86","unstructured":"ADVID (2023). Boletim Ano Vit\u00edcola 2023. Balanc\u0327o Final do Ano Vit\u00edcola, ADVID\u2014Associac\u0327\u00e3o para o Desenvolvimento e Viticultura Duriense."},{"key":"ref_87","doi-asserted-by":"crossref","unstructured":"Santoro, F.R., Ferreira J\u00fanior, W.S., Ara\u00fajo, T.A.d.S., Ladio, A.H., and Albuquerque, U.P. (2015). Does Plant Species Richness Guarantee the Resilience of Local Medical Systems? A Perspective from Utilitarian Redundancy. PLoS ONE, 10.","DOI":"10.1371\/journal.pone.0119826"},{"key":"ref_88","doi-asserted-by":"crossref","first-page":"18","DOI":"10.1046\/j.1523-1739.1992.610018.x","article-title":"Biodiversity and Ecological Redundancy","volume":"6","author":"Walker","year":"1992","journal-title":"Conserv. Biol."},{"key":"ref_89","doi-asserted-by":"crossref","first-page":"20142620","DOI":"10.1098\/rspb.2014.2620","article-title":"Functional identity and diversity of animals predict ecosystem functioning better than species-based indices","volume":"282","author":"Gagic","year":"2015","journal-title":"Proc. R. Soc. B Biol. Sci."},{"key":"ref_90","doi-asserted-by":"crossref","first-page":"857","DOI":"10.1111\/j.1461-0248.2005.00782.x","article-title":"Landscape perspectives on agricultural intensification and biodiversity\u2014Ecosystem service management","volume":"8","author":"Tscharntke","year":"2005","journal-title":"Ecol. Lett."},{"key":"ref_91","first-page":"8089","article-title":"Regenerative viticulture and climate change resilience","volume":"59","author":"Nesbitt","year":"2025","journal-title":"OENO One"},{"key":"ref_92","doi-asserted-by":"crossref","first-page":"70","DOI":"10.1007\/s13593-025-01074-7","article-title":"Cover crops promote regulating and supporting ecosystem services without compromising grape yield in temperate vineyards. A meta-analysis","volume":"45","author":"Calderone","year":"2025","journal-title":"Agron. Sustain. 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