{"status":"ok","message-type":"work","message-version":"1.0.0","message":{"indexed":{"date-parts":[[2026,10,1]],"date-time":"2026-10-01T06:49:47Z","timestamp":1790837387512,"version":"4.1.0"},"reference-count":193,"publisher":"MDPI AG","issue":"4","license":[{"start":{"date-parts":[[2018,4,10]],"date-time":"2018-04-10T00:00:00Z","timestamp":1523318400000},"content-version":"vor","delay-in-days":0,"URL":"https:\/\/creativecommons.org\/licenses\/by\/4.0\/"}],"content-domain":{"domain":[],"crossmark-restriction":false},"short-container-title":["Molecules"],"abstract":"<jats:p>Chitosan is a naturally occurring compound and is commercially produced from seafood shells. It has been utilized in the induction of the defense system in both pre and post-harvest fruits and vegetables against fungi, bacteria, viruses, and other abiotic stresses. In addition to that, chitosan effectively improves the physiological properties of plants and also enhances the shelf life of post-harvest produces. Moreover, chitosan treatment regulates several genes in plants, particularly the activation of plant defense signaling pathways. That includes the elicitation of phytoalexins and pathogenesis-related (PR) protein. Besides that, chitosan has been employed in soil as a plant nutrient and has shown great efficacy in combination with other industrial fertilizers without affecting the soil\u2019s beneficial microbes. Furthermore, it is helpful in reducing the fertilizer losses due to its coating ability, which is important in keeping the environmental pollution under check. Based on exhibiting such excellent properties, there is a striking interest in using chitosan biopolymers in agriculture systems. Therefore, our current review has been centered upon the multiple roles of chitosan in horticultural crops that could be useful in future crop improvement programs.<\/jats:p>","DOI":"10.3390\/molecules23040872","type":"journal-article","created":{"date-parts":[[2018,4,10]],"date-time":"2018-04-10T13:06:08Z","timestamp":1523365568000},"page":"872","update-policy":"https:\/\/doi.org\/10.3390\/mdpi_crossmark_policy","source":"Crossref","is-referenced-by-count":238,"title":["The Multifunctional Role of Chitosan in Horticultural Crops; A Review"],"prefix":"10.3390","volume":"23","author":[{"given":"Rahat","family":"Sharif","sequence":"first","affiliation":[{"name":"College of Horticulture, Northwest A&F University, Yangling 712100, China"}],"role":[{"vocabulary":"crossref","role":"author"}]},{"ORCID":"https:\/\/orcid.org\/0000-0001-8392-9226","authenticated-orcid":false,"given":"Muhammad","family":"Mujtaba","sequence":"additional","affiliation":[{"name":"Institute of Biotechnology, Ankara University, Ankara 06110, Turkey"}],"role":[{"vocabulary":"crossref","role":"author"}]},{"given":"Mati","family":"Ur Rahman","sequence":"additional","affiliation":[{"name":"State Key Laboratory of Crop Stress Biology in Arid Areas, College of Horticulture, Northwest A&amp;F University, Yangling 712100, China"}],"role":[{"vocabulary":"crossref","role":"author"}]},{"given":"Abdullah","family":"Shalmani","sequence":"additional","affiliation":[{"name":"State Key Laboratory of Crop Stress Biology in Arid Areas, College of Life sciences, Northwest A&amp;F University, Yangling 712100, China"}],"role":[{"vocabulary":"crossref","role":"author"}]},{"ORCID":"https:\/\/orcid.org\/0000-0003-0955-3683","authenticated-orcid":false,"given":"Husain","family":"Ahmad","sequence":"additional","affiliation":[{"name":"College of Horticulture, Northwest A&F University, Yangling 712100, China"}],"role":[{"vocabulary":"crossref","role":"author"}]},{"given":"Toheed","family":"Anwar","sequence":"additional","affiliation":[{"name":"Hubei Collaborative Innovation Center for Grain Industry\/Research Center of Crop Stresses Resistance Technologies, Yangtze University, Jingzhou 434025, China"}],"role":[{"vocabulary":"crossref","role":"author"}]},{"given":"Deng","family":"Tianchan","sequence":"additional","affiliation":[{"name":"School of Mechanical Aerospace and Civil Engineering, The University of Manchester, Manchester M13 9PL, UK"}],"role":[{"vocabulary":"crossref","role":"author"}]},{"given":"Xiping","family":"Wang","sequence":"additional","affiliation":[{"name":"State Key Laboratory of Crop Stress Biology in Arid Areas, College of Horticulture, Northwest A&amp;F University, Yangling 712100, China"},{"name":"Key Laboratory of Horticultural Plant Biology and Germplasm Innovation in Northwest China, Ministry of Agriculture, Northwest A&F University, Yangling 712100, China"}],"role":[{"vocabulary":"crossref","role":"author"}]}],"member":"1968","published-online":{"date-parts":[[2018,4,10]]},"reference":[{"key":"ref_1","doi-asserted-by":"crossref","first-page":"981","DOI":"10.1016\/j.progpolymsci.2011.02.001","article-title":"Chitosan\u2014A versatile semi-synthetic polymer in biomedical applications","volume":"36","author":"Dash","year":"2011","journal-title":"Prog. Polym. Sci."},{"key":"ref_2","doi-asserted-by":"crossref","first-page":"46","DOI":"10.1016\/j.ijbiomac.2013.04.043","article-title":"Chitosan-based nanomaterials: A state-of-the-art review","volume":"59","author":"Shukla","year":"2013","journal-title":"Int. J. Biol. Macromol."},{"key":"ref_3","doi-asserted-by":"crossref","first-page":"307","DOI":"10.3390\/ma2020307","article-title":"Biodegradable polymers","volume":"2","author":"Vroman","year":"2009","journal-title":"Materials"},{"key":"ref_4","unstructured":"Muzzarelli, R.A. (1973). Natural chelating polymers; alginic acid, chitin and chitosan. Natural Chelating Polymers; Alginic Acid, Chitin and Chitosan, Pergamon Press."},{"key":"ref_5","doi-asserted-by":"crossref","first-page":"603","DOI":"10.1016\/j.progpolymsci.2006.06.001","article-title":"Chitin and chitosan: Properties and applications","volume":"31","author":"Rinaudo","year":"2006","journal-title":"Prog. Polym. Sci."},{"key":"ref_6","doi-asserted-by":"crossref","first-page":"9","DOI":"10.1016\/j.carbpol.2015.06.008","article-title":"Fluctuation in physicochemical properties of chitins extracted from different body parts of honeybee","volume":"132","author":"Kaya","year":"2015","journal-title":"Carbohydr. Polym."},{"key":"ref_7","doi-asserted-by":"crossref","first-page":"64","DOI":"10.1016\/j.carbpol.2016.03.010","article-title":"Changes in physicochemical properties of chitin at developmental stages (larvae, pupa and adult) of Vespa crabro (wasp)","volume":"145","author":"Kaya","year":"2016","journal-title":"Carbohydr. Polym."},{"key":"ref_8","doi-asserted-by":"crossref","first-page":"443","DOI":"10.1016\/j.ijbiomac.2015.08.021","article-title":"Surface morphology of chitin highly related with the isolated body part of butterfly (Argynnis pandora)","volume":"81","author":"Kaya","year":"2015","journal-title":"Int. J. Biol. Macromol."},{"key":"ref_9","doi-asserted-by":"crossref","first-page":"397","DOI":"10.1002\/pi.2378","article-title":"Main properties and current applications of some polysaccharides as biomaterials","volume":"57","author":"Rinaudo","year":"2008","journal-title":"Polym. Int."},{"key":"ref_10","doi-asserted-by":"crossref","first-page":"316","DOI":"10.1006\/jcis.2002.8248","article-title":"Interaction of carboxylic acids with chitosan: Effect of pK and hydrocarbon chain length","volume":"249","author":"Shamov","year":"2002","journal-title":"J. Colloid Interface Sci."},{"key":"ref_11","doi-asserted-by":"crossref","first-page":"637","DOI":"10.1208\/s12249-011-9620-3","article-title":"Effects of spray drying on physicochemical properties of chitosan acid salts","volume":"12","author":"Cervera","year":"2011","journal-title":"AAPS PharmSciTech"},{"key":"ref_12","doi-asserted-by":"crossref","first-page":"236","DOI":"10.1016\/j.jiec.2016.11.038","article-title":"Incorporation of sporopollenin enhances acid\u2013base durability, hydrophobicity, and mechanical, antifungal and antioxidant properties of chitosan films","volume":"47","author":"Kaya","year":"2017","journal-title":"J. Ind. Eng. Chem."},{"key":"ref_13","doi-asserted-by":"crossref","first-page":"1401","DOI":"10.1016\/j.ijbiomac.2017.08.161","article-title":"Diatomite as a novel composite ingredient for chitosan film with enhanced physicochemical properties","volume":"105","author":"Akyuz","year":"2017","journal-title":"Int. J. Biol. Macromol."},{"key":"ref_14","doi-asserted-by":"crossref","first-page":"944","DOI":"10.1016\/j.ijbiomac.2017.06.127","article-title":"Utilization of flax (Linum usitatissimum) cellulose nanocrystals as reinforcing material for chitosan films","volume":"104","author":"Mujtaba","year":"2017","journal-title":"Int. J. Biol. Macromol."},{"key":"ref_15","doi-asserted-by":"crossref","first-page":"285","DOI":"10.1016\/S0147-5975(79)80054-7","article-title":"The fungicidal effect of chitosan on fungi of varying cell wall composition","volume":"3","author":"Allan","year":"1979","journal-title":"Exp. Mycol."},{"key":"ref_16","doi-asserted-by":"crossref","first-page":"3","DOI":"10.1016\/0885-5765(89)90012-X","article-title":"Chitin oligosaccharides elicit lignification in wounded wheat leaves","volume":"34","author":"Barber","year":"1989","journal-title":"Physiol. Mol. Plant Pathol."},{"key":"ref_17","unstructured":"Chaouat, C. (2013). Conception de Nouveaux Syst\u00e8mes de Formulation d'actifs D\u00e9pigmentants, en vue de leur Utilisation par voie Cutan\u00e9e, Universit\u00e9 de Toulouse, Universit\u00e9 Toulouse III-Paul Sabatier."},{"key":"ref_18","doi-asserted-by":"crossref","first-page":"21","DOI":"10.1080\/03235409409383089","article-title":"Chitosan inhibits systemic infections caused by DNA-containing plant viruses","volume":"29","author":"Chirkov","year":"1994","journal-title":"Arch. Phytopathol. Plant Prot."},{"key":"ref_19","doi-asserted-by":"crossref","first-page":"1010","DOI":"10.1002\/jsfa.5812","article-title":"Effect of chitosan solution on the inhibition of Acidovorax citrulli causing bacterial fruit blotch of watermelon","volume":"93","author":"Li","year":"2013","journal-title":"J. Sci. Food Agric."},{"key":"ref_20","doi-asserted-by":"crossref","first-page":"7830182","DOI":"10.1155\/2016\/7830182","article-title":"Synergistic effects of plant growth promoting rhizobacteria and chitosan on in vitro seeds germination, greenhouse growth, and nutrient uptake of maize (Zea mays L.)","volume":"2016","author":"Agbodjato","year":"2016","journal-title":"Biotechnol. Res. Int."},{"key":"ref_21","doi-asserted-by":"crossref","first-page":"72","DOI":"10.1016\/j.msec.2014.09.004","article-title":"Physicochemical comparison of chitin and chitosan obtained from larvae and adult Colorado potato beetle (Leptinotarsa decemlineata)","volume":"45","author":"Kaya","year":"2014","journal-title":"Mater. Sci. Eng. C"},{"key":"ref_22","doi-asserted-by":"crossref","first-page":"58","DOI":"10.1007\/s12257-013-0488-9","article-title":"New chitin, chitosan, and O-carboxymethyl chitosan sources from resting eggs of Daphnia longispina (Crustacea); with physicochemical characterization, and antimicrobial and antioxidant activities","volume":"19","author":"Kaya","year":"2014","journal-title":"Biotechnol. Bioprocess Eng."},{"key":"ref_23","doi-asserted-by":"crossref","first-page":"6017","DOI":"10.1021\/cr030441b","article-title":"Chitosan chemistry and pharmaceutical perspectives","volume":"104","author":"Kumar","year":"2004","journal-title":"Chem. Rev."},{"key":"ref_24","doi-asserted-by":"crossref","first-page":"1457","DOI":"10.1021\/bm034130m","article-title":"Chitosan as antimicrobial agent: Applications and mode of action","volume":"4","author":"Rabea","year":"2003","journal-title":"Biomacromolecules"},{"key":"ref_25","doi-asserted-by":"crossref","first-page":"189","DOI":"10.1016\/j.chroma.2006.03.068","article-title":"High-performance liquid chromatographic determination of chitin in the snow crab, Chionoecetes opilio","volume":"1116","author":"Crespo","year":"2006","journal-title":"J. Chromatogr. A"},{"key":"ref_26","first-page":"72","article-title":"Extraction of chitin from trash crabs (Podophthalmus vigil) by an eccentric method","volume":"2","author":"Das","year":"2010","journal-title":"Curr. Res. Biol. Sci."},{"key":"ref_27","doi-asserted-by":"crossref","first-page":"583","DOI":"10.1016\/j.dci.2008.10.010","article-title":"Characterization of crustins from the hemocytes of the spider crab, Hyas araneus, and the red king crab, Paralithodes camtschaticus","volume":"33","author":"Sperstad","year":"2009","journal-title":"Dev. Comp. Immunol."},{"key":"ref_28","doi-asserted-by":"crossref","first-page":"298","DOI":"10.1016\/j.ijbiomac.2014.01.045","article-title":"Structural differences between chitin and chitosan extracted from three different marine sources","volume":"65","author":"Hajji","year":"2014","journal-title":"Int. J. Biol. Macromol."},{"key":"ref_29","doi-asserted-by":"crossref","first-page":"553","DOI":"10.1016\/j.ijbiomac.2014.02.010","article-title":"A physicochemical characterization of fully acetylated chitin structure isolated from two spider species: With new surface morphology","volume":"65","author":"Kaya","year":"2014","journal-title":"Int. J. Biol. Macromol."},{"key":"ref_30","doi-asserted-by":"crossref","unstructured":"Davies, G.J., Knight, D.P., and Vollrath, F. (2013). Chitin in the silk gland ducts of the spider Nephila edulis and the silkworm Bombyx mori. PLoS ONE, 8.","DOI":"10.1371\/journal.pone.0073225"},{"key":"ref_31","doi-asserted-by":"crossref","first-page":"605","DOI":"10.1016\/S0308-8146(97)00039-3","article-title":"Mycelia of Mucor rouxii as a source of chitin and chitosan","volume":"60","author":"Synowiecki","year":"1997","journal-title":"Food Chem."},{"key":"ref_32","doi-asserted-by":"crossref","first-page":"938","DOI":"10.1021\/ed067p938","article-title":"Chitin and chitosan, versatile polysaccharides from marine animals","volume":"67","author":"Mathur","year":"1990","journal-title":"J. Chem. Educ."},{"key":"ref_33","doi-asserted-by":"crossref","first-page":"204","DOI":"10.1002\/cben.201400025","article-title":"A review on chitin and chitosan polymers: Structure, chemistry, solubility, derivatives, and applications","volume":"2","author":"Zargar","year":"2015","journal-title":"ChemBioEng Rev."},{"key":"ref_34","first-page":"190","article-title":"Phytotherapeutic aspects of diseases of the circulatory system. 4. Chitin and chitosan","volume":"44","author":"Chobot","year":"1995","journal-title":"Ceska a Slovenska Farmacie: Casopis Ceske Farmaceuticke Spolecnosti a Slovenske farmaceuticke Spolecnosti"},{"key":"ref_35","doi-asserted-by":"crossref","first-page":"678","DOI":"10.1016\/j.jbiosc.2015.11.005","article-title":"Comparison of antimicrobial activities of newly obtained low molecular weight scorpion chitosan and medium molecular weight commercial chitosan","volume":"121","author":"Kaya","year":"2016","journal-title":"J. Biosci. Bioeng."},{"key":"ref_36","doi-asserted-by":"crossref","first-page":"1359","DOI":"10.1016\/j.biortech.2007.01.051","article-title":"Extraction and characterization of chitin and chitosan from local sources","volume":"99","author":"Abdou","year":"2008","journal-title":"Bioresour. Technol."},{"key":"ref_37","doi-asserted-by":"crossref","first-page":"414","DOI":"10.1016\/j.nimb.2007.09.015","article-title":"Gamma irradiation-aided chitin\/chitosan extraction from prawn shells","volume":"265","author":"Mahlous","year":"2007","journal-title":"Nucl. Instrum. Methods Phys. Res. Sec. B Beam Interact. Mater. At."},{"key":"ref_38","doi-asserted-by":"crossref","first-page":"99","DOI":"10.1016\/S0141-8130(99)00123-3","article-title":"Structure of insect chitin isolated from beetle larva cuticle and silkworm (Bombyx mori) pupa exuvia","volume":"27","author":"Zhang","year":"2000","journal-title":"Int. J. Biol. Macromol."},{"key":"ref_39","doi-asserted-by":"crossref","first-page":"4604","DOI":"10.3390\/molecules17044604","article-title":"Extraction and characterization of chitin from the beetle Holotrichia parallela motschulsky","volume":"17","author":"Liu","year":"2012","journal-title":"Molecules"},{"key":"ref_40","doi-asserted-by":"crossref","first-page":"7","DOI":"10.1016\/j.ijbiomac.2015.01.015","article-title":"Description of a new surface morphology for chitin extracted from wings of cockroach (Periplaneta americana)","volume":"75","author":"Kaya","year":"2015","journal-title":"Int. J. Biol. Macromol."},{"key":"ref_41","doi-asserted-by":"crossref","first-page":"145","DOI":"10.1007\/s11483-013-9327-y","article-title":"Extraction and characterization of \u03b1-chitin and chitosan from six different aquatic invertebrates","volume":"9","author":"Kaya","year":"2014","journal-title":"Food Biophys."},{"key":"ref_42","doi-asserted-by":"crossref","first-page":"119","DOI":"10.1295\/polymj.20.119","article-title":"Composite texture of chitin and keratin in an animal organ, Lingula seta","volume":"20","author":"Tanaka","year":"1988","journal-title":"Polym. J."},{"key":"ref_43","doi-asserted-by":"crossref","first-page":"559","DOI":"10.1021\/bm034401t","article-title":"New aspects of the extraction of chitin from squid pens","volume":"5","author":"Chaussard","year":"2004","journal-title":"Biomacromolecules"},{"key":"ref_44","doi-asserted-by":"crossref","first-page":"1919","DOI":"10.1021\/bm800178b","article-title":"Preparation of chitin nanofibers from squid pen \u03b2-chitin by simple mechanical treatment under acid conditions","volume":"9","author":"Fan","year":"2008","journal-title":"Biomacromolecules"},{"key":"ref_45","doi-asserted-by":"crossref","first-page":"93","DOI":"10.1016\/S1043-4526(05)49003-8","article-title":"Chitin, chitosan, and co-products: Chemistry, production, applications, and health effects","volume":"49","author":"Shahidi","year":"2005","journal-title":"Adv. Food Nutr. Res."},{"key":"ref_46","doi-asserted-by":"crossref","first-page":"3423","DOI":"10.1002\/pola.20176","article-title":"Physicochemical characterization of \u03b1-chitin, \u03b2-chitin, and \u03b3-chitin separated from natural resources","volume":"42","author":"Jang","year":"2004","journal-title":"J. Polym. Sci. Part A Polym. Chem."},{"key":"ref_47","doi-asserted-by":"crossref","first-page":"283","DOI":"10.1163\/1937240X-00002221","article-title":"Physicochemical characterization of chitin and chitosan obtained from resting eggs of Ceriodaphnia quadrangula (Branchiopoda: Cladocera: Daphniidae)","volume":"34","author":"Kaya","year":"2014","journal-title":"J. Crustacean Biol."},{"key":"ref_48","doi-asserted-by":"crossref","first-page":"300","DOI":"10.1023\/B:ABIM.0000025956.98250.30","article-title":"Hydrolysis of chitosan in lactic acid","volume":"40","author":"Varlamov","year":"2004","journal-title":"Appl. Biochem. Microbiol."},{"key":"ref_49","doi-asserted-by":"crossref","first-page":"969","DOI":"10.1021\/bp020011r","article-title":"Factors affecting the composition of oligosaccharides produced in chitosan hydrolysis using immobilized chitosanases","volume":"18","author":"Kuroiwa","year":"2002","journal-title":"Biotechnol. Prog."},{"key":"ref_50","doi-asserted-by":"crossref","first-page":"45","DOI":"10.1016\/j.ijpharm.2004.05.019","article-title":"The depolymerization of chitosan: Effects on physicochemical and biological properties","volume":"281","author":"Mao","year":"2004","journal-title":"Int. J. Pharm."},{"key":"ref_51","first-page":"361","article-title":"Changes of polydispersity and limiting molecular weight of ultrasound-treated chitosan","volume":"4","author":"Chen","year":"2000","journal-title":"Adv. Chitin Sci."},{"key":"ref_52","doi-asserted-by":"crossref","first-page":"557","DOI":"10.1007\/s10967-011-1310-9","article-title":"Water-soluble derivatives of chitosan as a target delivery system of 99mTc to some organs in vivo for nuclear imaging and biodistribution","volume":"290","author":"Hawary","year":"2011","journal-title":"J. Radioanal. Nucl. Chem."},{"key":"ref_53","doi-asserted-by":"crossref","first-page":"94","DOI":"10.1016\/j.lwt.2013.02.012","article-title":"Combined effect of N, O-carboxymethyl chitosan and oregano essential oil to extend shelf life and control Listeria monocytogenes in raw chicken meat fillets","volume":"53","author":"Khanjari","year":"2013","journal-title":"LWT-Food Sci. Technol."},{"key":"ref_54","doi-asserted-by":"crossref","first-page":"168","DOI":"10.1016\/j.colsurfb.2011.10.053","article-title":"Synthesis and characterization of N-(2-hydroxy)propyl-3-trimethyl ammonium chitosan chloride for potential application in gene delivery","volume":"91","author":"Xiao","year":"2012","journal-title":"Colloids Surf. B Biointerfaces"},{"key":"ref_55","doi-asserted-by":"crossref","first-page":"241","DOI":"10.1590\/S0104-14282009000300013","article-title":"A review of the antimicrobial activity of chitosan","volume":"19","author":"Goy","year":"2009","journal-title":"Pol\u00edmeros"},{"key":"ref_56","doi-asserted-by":"crossref","first-page":"427","DOI":"10.1016\/j.ijbiomac.2014.07.012","article-title":"Structure and antimicrobial mechanism of \u025b-polylysine\u2013chitosan conjugates through Maillard reaction","volume":"70","author":"Liang","year":"2014","journal-title":"Int. J. Biol. Macromol."},{"key":"ref_57","doi-asserted-by":"crossref","first-page":"33","DOI":"10.1080\/09583159830414","article-title":"EVect of Chitosan on Growth and Toxin Production by Alternaria alternata f. sp. lycopersici","volume":"8","author":"Reddy","year":"1998","journal-title":"Biocontrol Sci. Technol."},{"key":"ref_58","doi-asserted-by":"crossref","first-page":"122","DOI":"10.1016\/j.bjp.2015.09.010","article-title":"Evaluation of the antimicrobial activity of chitosan and its quaternized derivative on E. coli and S. aureus growth","volume":"26","author":"Goy","year":"2016","journal-title":"Revista Brasileira de Farmacognosia"},{"key":"ref_59","unstructured":"Jinasena, D., Pathirathna, P., Wickramarachchi, S., and Marasinghe, E. (2011, January 2). Use of chitosan to control anthracnose on \u201cEmbul\u201d banana. Proceedings of the 2011 International Conference on Asia Agriculture and Animal IPCBEE, Hong Kong, China."},{"key":"ref_60","doi-asserted-by":"crossref","first-page":"89","DOI":"10.1080\/01140671.1997.9513992","article-title":"Chitosan coating for inhibition of sclerotinia rot of carrots","volume":"25","author":"Cheah","year":"1997","journal-title":"N. Z. J. Crop Hortic. Sci."},{"key":"ref_61","doi-asserted-by":"crossref","first-page":"99","DOI":"10.1007\/BF02979773","article-title":"Further suppression of Botrytis cinerea disease in cucumber seedlings by chitosan-copper complex as compared with chitosan alone","volume":"31","author":"Fallik","year":"2003","journal-title":"Phytoparasitica"},{"key":"ref_62","unstructured":"Moret, A., Mu\u00f1oz, Z., and Garc\u00e9s, S. (2009). Control of powdery mildew on cucumber cotyledons by chitosan. J. Plant Pathol., 375\u2013380."},{"key":"ref_63","doi-asserted-by":"crossref","unstructured":"Long, L.T., Tan, L.V., Boi, V.N., and Trung, T.S. (2017). Antifungal activity of water-soluble chitosan against Colletotrichum capsici in post-harvest chili pepper. J. Food Process. Preserv.","DOI":"10.1111\/jfpp.13339"},{"key":"ref_64","doi-asserted-by":"crossref","first-page":"8038","DOI":"10.5897\/AJB10.984","article-title":"Induction of phenolics, lignin and key defense enzymes in eggplant (Solanum melongena L.) roots in response to elicitors","volume":"9","author":"Mandal","year":"2010","journal-title":"Afr. J. Biotechnol."},{"key":"ref_65","doi-asserted-by":"crossref","first-page":"211","DOI":"10.1080\/01140670709510187","article-title":"Effect of chitosan on ripening, enzymatic activity, and disease development in mango (Mangifera indica) fruit","volume":"35","author":"Jitareerat","year":"2007","journal-title":"N. Z. J. Crop Hortic. Sci."},{"key":"ref_66","doi-asserted-by":"crossref","first-page":"223","DOI":"10.1016\/j.scienta.2010.07.017","article-title":"Induction of disease resistance and ROS metabolism in navel oranges by chitosan","volume":"126","author":"Zeng","year":"2010","journal-title":"Sci. Hortic."},{"key":"ref_67","doi-asserted-by":"crossref","first-page":"70","DOI":"10.1016\/j.carbpol.2010.01.057","article-title":"Effects of chitosan and oligochitosan on growth of two fungal pathogens and physiological properties in pear fruit","volume":"81","author":"Meng","year":"2010","journal-title":"Carbohydr. Polym."},{"key":"ref_68","doi-asserted-by":"crossref","first-page":"1087","DOI":"10.1016\/S0261-2194(03)00117-0","article-title":"Effects of chitosan and plant extracts on growth of Colletotrichum gloeosporioides, anthracnose levels and quality of papaya fruit","volume":"22","author":"Wilson","year":"2003","journal-title":"Crop Prot."},{"key":"ref_69","first-page":"195","article-title":"Chitosan, antifungal product against Fusarium oxysporum f. sp. albedinis and elicitor of defence reactions in date palm roots","volume":"43","author":"Hassni","year":"2004","journal-title":"Phytopathol. Mediterr."},{"key":"ref_70","doi-asserted-by":"crossref","first-page":"272","DOI":"10.1016\/j.carbpol.2013.01.012","article-title":"Chitosan and oligochitosan enhance the resistance of peach fruit to brown rot","volume":"94","author":"Ma","year":"2013","journal-title":"Carbohydr. Polym."},{"key":"ref_71","doi-asserted-by":"crossref","first-page":"1777","DOI":"10.1080\/03235408.2013.858423","article-title":"Chitosan-induced defence responses in tomato plants against early blight disease caused by Alternaria solani (Ellis and Martin) Sorauer","volume":"47","author":"Sathiyabama","year":"2014","journal-title":"Arch. Phytopathol. Plant Prot."},{"key":"ref_72","doi-asserted-by":"crossref","first-page":"400","DOI":"10.1016\/j.carbpol.2015.07.066","article-title":"Fungal cell wall polymer based nanoparticles in protection of tomato plants from wilt disease caused by Fusarium oxysporum f. sp. lycopersici","volume":"133","author":"Sathiyabama","year":"2015","journal-title":"Carbohydr. Polym."},{"key":"ref_73","doi-asserted-by":"crossref","unstructured":"Escudero, N., Lopez-Moya, F., Ghahremani, Z., Zavala-Gonzalez, E.A., Alaguero-Cordovilla, A., Ros-Iba\u00f1ez, C., Lacasa, A., Sorribas, F.J., and Lopez-Llorca, L.V. (2017). Chitosan increases tomato root colonization by Pochonia chlamydosporia and their combination reduces root-knot nematode damage. Front. Plant Sci., 8.","DOI":"10.3389\/fpls.2017.01415"},{"key":"ref_74","first-page":"593","article-title":"Effects of Paenibacillus strains and chitosan on plant growth promotion and control of Ralstonia wilt in tomato","volume":"92","author":"Algam","year":"2010","journal-title":"J. Plant Pathol."},{"key":"ref_75","doi-asserted-by":"crossref","unstructured":"Nicaise, V. (2014). Crop immunity against viruses: Outcomes and future challenges. Front. Plant Sci., 5.","DOI":"10.3389\/fpls.2014.00660"},{"key":"ref_76","unstructured":"Faoro, F. (2018, April 08). Induced Systemic Resistance against Systemic Viruses: A Feasible Approach?. Available online: https:\/\/air.unimi.it\/retrieve\/handle\/2434\/229167\/299904\/Faoro%20F%20_%20IOBC%20Bull.%2089.pdf."},{"key":"ref_77","doi-asserted-by":"crossref","first-page":"774","DOI":"10.1023\/A:1012508625017","article-title":"Effect of chitosan on systemic viral infection and some defense responses in potato plants","volume":"48","author":"Chirkov","year":"2001","journal-title":"Russ. J. Plant Physiol."},{"key":"ref_78","first-page":"1520","article-title":"Response of Tomato Plants to Salicyli c Acid and Chitosan under Infection with Tomato mosaic virus","volume":"15","author":"Bondok","year":"2015","journal-title":"Am.-Eur. J. Agric. Environ. Sci."},{"key":"ref_79","first-page":"347","article-title":"Biocontrol of tomato leaf curl virus (ToLCV) in tomato with chitosan supplemented formulations of Pseudomonas sp. under field conditions","volume":"8","author":"Mishra","year":"2014","journal-title":"Aust. J. Crop Sci."},{"key":"ref_80","doi-asserted-by":"crossref","first-page":"148","DOI":"10.3923\/ajppaj.2017.148.155","article-title":"Use of Chitosan and Plant Growth Promoting Rhizobacteria to Control Squash Mosaic Virus on Cucumber Plants","volume":"11","author":"Firmansyah","year":"2017","journal-title":"Asian J. Plant Pathol."},{"key":"ref_81","doi-asserted-by":"crossref","first-page":"250","DOI":"10.1007\/s12250-014-3452-8","article-title":"Effect of chitosan on tobacco mosaic virus (TMV) accumulation, hydrolase activity, and morphological abnormalities of the viral particles in leaves of N. tabacum L. cv. Samsun","volume":"29","author":"Nagorskaya","year":"2014","journal-title":"Virol. Sin."},{"key":"ref_82","doi-asserted-by":"crossref","unstructured":"Jia, X., Meng, Q., Zeng, H., Wang, W., and Yin, H. (2016). Chitosan oligosaccharide induces resistance to Tobacco mosaic virus in Arabidopsis via the salicylic acid-mediated signalling pathway. Sci. Rep., 6.","DOI":"10.1038\/srep26144"},{"key":"ref_83","doi-asserted-by":"crossref","first-page":"951","DOI":"10.1002\/ps.1085","article-title":"Insecticidal and fungicidal activity of new synthesized chitosan derivatives","volume":"61","author":"Rabea","year":"2005","journal-title":"Pest Manag. Sci."},{"key":"ref_84","first-page":"589","article-title":"Synthesis, antifungal and insecticidal potential of Chitosan (CS)-g-poly (acrylic acid)(PAA) nanoparticles against some seed borne fungi and insects of soybean","volume":"8","author":"Sahab","year":"2015","journal-title":"Int. J. Chem. Tech. Res."},{"key":"ref_85","doi-asserted-by":"crossref","unstructured":"Li, Y., Qin, Y., Liu, S., Xing, R., Yu, H., Li, K., and Li, P. (2016). Preparation, Characterization, and Insecticidal Activity of Avermectin-Grafted-Carboxymethyl Chitosan. BioMed Res. Int., 2016.","DOI":"10.1155\/2016\/9805675"},{"key":"ref_86","doi-asserted-by":"crossref","first-page":"41","DOI":"10.1007\/s10343-016-0361-9","article-title":"Biochemical and Ultrastructural Changes of Some Tomato Cultivars after Infestation with Aphis gossypii Glover (Hemiptera: Aphididae) at Qalyubiyah, Egypt","volume":"68","author":"Helmi","year":"2016","journal-title":"Gesunde Pflanzen"},{"key":"ref_87","doi-asserted-by":"crossref","first-page":"175","DOI":"10.1007\/s13355-014-0319-2","article-title":"Tomato treatment with chemical inducers reduces the performance of Spodoptera littoralis (Lepidoptera: Noctuidae)","volume":"50","author":"Sobhy","year":"2015","journal-title":"Appl. Entomol. Zool."},{"key":"ref_88","doi-asserted-by":"crossref","first-page":"1379","DOI":"10.1603\/EC10120","article-title":"Effect of vermicompost and cucumber cultivar on population growth attributes of the melon aphid (Hemiptera: Aphididae)","volume":"104","author":"Razmjou","year":"2011","journal-title":"J. Econ. Entomol."},{"key":"ref_89","doi-asserted-by":"crossref","first-page":"2198","DOI":"10.1111\/j.1365-294X.2009.04190.x","article-title":"Ecological specialization of the aphid Aphis gossypii Glover on cultivated host plants","volume":"18","author":"Carletto","year":"2009","journal-title":"Mol. Ecol."},{"key":"ref_90","first-page":"891","article-title":"Optimization and application of microencapsulated artificial diet for Orius sauteri (Hemiptera: Anthocoridae)","volume":"53","author":"Tan","year":"2010","journal-title":"Acta Entomol. Sin."},{"key":"ref_91","first-page":"1","article-title":"Drought Stress in Plants: Causes, Consequences, and Tolerance","volume":"Volume 1","year":"2016","journal-title":"Drought Stress Tolerance in Plants"},{"key":"ref_92","doi-asserted-by":"crossref","first-page":"131","DOI":"10.1007\/s10725-009-9361-4","article-title":"Chitosan enhances leaf membrane stability and antioxidant enzyme activities in apple seedlings under drought stress","volume":"58","author":"Yang","year":"2009","journal-title":"Plant Growth Regul."},{"key":"ref_93","doi-asserted-by":"crossref","first-page":"293","DOI":"10.1007\/s11540-012-9223-8","article-title":"Effects of exogenous chitosan on physiological characteristics of potato seedlings under drought stress and rehydration","volume":"55","author":"Jiao","year":"2012","journal-title":"Potato Res."},{"key":"ref_94","first-page":"90","article-title":"Effects of exogenous Chitosan on physiological characteristics of phalaenopsis seedlings under draught stress","volume":"24","author":"Gu","year":"2011","journal-title":"Southwest China J. Agric. Sci."},{"key":"ref_95","doi-asserted-by":"crossref","first-page":"159","DOI":"10.1007\/s10725-013-9789-4","article-title":"The role of hydrogen peroxide in chitosan-induced resistance to osmotic stress in rice (Oryza sativa L.)","volume":"70","author":"Pongprayoon","year":"2013","journal-title":"Plant Growth Regul."},{"key":"ref_96","doi-asserted-by":"crossref","first-page":"3039","DOI":"10.1021\/acs.jproteome.7b00334","article-title":"Metabolic pathways regulated by chitosan contributing to drought resistance in white clover","volume":"16","author":"Li","year":"2017","journal-title":"J. Proteome Res."},{"key":"ref_97","first-page":"333","article-title":"The effect of chitosan on rooting of grapevine cuttings and on subsequent plant growth under drought and temperature stress","volume":"16","author":"Grzesik","year":"2008","journal-title":"J. Fruit Ornam. Plant Res."},{"key":"ref_98","doi-asserted-by":"crossref","first-page":"1106","DOI":"10.1016\/j.plaphy.2008.08.002","article-title":"Abscisic acid is involved in chitosan-induced resistance to tobacco necrosis virus (TNV)","volume":"46","author":"Iriti","year":"2008","journal-title":"Plant Physiol. Biochem."},{"key":"ref_99","doi-asserted-by":"crossref","first-page":"15251","DOI":"10.3390\/ijms160715251","article-title":"Function of ABA in stomatal defense against biotic and drought stresses","volume":"16","author":"Lim","year":"2015","journal-title":"Int. J. Mol. Sci."},{"key":"ref_100","unstructured":"McKersie, B.D., and Lesheim, Y. (2013). Stress and Stress Coping in Cultivated Plants, Springer."},{"key":"ref_101","doi-asserted-by":"crossref","first-page":"101","DOI":"10.1016\/j.scienta.2014.11.010","article-title":"Effect of zinc foliar spray alone and combined with humic acid or\/and chitosan on growth, nutrient elements content and yield of dry bean (Phaseolus vulgaris L.) plants sown at different dates","volume":"184","author":"Ibrahim","year":"2015","journal-title":"Sci. Hortic."},{"key":"ref_102","doi-asserted-by":"crossref","first-page":"567","DOI":"10.1038\/aps.2014.5","article-title":"Abscisic acid perception and signaling: Structural mechanisms and applications","volume":"35","author":"Ng","year":"2014","journal-title":"Acta Pharmacol. Sin."},{"key":"ref_103","doi-asserted-by":"crossref","first-page":"839","DOI":"10.1093\/jxb\/erm364","article-title":"Water deficits and heat shock effects on photosynthesis of a transgenic Arabidopsis thaliana constitutively expressing ABP9, a bZIP transcription factor","volume":"59","author":"Zhang","year":"2008","journal-title":"J. Exp. Bot."},{"key":"ref_104","doi-asserted-by":"crossref","first-page":"165","DOI":"10.1007\/s11816-012-0245-0","article-title":"Overexpression of ArabidopsisABF3 gene confers enhanced tolerance to drought and heat stress in creeping bentgrass","volume":"7","author":"Choi","year":"2013","journal-title":"Plant Biotechnol. Rep."},{"key":"ref_105","first-page":"673","article-title":"Effect of Nano-chitosan on Vegetative Growth, Fruiting and Resistance of Malformation of Mango","volume":"6","author":"Zagzog","year":"2017","journal-title":"Trends Hortic. Res."},{"key":"ref_106","doi-asserted-by":"crossref","first-page":"882","DOI":"10.1111\/j.1365-3059.2010.02312.x","article-title":"Inhibition of Botrytis cinerea growth and suppression of botrytis bunch rot in grapes using chitosan","volume":"59","author":"Reglinski","year":"2010","journal-title":"Plant Pathol."},{"key":"ref_107","doi-asserted-by":"crossref","first-page":"501","DOI":"10.1016\/j.foodchem.2007.06.012","article-title":"Physiological responses and quality attributes of table grape fruit to chitosan preharvest spray and post-harvest coating during storage","volume":"106","author":"Meng","year":"2008","journal-title":"Food Chem."},{"key":"ref_108","doi-asserted-by":"crossref","first-page":"1473","DOI":"10.1016\/j.foodchem.2010.07.114","article-title":"Chitosan elicits mono-glucosylated stilbene production and release in fed-batch bioreactor cultures of grape cells","volume":"124","author":"Ferri","year":"2011","journal-title":"Food Chem."},{"key":"ref_109","doi-asserted-by":"crossref","first-page":"887","DOI":"10.1007\/s10658-014-0515-5","article-title":"Field efficacy of chitosan to control Pseudomonas syringae pv. actinidiae, the causal agent of kiwifruit bacterial canker","volume":"140","author":"Scortichini","year":"2014","journal-title":"Eur. J. Plant Pathol."},{"key":"ref_110","doi-asserted-by":"crossref","first-page":"220","DOI":"10.1590\/1413-70542017412005917","article-title":"Pre-harvest application of calcium chloride and chitosan on fruit quality and storability of \u2018Early Swelling\u2019peach during cold storage","volume":"41","author":"Gayed","year":"2017","journal-title":"Ci\u00eancia Agrotecnol."},{"key":"ref_111","first-page":"675","article-title":"Effect of preharvest and post-harvest application of chitosan coating on storage quality of nectarines","volume":"1084","author":"Giacalone","year":"2013","journal-title":"Acta Hortic."},{"key":"ref_112","doi-asserted-by":"crossref","first-page":"434","DOI":"10.1590\/S1982-56762014000600003","article-title":"High-density chitosan reduces the severity of bacterial spot and activates the defense mechanisms of tomato plants","volume":"39","author":"Jail","year":"2014","journal-title":"Trop. Plant Pathol."},{"key":"ref_113","doi-asserted-by":"crossref","first-page":"742","DOI":"10.21273\/JASHS.125.6.742","article-title":"Chitosan effects on blackmold rot and pathogenic factors produced by Alternaria alternata in post-harvest tomatoes","volume":"125","author":"Reddy","year":"2000","journal-title":"J. Am. Soc. Hortic. Sci."},{"key":"ref_114","unstructured":"Tsugita, T., Takahashi, K., Muraoka, T., and Fukui, H. The application of chitin\/chitosan for agriculture. Proceedings of the Special Session of the 7th Symposium on Chitin and Chitosan, (In Japanese)."},{"key":"ref_115","doi-asserted-by":"crossref","first-page":"261","DOI":"10.1007\/BF02070695","article-title":"Chitin biodegradation and wound healing in tree bark tissues","volume":"4","author":"Hirano","year":"1996","journal-title":"J. Polym. Environ."},{"key":"ref_116","doi-asserted-by":"crossref","first-page":"3696","DOI":"10.1021\/jf0480804","article-title":"Effect of chitosan on the biological properties of sweet basil (Ocimum basilicum L.)","volume":"53","author":"Kim","year":"2005","journal-title":"J. Agric. Food Chem."},{"key":"ref_117","doi-asserted-by":"crossref","first-page":"608","DOI":"10.1007\/s00299-003-0733-3","article-title":"Chitosan improves development, and protects Vitis vinifera L. against Botrytis cinerea","volume":"22","author":"Barka","year":"2004","journal-title":"Plant Cell Rep."},{"key":"ref_118","unstructured":"Uragami, T., Kurita, K., and Fukamizo, T. (2000, January 21\u201323). Effects of Chitosan on the growth of Gerbera flower plant (Gerbera jamesonii). Proceedings of the Eighth International Chitin and Chitosan Conference and Fourth Asia Pacific Chitin and Chitosan Symposium, Yamaguchi, Japan."},{"key":"ref_119","first-page":"458","article-title":"The application of chitin and chitosan in agriculture in Thailand","volume":"5","author":"Chandrkrachang","year":"2002","journal-title":"Adv. Chitin Sci."},{"key":"ref_120","first-page":"441","article-title":"Effects of chitosan treatment on physiological and biochemical characteristics in cucumber seedlings under low temperature","volume":"30","author":"Xue","year":"2004","journal-title":"J. Plant Physiol. Mol. Biol."},{"key":"ref_121","doi-asserted-by":"crossref","unstructured":"Chookhongkha, N., Sopondilok, T., and Photchanachai, S. (2012, January 21). Effect of chitosan and chitosan nanoparticles on fungal growth and chilli seed quality. Proceedings of the International Conference on Post-harvest Pest and Disease Management in Exporting Horticultural Crops-PPDM2012 973, Bangkok, Thailand.","DOI":"10.17660\/ActaHortic.2013.973.32"},{"key":"ref_122","doi-asserted-by":"crossref","first-page":"853","DOI":"10.1016\/j.cropro.2010.03.002","article-title":"Application of bioactive coatings based on chitosan for artichoke seed protection","volume":"29","author":"Ziani","year":"2010","journal-title":"Crop Prot."},{"key":"ref_123","unstructured":"Chookhongkha, N., Miyagawa, S., Jirakiattikul, Y., and Photchanachai, S. (2012, January 17\u201318). Chili growth and seed productivity as affected by chitosan. Proceedings of the International Conference on Agriculture Technology and Food Sciences (ICATFS\u20192012), Manila, Philippines."},{"key":"ref_124","first-page":"63","article-title":"Response of cucumber plants to foliar application of chitosan and yeast under greenhouse conditions","volume":"6","author":"Shehata","year":"2012","journal-title":"Aust. J. Basic Appl. Sci."},{"key":"ref_125","doi-asserted-by":"crossref","first-page":"289","DOI":"10.1016\/j.bcab.2013.06.001","article-title":"Study on chitosan nanoparticles on biophysical characteristics and growth of Robusta coffee in green house","volume":"2","author":"Van","year":"2013","journal-title":"Biocatal. Agric. Biotechnol."},{"key":"ref_126","first-page":"673","article-title":"Effect of chitosan on seed germation, seedling growth and the clubroot control in Chinese cabbage","volume":"10","author":"Wang","year":"2012","journal-title":"J. Food Agric. Environ."},{"key":"ref_127","doi-asserted-by":"crossref","first-page":"239","DOI":"10.1016\/j.scienta.2009.11.019","article-title":"Chitosan specificity for the in vitro seed germination of two Dendrobium orchids (Asparagales: Orchidaceae)","volume":"124","author":"Kananont","year":"2010","journal-title":"Sci. Hortic."},{"key":"ref_128","first-page":"918","article-title":"Effect of foliar application of chitosan on growth and yield in okra","volume":"6","author":"Mondal","year":"2012","journal-title":"Aust. J. Crop Sci."},{"key":"ref_129","first-page":"570","article-title":"Induced resistance in potato plants against verticillium wilt invoked by chitosan and Acibenzolar-S-methyl","volume":"9","author":"Amini","year":"2015","journal-title":"Aust. J. Crop Sci."},{"key":"ref_130","first-page":"99","article-title":"Protective effect of humic acid and chitosan on radish (Raphanus sativus, L. var. sativus) plants subjected to cadmium stress","volume":"7","author":"Farouk","year":"2011","journal-title":"J. Stress Physiol. Biochem."},{"key":"ref_131","doi-asserted-by":"crossref","first-page":"448","DOI":"10.1016\/j.foodchem.2015.05.107","article-title":"Effects of preharvest applications of methyl jasmonate and chitosan on post-harvest decay, quality and chemical attributes of Fragaria chiloensis fruit","volume":"190","author":"Saavedra","year":"2016","journal-title":"Food Chem."},{"key":"ref_132","doi-asserted-by":"crossref","first-page":"93","DOI":"10.1166\/jcc.2014.1056","article-title":"Chitosan Increased Phenolic Compound Contents in Tea (Camellia sinensis) Leaves by Pre-and Post-Treatments","volume":"2","author":"Srisornkompon","year":"2014","journal-title":"J. Chitin Chitosan Sci."},{"key":"ref_133","doi-asserted-by":"crossref","unstructured":"Gonz\u00e1lez G\u00f3mez, H., Ram\u00edrez Godina, F., Ortega Ortiz, H., Benavides Mendoza, A., Robledo Torres, V., and Cabrera De la Fuente, M. (2017). Use of chitosan-PVA hydrogels with copper nanoparticles to improve the growth of grafted watermelon. Molecules, 22.","DOI":"10.3390\/molecules22071031"},{"key":"ref_134","first-page":"361","article-title":"New safe methods for controlling anthracnose disease of mango (Mangifera indica L.) fruits caused by Colletotrichum gloeosporioides (Penz.)","volume":"8","author":"Wafaa","year":"2010","journal-title":"J. Am. Sci."},{"key":"ref_135","first-page":"343","article-title":"Post-harvest quality of mango (Mangifera indica L.) fruit as affected by chitosan coating","volume":"41","author":"Abbasi","year":"2009","journal-title":"Pak. J. Bot"},{"key":"ref_136","doi-asserted-by":"crossref","first-page":"770","DOI":"10.1111\/j.1745-4549.2008.00213.x","article-title":"Effects of chitosan coating on post-harvest quality of mango (Mangifera indica L. cv. Tainong) fruits","volume":"32","author":"Zhu","year":"2008","journal-title":"J. Food Process. Preserv."},{"key":"ref_137","doi-asserted-by":"crossref","first-page":"435","DOI":"10.21608\/jfds.2016.46049","article-title":"Effect of Chitosan Edible Coating on Quality Attributes of Pomegranate Arils During Cold Storage","volume":"7","author":"Ashoush","year":"2016","journal-title":"J. Food Dairy Sci. Mansoura Univ."},{"key":"ref_138","doi-asserted-by":"crossref","first-page":"368","DOI":"10.1002\/jsfa.5770","article-title":"Effect of chitosan coating on maintenance of aril quality, microbial population and PPO activity of pomegranate (Punica granatum L. cv. Tarom) at cold storage temperature","volume":"93","author":"Ghasemnezhad","year":"2013","journal-title":"J. Sci. Food Agric."},{"key":"ref_139","doi-asserted-by":"crossref","first-page":"394","DOI":"10.1007\/s11947-014-1411-x","article-title":"The effect of chitosan coating on the quality and nutraceutical traits of sweet cherry during post-harvest life","volume":"8","author":"Petriccione","year":"2015","journal-title":"Food Bioprocess Technol."},{"key":"ref_140","doi-asserted-by":"crossref","first-page":"501","DOI":"10.3390\/foods4040501","article-title":"Effect of chitosan coating on the post-harvest quality and antioxidant enzyme system response of strawberry fruit during cold storage","volume":"4","author":"Petriccione","year":"2015","journal-title":"Foods"},{"key":"ref_141","doi-asserted-by":"crossref","first-page":"317","DOI":"10.1016\/j.apcbee.2014.03.047","article-title":"Effect of chitosan on the quality of rose apples (Syzygium agueum Alston) cv. Tabtim Chan stored at an ambient temperature","volume":"8","author":"Plainsirichai","year":"2014","journal-title":"APCBEE Procedia"},{"key":"ref_142","first-page":"25","article-title":"Effect of chitosan coatings on some quality indices of apricot (Prunus armeniaca L.) during cold storage","volume":"8","author":"Ghasemnezhad","year":"2010","journal-title":"Casp. J. Environ. Sci."},{"key":"ref_143","doi-asserted-by":"crossref","first-page":"113","DOI":"10.1016\/j.proche.2014.05.014","article-title":"Improving shelf-life of cavendish banana using chitosan edible coating","volume":"9","author":"Suseno","year":"2014","journal-title":"Procedia Chem."},{"key":"ref_144","doi-asserted-by":"crossref","unstructured":"El Guilli, M., Hamza, A., Cl\u00e9ment, C., Ibriz, M., and Ait Barka, E. (2016). Effectiveness of post-harvest treatment with chitosan to control citrus green mold. Agriculture, 6.","DOI":"10.3390\/agriculture6020012"},{"key":"ref_145","doi-asserted-by":"crossref","first-page":"269","DOI":"10.1016\/j.carbpol.2017.06.002","article-title":"Effect of molecular weight of chitosan on the shelf life and other quality parameters of three different cultivars of Actinidia kolomikta (kiwifruit)","volume":"173","author":"Drevinskas","year":"2017","journal-title":"Carbohydr. Polym."},{"key":"ref_146","doi-asserted-by":"crossref","first-page":"238","DOI":"10.9755\/ejfa.v26i3.16620","article-title":"Effect of edible chitosan\/zeolite coating on tomatoes quality during refrigerated storage","volume":"26","author":"Casariego","year":"2014","journal-title":"Emirates J. Food Agric."},{"key":"ref_147","first-page":"141","article-title":"Use of chitosan film coatings in the storage of carrots (Daucus carota)","volume":"13","author":"Zlotek","year":"2008","journal-title":"Prog. Chem. Appl. Chitin Deriv."},{"key":"ref_148","doi-asserted-by":"crossref","first-page":"M367","DOI":"10.1111\/j.1750-3841.2011.02210.x","article-title":"Effectiveness of edible coatings combined with mild heat shocks on microbial spoilage and sensory quality of fresh cut broccoli (Brassica oleracea L.)","volume":"76","author":"Moreira","year":"2011","journal-title":"J. Food Sci."},{"key":"ref_149","doi-asserted-by":"crossref","first-page":"29","DOI":"10.1016\/j.postharvbio.2015.11.004","article-title":"Chitosan coating with trans-cinnamaldehyde improves structural integrity and antioxidant metabolism of fresh-cut melon","volume":"113","author":"Carvalho","year":"2016","journal-title":"Post-harvest Biol. Technol."},{"key":"ref_150","doi-asserted-by":"crossref","first-page":"558","DOI":"10.1016\/j.foodchem.2014.11.106","article-title":"Post-harvest chitosan-g-salicylic acid application alleviates chilling injury and preserves cucumber fruit quality during cold storage","volume":"174","author":"Zhang","year":"2015","journal-title":"Food Chem."},{"key":"ref_151","doi-asserted-by":"crossref","unstructured":"Hadwiger, L., Kendra, D., Fristensky, B., and Wagoner, W. (1986). Chitosan both activates genes in plants and inhibits RNA synthesis in fungi. Chitin in Nature and Technology, Springer.","DOI":"10.1007\/978-1-4613-2167-5_28"},{"key":"ref_152","doi-asserted-by":"crossref","first-page":"65","DOI":"10.1016\/j.scienta.2007.10.034","article-title":"Chitosan effects on floral production, gene expression, and anatomical changes in the Dendrobium orchid","volume":"116","author":"Limpanavech","year":"2008","journal-title":"Sci. Hortic."},{"key":"ref_153","doi-asserted-by":"crossref","first-page":"42","DOI":"10.1016\/j.plantsci.2013.03.007","article-title":"Multiple effects of chitosan on plant systems: Solid science or hype","volume":"208","author":"Hadwiger","year":"2013","journal-title":"Plant Sci."},{"key":"ref_154","doi-asserted-by":"crossref","first-page":"85","DOI":"10.1006\/pmpp.1999.0213","article-title":"The families of pathogenesis-related proteins, their activities, and comparative analysis of PR-1 type proteins","volume":"55","year":"1999","journal-title":"Physiol. Mol. Plant Pathol."},{"key":"ref_155","doi-asserted-by":"crossref","first-page":"274","DOI":"10.1111\/aab.12299","article-title":"Bio-efficacy of a chitosan based elicitor on Alternaria solani and Xanthomonas vesicatoria infections in tomato under tropical conditions","volume":"169","author":"Ramkissoon","year":"2016","journal-title":"Ann. Appl. Biol."},{"key":"ref_156","first-page":"461","article-title":"Expression of Plant Defense Genes in Pepper Seedlings Treated with Chitosan Solution","volume":"1088","author":"Jitareerat","year":"2013","journal-title":"Southeast Asia Symp. Qual. Manag. Post-harvest Syst."},{"key":"ref_157","doi-asserted-by":"crossref","first-page":"3047","DOI":"10.1021\/jf404423x","article-title":"Expression of defense genes in strawberry fruits treated with different resistance inducers","volume":"62","author":"Landi","year":"2014","journal-title":"J. Agric. Food Chem."},{"key":"ref_158","doi-asserted-by":"crossref","first-page":"71","DOI":"10.1016\/j.lwt.2012.05.003","article-title":"Effect of chitosan-based edible coating on antioxidants, antioxidant enzyme system, and post-harvest fruit quality of strawberries (Fragaria x aranassa Duch.)","volume":"52","author":"Wang","year":"2013","journal-title":"LWT-Food Sci. Technol."},{"key":"ref_159","doi-asserted-by":"crossref","first-page":"763","DOI":"10.1080\/14620316.1998.11511045","article-title":"Antifungal effects of chitosan coating on fresh strawberries and raspberries during storage","volume":"73","author":"Zhang","year":"1998","journal-title":"J. Hortic. Sci. Biotechnol."},{"key":"ref_160","doi-asserted-by":"crossref","first-page":"307","DOI":"10.1094\/PDIS-12-11-1043-RE","article-title":"Preharvest fungicide, potassium sorbate, or chitosan use on quality and storage decay of table grapes","volume":"97","author":"Feliziani","year":"2013","journal-title":"Plant Dis."},{"key":"ref_161","doi-asserted-by":"crossref","unstructured":"Berg, J.A., Appiano, M., Bijsterbosch, G., Visser, R.G., Schouten, H.J., and Bai, Y. (2017). Functional characterization of cucumber (Cucumis sativus L.) Clade V MLO genes. BMC Plant Biol., 17.","DOI":"10.1186\/s12870-017-1029-z"},{"key":"ref_162","doi-asserted-by":"crossref","first-page":"364","DOI":"10.1016\/j.ijbiomac.2014.06.060","article-title":"Quaternized chitosan oligomers as novel elicitors inducing protection against B. cinerea in Arabidopsis","volume":"72","author":"Feng","year":"2015","journal-title":"Int. J. Biol. Macromol."},{"key":"ref_163","doi-asserted-by":"crossref","first-page":"10178","DOI":"10.3390\/ijms140510178","article-title":"Oxidative and molecular responses in Capsicum annuum L. after hydrogen peroxide, salicylic acid and chitosan foliar applications","volume":"14","year":"2013","journal-title":"Int. J. Mol. Sci."},{"key":"ref_164","first-page":"2807","article-title":"Cloning and expression analysis of transcription factor gene DoWRKY1 in Dendrobium officinale","volume":"40","author":"Zhao","year":"2015","journal-title":"China J. Chin. Mater. Med."},{"key":"ref_165","doi-asserted-by":"crossref","first-page":"474","DOI":"10.1016\/j.carbpol.2016.05.103","article-title":"Chitosan and oligochitosan enhance ginger (Zingiber officinale Roscoe) resistance to rhizome rot caused by Fusarium oxysporum in storage","volume":"151","author":"Liu","year":"2016","journal-title":"Carbohydr. Polym."},{"key":"ref_166","first-page":"16","article-title":"Effect of chitosan on the induction of disease resistance against Colletotrichum sp. in mango (Mangifera indica L) cv","volume":"66","year":"2015","journal-title":"Tommy Atkins"},{"key":"ref_167","doi-asserted-by":"crossref","first-page":"174","DOI":"10.1016\/j.carbpol.2014.06.077","article-title":"Quaternary chitosan oligomers enhance resistance and biocontrol efficacy of Rhodosporidium paludigenum to green mold in satsuma orange","volume":"113","author":"Lu","year":"2014","journal-title":"Carbohydr. Polym."},{"key":"ref_168","doi-asserted-by":"crossref","unstructured":"Coqueiro, D.S.O., de Souza, A.A., Takita, M.A., Rodrigues, C.M., Kishi, L.T., and Machado, M.A. (2015). Transcriptional profile of sweet orange in response to chitosan and salicylic acid. BMC Genom., 16.","DOI":"10.1186\/s12864-015-1440-5"},{"key":"ref_169","doi-asserted-by":"crossref","first-page":"1340","DOI":"10.1007\/s13197-017-2554-3","article-title":"Effect of chitosan treatment on strawberry allergen-related gene expression during ripening stages","volume":"54","author":"Petriccione","year":"2017","journal-title":"J. Food Sci. Technol."},{"key":"ref_170","doi-asserted-by":"crossref","first-page":"113","DOI":"10.1093\/mp\/ssp087","article-title":"The strawberry fruit Fra a allergen functions in flavonoid biosynthesis","volume":"3","author":"Hoffmann","year":"2010","journal-title":"Mol. Plant"},{"key":"ref_171","doi-asserted-by":"crossref","first-page":"374","DOI":"10.1016\/j.plaphy.2016.06.034","article-title":"Induction of aromatic amino acids and phenylpropanoid compounds in Scrophularia striata Boiss. cell culture in response to chitosan-induced oxidative stress","volume":"107","author":"Kamalipourazad","year":"2016","journal-title":"Plant Physiol. Biochem."},{"key":"ref_172","doi-asserted-by":"crossref","first-page":"7399","DOI":"10.1021\/acs.jafc.5b01566","article-title":"Chitosan controls post-harvest decay on cherry tomato fruit possibly via the mitogen-activated protein kinase signaling pathway","volume":"63","author":"Zhang","year":"2015","journal-title":"J. Agric. Food Chem."},{"key":"ref_173","doi-asserted-by":"crossref","first-page":"322","DOI":"10.1094\/PDIS.1998.82.3.322","article-title":"Effects of chitin and chitosan on the incidence and severity of Fusarium yellows of celery","volume":"82","author":"Bell","year":"1998","journal-title":"Plant Dis."},{"key":"ref_174","doi-asserted-by":"crossref","first-page":"153","DOI":"10.1016\/S0929-1393(00)00091-3","article-title":"Stimulation of wild strawberry (Fragaria vesca) arbuscular mycorrhizas by addition of shellfish waste to the growth substrate: Interaction between mycorrhization, substrate amendment and susceptibility to red core (Phytophthora fragariae)","volume":"15","author":"Murphy","year":"2000","journal-title":"Appl. Soil Ecol."},{"key":"ref_175","unstructured":"Inc, N.T. (2010). Efficacit\u00b4e du Lysozyme dans le Contr\u02c6olede la Croissance des Agents Pathog`enes des Plants de Serres, Neova Technologies Inc."},{"key":"ref_176","doi-asserted-by":"crossref","first-page":"201","DOI":"10.1007\/BF02803152","article-title":"The effects of arbuscular mycorrhizal fungi and chitosan sprays on yield and late blight resistance in potato crops from microplants","volume":"38","author":"Duffy","year":"2003","journal-title":"Folia Geobot."},{"key":"ref_177","doi-asserted-by":"crossref","first-page":"63","DOI":"10.2503\/jjshs.69.63","article-title":"Effects of chitosan with or without nitrogen treatments on seedling growth in Eustoma grandiflorum (Raf.) Shinn. Cv. Kairyou Wakamurasaki","volume":"69","author":"Ohta","year":"2000","journal-title":"J. Jpn. Soc. Hortic. Sci."},{"key":"ref_178","doi-asserted-by":"crossref","first-page":"621","DOI":"10.1080\/14620316.1988.11515902","article-title":"Evaluation of a protein-chitin derivative of crustacean shells as a slow-release nitrogen fertilizer on Chinese cabbage","volume":"63","author":"Spiegel","year":"1988","journal-title":"J. Hortic. Sci."},{"key":"ref_179","doi-asserted-by":"crossref","first-page":"1","DOI":"10.1007\/s13580-016-0119-4","article-title":"A combination of chitosan and chemical fertilizers improves growth and disease resistance in Begonia\u00d7 hiemalis Fotsch","volume":"57","author":"Chen","year":"2016","journal-title":"Hortic. Environ. Biotechnol."},{"key":"ref_180","doi-asserted-by":"crossref","first-page":"618","DOI":"10.1139\/w01-057","article-title":"Chitin-supplemented formulations improve biocontrol and plant growth promoting efficiency of Bacillus subtilis AF 1","volume":"47","author":"Manjula","year":"2001","journal-title":"Can. J. Microbiol."},{"key":"ref_181","unstructured":"Bakiyalakshmi, S.V., Valli, V., and Swarnila, R.D.L. (2016). Isolation and Application of Chitin and Chitosan from crab shell. Int. J. Curr. Microbiol. Appl. Sci., 91\u201399."},{"key":"ref_182","doi-asserted-by":"crossref","first-page":"236","DOI":"10.1016\/j.scienta.2016.06.027","article-title":"The impact of biofertilizers with diazotrophic bacteria and fungi chitosan on melon characteristics and nutrient uptake as an alternative for conventional fertilizers","volume":"209","author":"Silva","year":"2016","journal-title":"Sci. Hortic."},{"key":"ref_183","unstructured":"Salachna, P., Wilas, J., and Zawadzi\u0144ska, A. (2014, January 17\u201322). The effect of chitosan coating of bulbs on the growth and flowering of Ornithogalum saundersiae. Proceedings of the XXIX International Horticultural Congress on Horticulture: Sustaining Lives, Livelihoods and Landscapes (IHC2014), Brisbane, Australia."},{"key":"ref_184","doi-asserted-by":"crossref","unstructured":"Salachna, P., Grzeszczuk, M., and Sob\u00f3l, M. (2017). Effects of Chitooligosaccharide Coating Combined with Selected Ionic Polymers on the Stimulation of Ornithogalum saundersiae Growth. Molecules, 22.","DOI":"10.3390\/molecules22111903"},{"key":"ref_185","unstructured":"Trenkel, M.E. (1997). Controlled-Release and Stabilized Fertilizers in Agriculture, International Fertilizer Industry Association Paris."},{"key":"ref_186","doi-asserted-by":"crossref","first-page":"287","DOI":"10.1016\/j.apcbee.2012.03.047","article-title":"Investigation of effect of chemical fertilizers on environment","volume":"1","author":"Savci","year":"2012","journal-title":"Apcbee Procedia"},{"key":"ref_187","doi-asserted-by":"crossref","first-page":"355","DOI":"10.1080\/02652030500482363","article-title":"Nitrates and nitrites in vegetables and vegetable-based products and their intakes by the Estonian population","volume":"23","author":"Tamme","year":"2006","journal-title":"Food Addit. Contam."},{"key":"ref_188","doi-asserted-by":"crossref","first-page":"113","DOI":"10.4141\/CJSS07102","article-title":"Opportunities for improved fertilizer nitrogen management in production of arable crops in eastern Canada: A review","volume":"89","author":"Zebarth","year":"2009","journal-title":"Can. J. Soil Sci."},{"key":"ref_189","doi-asserted-by":"crossref","first-page":"240","DOI":"10.1016\/j.carbpol.2007.08.020","article-title":"Preparation and properties of chitosan-coated NPK compound fertilizer with controlled-release and water-retention","volume":"72","author":"Wu","year":"2008","journal-title":"Carbohydr. Polym."},{"key":"ref_190","doi-asserted-by":"crossref","first-page":"509","DOI":"10.3144\/expresspolymlett.2010.64","article-title":"A preliminary study of the incorparation of NPK fertilizer into chitosan nanoparticles","volume":"4","author":"Corradini","year":"2010","journal-title":"Express Polym. Lett."},{"key":"ref_191","doi-asserted-by":"crossref","first-page":"473","DOI":"10.1007\/s13726-012-0051-0","article-title":"Controlled release of urea from chitosan microspheres prepared by emulsification and cross-linking method","volume":"21","author":"Hussain","year":"2012","journal-title":"Iranian Polym. J."},{"key":"ref_192","unstructured":"H\u00e4hndel, R., and Aktiengesellschaft, B. Personal communication."},{"key":"ref_193","doi-asserted-by":"crossref","first-page":"1061","DOI":"10.1016\/j.ijbiomac.2017.11.092","article-title":"Synthesis, characterization, and the antifungal activity of chitosan derivatives containing urea groups","volume":"109","author":"Zhang","year":"2018","journal-title":"Int. J. Biol. Macromol."}],"container-title":["Molecules"],"original-title":[],"language":"en","link":[{"URL":"https:\/\/www.mdpi.com\/1420-3049\/23\/4\/872\/pdf","content-type":"unspecified","content-version":"vor","intended-application":"similarity-checking"}],"deposited":{"date-parts":[[2025,10,11]],"date-time":"2025-10-11T15:00:16Z","timestamp":1760194816000},"score":1,"resource":{"primary":{"URL":"https:\/\/www.mdpi.com\/1420-3049\/23\/4\/872"}},"subtitle":[],"short-title":[],"issued":{"date-parts":[[2018,4,10]]},"references-count":193,"journal-issue":{"issue":"4","published-online":{"date-parts":[[2018,4]]}},"alternative-id":["molecules23040872"],"URL":"https:\/\/doi.org\/10.3390\/molecules23040872","relation":{},"ISSN":["1420-3049"],"issn-type":[{"value":"1420-3049","type":"electronic"}],"subject":[],"published":{"date-parts":[[2018,4,10]]}}}